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Mantis shrimp

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853:(tiers), each for different wavelengths. The three tiers in rows 2 and 3 are separated by colour filters (intrarhabdomal filters) that can be divided into four distinct classes, two classes in each row. Each consists of a tier, a colour filter of one class, a tier again, a colour filter of another class, and then a last tier. These colour filters allow the mantis shrimp to see with diverse colour vision. Without the filters, the pigments themselves range only a small segment of the visual spectrum, about 490 to 550 nm. Rows 5 and 6 are also segregated into different tiers, but have only one class of visual pigment, the ninth class, and are specialised for polarisation vision. Depending upon the species, they can detect circularly polarised light, linearly polarised light, or both. A tenth class of visual pigment is found in the upper and lower hemispheres of the eye. 592:
mantis shrimp into two main types: those that hunt by impaling their prey with spear-like structures and those that smash prey with a powerful blow from a heavily mineralised club-like appendage. A considerable amount of damage can be inflicted after impact with these robust, hammer-like claws. This club is further divided into three subregions: the impact region, the periodic region, and the striated region. Mantis shrimp are commonly separated into many (most fall into spears and smashers but there are some outliers) distinct groups determined by the type of claws they possess:
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the human brain, the inferior temporal cortex has a huge number of colour-specific neurons, which process visual impulses from the eyes to extract colour information. The mantis shrimp instead uses the different types of photoreceptors in its eyes to perform the same function as the human brain neurons, resulting in a hardwired and more efficient system for an animal that requires rapid colour identification. Humans have fewer types of photoreceptors, but more colour-tuned neurons, while mantis shrimp appear to have fewer colour neurons and more classes of photoreceptors.
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muscles divided into six functional groups. By using these muscles to scan the surroundings with the midband, they can add information about forms, shapes, and landscape, which cannot be detected by the upper and lower hemispheres of the eyes. They can also track moving objects using large, rapid eye movements where the two eyes move independently. By combining different techniques, including movements in the same direction, the midband can cover a very wide range of the visual field.
541: 773: 155: 806:, clusters of photoreceptor cells. Each eye consists of two flattened hemispheres separated by parallel rows of specialised ommatidia, collectively called the midband. The number of omatidial rows in the midband ranges from two to six. This divides the eye into three regions. This configuration enables mantis shrimp to see objects that are near the mid-plane of an eye with three parts of the same eye (as can be seen in some photos showing three 1350: 113: 907:, which yield a full description of polarisation. It is thus believed to have optimal polarisation vision. It is the only animal known to have dynamic polarisation vision. This is achieved by rotational eye movements to maximise the polarisation contrast between the object in focus and its background. Since each eye moves independently from the other, it creates two separate streams of visual information. 1662: 785: 1118: 991: 571: 1210: 916: 700: 1075: 764:
lost the ancestral phenotype, although some still maintain 16 distinct photoreceptors and four light filters. Species that live in a variety of photic environments have high selective pressure for photoreceptor diversity, and maintain ancestral phenotypes better than species that live in murky waters or are primarily nocturnal.
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In a lifetime, they can have as many as 20 or 30 breeding episodes. Depending on the species, the eggs can be laid and kept in a burrow, or they can be carried around under the female's tail until they hatch. Also depending on the species, males and females may come together only to mate, or they may
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patterns on their bodies for signalling with their own and maybe even other species, expanding their range of behavioural signals. They can learn and remember well, and are able to recognise individual neighbours with which they frequently interact. They can recognise them by visual signs and even by
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The shrimps use a form of reflector of polarised light not seen in nature or human technology before. It allows the manipulation of light across the structure rather than through its depth, the typical way polarisers work. This allows the structure to be both small and microscopically thin, and still
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What advantage sensitivity to polarisation confers is unclear; however, polarisation vision is used by other animals for sexual signaling and secret communication that avoids the attention of predators. This mechanism could provide an evolutionary advantage; it only requires small changes to the cell
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events. One consequence of this duplication is the lack of correlation between opsin transcript number and physiologically expressed photoreceptors. One species may have six different opsin genes, but only express one spectrally distinct photoreceptor. Over the years, some mantis shrimp species have
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and can move independently of each other. The extreme mobility allows them to be rotated in all three dimensions, yet the position of their eyes has shown to have no effect on the perception of their surroundings. They are thought to have the most complex eyes in the animal kingdom and have the most
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possess a much more developed club and a more rudimentary spear (which is nevertheless quite sharp and still used in fights between their own kind); the club is used to bludgeon and smash their meals apart. The inner aspect of the terminal portion of the appendage can also possess a sharp edge, used
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Mantis shrimp possess one of the most complex visual systems among all animals, with compound eyes capable of detecting a wider range of colors and polarized light than humans. Additionally, their eyes are capable of independently moving and focusing on different objects, allowing for simultaneous
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colour perception used by other species, thus requiring more time and energy. While the eyes themselves are complex and not yet fully understood, the principle of the system appears to be simple. It has a similar set of sensitivities to the human visual system, but works in the opposite manner. In
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Six species of mantis shrimp have been reported to be able to detect circularly polarised light, which has not been documented in any other animal, and whether it is present across all species is unknown. They perform this feat by converting circularly polarized light into linearly polarized light
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The midband covers only about 5 to 10° of the visual field at any given instant, but like most crustaceans, mantis shrimps' eyes are mounted on stalks. In mantis shrimps, the movement of the stalked eye is unusually free, and can be driven up to 70° in all possible axes of movement by eight eyecup
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Some species have at least 16 photoreceptor types, which are divided into four classes (their spectral sensitivity is further tuned by colour filters in the retinas), 12 for colour analysis in the different wavelengths (including six which are sensitive to ultraviolet light) and four for analysing
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and the smashing species make burrows in hard substrata or coral cavities. These two habitats are crucial for their ecology since they use burrows as sites for retreat and as locations for consuming their prey. Burrows and coral cavities are also used as sites for mating and for keeping their eggs
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Rows 1 to 4 of the midband are specialised for colour vision, from deep ultraviolet to far red. Their UV vision can detect five different frequency bands in the deep ultraviolet. To do this, they use two photoreceptors in combination with four different colour filters. They are currently believed
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The mantis shrimp's second pair of thoracic appendages has been highly adapted for powerful close-range combat. These claws can accelerate at a rate comparable to that of a 0.22 caliber bullet when fired, having around 1500 newtons of force with each swing/attack. The appendage differences divide
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trade and is often collected. A piece of live rock not uncommonly conveys a live mantis shrimp into an aquarium. Once inside the tank, it may feed on fish and other inhabitants, and is notoriously difficult to catch when established in a well-stocked tank. While there are accounts of this shrimp
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that humans possess in their eyes, the eyes of a mantis shrimp have between 12 and 16 types of photoreceptor cells. Furthermore, some of these stomatopods can tune the sensitivity of their long-wavelength colour vision to adapt to their environment. This phenomenon, called "spectral tuning", is
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apart. It is suggested that not discriminating between closely positioned wavelengths allows these organisms to make determinations of its surroundings with little processing delay. Having little delay in evaluating surroundings is important for mantis shrimp, since they are territorial and
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that are caused by the impact of the appendage against the striking surface, which means that the prey is hit twice by a single strike; first by the claw and then by the collapsing cavitation bubbles that immediately follow. Even if the initial strike misses the prey, the resulting
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safe. Stomatopod body size undergoes periodic growth which necessitates finding a new cavity or burrow that will fit the animal's new diameter. Some spearing species can modify their pre-established habitat if the burrow is made of silt or mud, which can be expanded.
1980: 1031:, the female lays two clutches of eggs – one that the male tends and one that the female tends. In other species, the female looks after the eggs while the male hunts for both of them. After the eggs hatch, the offspring may spend up to three months as 736:, a species with a variety of habitats ranging from a depth of 5 to 10 m (although it can be found down to 20 m below the surface), spectral tuning was observed, but the ability to alter wavelengths of maximum absorbance was not as pronounced as in 961:
Researchers suspect that the broader variety of photoreceptors in the eyes of mantis shrimps allows visual information to be preprocessed by the eyes instead of the brain, which would otherwise have to be larger to deal with the complex task of
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In the monogamous species, the mantis shrimps remain with the same partner up to 20 years. They share the same burrow and may be able to coordinate their activities. Both sexes often take care of the eggs (bi-parental care). In
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is often stranded by its short rear legs, which are sufficient for movement when the body is supported by water, but not on dry land. The mantis shrimp then performs a forward flip in an attempt to roll towards the next tide pool.
890:. Some of their biological quarter-waveplates perform more uniformly over the visual spectrum than any current man-made polarising optics, and this could inspire new types of optical media that would outperform early 21st century 1368:
While some aquarists value mantis shrimps, others consider them harmful pests, because they are voracious predators, eating other desirable inhabitants of the tank. Additionally, some rock-burrowing species can do more damage to
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These aggressive and typically solitary sea creatures spend most of their time hiding in rock formations or burrowing intricate passageways in the sea bed. They rarely exit their homes except to feed and relocate, and can be
1265:, and like lobsters, their shells are quite hard and require some pressure to crack. One common preparation is first deep-frying, then stir-frying with garlic and chili peppers. They may also be boiled or steamed. 524:(active at twilight), depending on the species. Unlike most crustaceans, they sometimes hunt, chase, and kill prey. Although some live in temperate seas, most species live in tropical and subtropical waters in the 630:
claws at the prey, and can inflict serious damage on victims significantly greater in size than themselves. In smashers, these two weapons are employed with blinding quickness, with an acceleration of 10,400 
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breaking glass tanks, they are rare and are usually the result of the shrimp being kept in too small a tank. While stomatopods do not eat coral, smashers can damage it if they try to make a home within it.
1849: 833:. In mantis shrimp in the superfamilies Gonodactyloidea, Lysiosquilloidea, and Hemisquilloidea, the midband is made up of six ommatidial rows. Rows 1 to 4 process colours, while rows 5 and 6 detect 857:
polarised light. By comparison, most humans have only four visual pigments, of which three are dedicated to see colour, and human lenses block ultraviolet light. The visual information leaving the
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Roberts, Nicholas W.; Chiou, Tsyr-Huei; Marshall, Nicholas Justin; Cronin, Thomas W. (2009). "A biological quarter-wave retarder with excellent achromaticity in the visible wavelength region".
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may, therefore, help prevent wasted mating efforts. It may also give these shrimps information about the size of the tide, which is important to species living in shallow water near the shore.
2095: 2273: 3457: 2576: 3641: 740:, a species with much higher ecological/photic habitat diversity. The diversity of spectral tuning in Stomatopoda is also hypothesised to be directly linked to mutations in the 439:
around 340 million years ago. Mantis shrimp typically grow to around 10 cm (3.9 in) in length, while a few can reach up to 38 cm (15 in). A mantis shrimp's
2149: 485:. Some mantis shrimp species have specialised calcified 'clubs' that can strike with great power, while others have sharp forelimbs used to seize the prey (hence the term " 2980: 4059: 936:
The eyes of mantis shrimps may enable them to recognise different types of coral, prey species (which are often transparent or semitransparent), or predators, such as
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Despite the impressive range of wavelengths that mantis shrimp have the ability to see, they do not have the ability to discriminate wavelengths less than 25 
2066: 1857: 818:, for objects near its mid-plane. The upper and lower hemispheres are used primarily for recognition of form and motion, like the eyes of many other crustaceans. 1426:, the latter of which are more closely related to modern mantis shrimp, which are assigned to the clade Unipeltata. The oldest members of Unipeltata date to the 648:) from a standing start. Because they strike so rapidly, they generate vapor-filled bubbles in the water between the appendage and the striking surface—known as 6151: 3029: 2597:
Porter, Megan L.; Bok, Michael J.; Robinson, Phyllis R.; Cronin, Thomas W. (1 May 2009). "Molecular diversity of visual pigments in Stomatopoda (Crustacea)".
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How, M. J.; Porter, M. L.; Radford, A. N.; Feller, K. D.; Temple, S. E.; Caldwell, R. L.; Marshall, N. J.; Cronin, T. W.; Roberts, N. W. (7 August 2014).
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However, a study from 2022 failed to find unequivocal evidence for a solely "barcode"-like visual system as described above. Stomatopods of the species
947:, and the wavelength of this fluorescence matches the wavelengths detected by their eye pigments. Females are only fertile during certain phases of the 477:, and now sometimes referred to as "thumb splitters" due to their ability to inflict painful wounds if handled incautiously mantis shrimp have powerful 447:. Varieties range in colour from shades of brown to vivid colours, with more than 520 species of mantis shrimp known. They are among the most important 3376: 3119:
Chiou, Tsyr-Huei; Kleinlogel, Sanja; Cronin, Tom; Caldwell, Roy; Loeffler, Birte; Siddiqi, Afsheen; Goldzien, Alan; Marshall, Justin (March 25, 2008).
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Thoen, Hanne H.; How, Martin J.; Chiou, Tsyr-Huei; Marshall, Nicholas Justin (January 24, 2014). "A Different Form of Color Vision in Mantis Shrimp".
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Smith, C.P.A.; Aubier, P.; Charbonnier, S.; Laville, T.; Olivier, N.; Escarguel, G.; Jenks, J.F.; Bylund, K.G.; Fara, E.; Brayard, A. (2023-03-31).
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bubbles. The collapse of these cavitation bubbles produces measurable forces on their prey in addition to the instantaneous forces of 1,500 
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frequently in combat. However, some mantis shrimp have been found capable of distinguishing between high-saturation and low-saturation colors.
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are armed with spiny appendages - the spines having barbed tips - used to stab and snag prey. These raptorial appendages resemble those of
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is the only organism known to simultaneously detect the four linear and two circular polarisation components required to measure all four
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were able to distinguish high and low-saturation colors from grey, contravening Thoen and colleagues. It may be that some combination of
3784: 3665: 1646:; the collection of stomatopods he amassed is the largest in the world, covering 90% of the known species whilst 10% are still unknown. 463:. However, despite being common, they are poorly understood, as many species spend most of their lives sheltering in burrows and holes. 4093: 3795: 3686: 3254:"Circularly polarized light detection in stomatopod crustaceans: a comparison of photoreceptors and possible function in six species" 2848:
Marshall, Nicholas Justin; Oberwinkler, Johannes (October 28, 1999). "Ultraviolet vision: the colourful world of the mantis shrimp".
2528:"Evolutionary variation in the expression of phenotypically plastic color vision in Caribbean mantis shrimps, genus Neogonodactylus" 1964: 1955:
Mead, K.; Caldwell, R. (2001). "Mantis Shrimp: Olfactory Apparatus and Chemosensory Behavior". In Breithaupt, T.; Thiel, M. (eds.).
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David Cowles; Jaclyn R. Van Dolson; Lisa R. Hainey; Dallas M. Dick (2006). "The use of different eye regions in the mantis shrimp
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has been observed to roll repeatedly for 2 m (6.6 ft), but specimens typically travel less than 1 m (3.3 ft).
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have been suggested to be early stomatopods, their fragmentary known remains make the referral uncertain. The oldest unambiguous
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C. H. Mazel; T. W. Cronin; R. L. Caldwell; N. J. Marshall (2004). "Fluorescent enhancement of signaling in a mantis shrimp".
3481:"Out of the blue: the evolution of horizontally polarized signals in Haptosquilla (Crustacea, Stomatopoda, Protosquillidae)" 3098: 557:
Mantis shrimp live in burrows where they spend the majority of their time. The spearing species build their habitat in soft
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Porter, Megan L.; Speiser, Daniel I.; Zaharoff, Alexander K.; Caldwell, Roy L.; Cronin, Thomas W.; Oakley, Todd H. (2013).
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insensitive to infrared light. The optical elements in these rows have eight different classes of visual pigments and the
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Templin, Rachel M.; How, Martin J.; Roberts, Nicholas W.; Chiou, Tsyr-Huei; Marshall, Justin (15 September 2017).
6271: 5277: 3341: 1628: 1513: 1468: 1122: 1052:, has been observed flipping itself into a crude wheel. The species lives in shallow, sandy areas. At low tides, 1048: 221: 3838: 1922: 5722: 5417: 5371: 1917: 1535: 1475: 1362: 1354: 728: 121: 1994: 1186:, named for its resemblance to a scrub brush. The shrimp can be steamed, boiled, grilled, or dried, used with 587:
Strike mechanics and spearing movement of the 2nd maxilliped (raptorial claw, ballistic claw) of mantis shrimp
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Mantis shrimp are long-lived and exhibit complex behaviour, such as ritualised fighting. Some species use
550: 394: 2224:"Extreme impact and cavitation forces of a biological hammer: strike forces of the peacock mantis shrimp" 1423: 6187: 5885: 5270: 5063: 4304: 4194: 4053: 1575: 1559: 1419: 1142: 921: 834: 790: 6205: 6026: 3711:
Caldwell, Roy L. (1979). "A unique form of locomotion in a stomatopod – backward somersaulting".
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The capacity to see UV light may enable observation of otherwise hard-to-detect prey on coral reefs.
899: 830: 258: 6228: 2769:"The molecular genetics and evolution of colour and polarization vision in stomatopod crustaceans". 2026: 5797: 5736: 5658: 5630: 5624: 5582: 5575: 5459: 5431: 5365: 5048: 5023: 4957: 4789: 4662: 4647: 4642: 4587: 4552: 4547: 4410: 4261: 4176: 1326: 1012: 1007:
individual smell. Many have developed complex social behaviours to defend their space from rivals.
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The appendages are being studied as a microscale analogue for new macroscale material structures.
5804: 5715: 5652: 5642: 5480: 5466: 5298: 5073: 4617: 4267: 4003: 3938:"Closing a major gap in mantis shrimp evolution - first fossils of Stomatopoda from the Triassic" 3738: 3662: 3614: 3583: 3196: 3160: 3090: 3074: 2883: 2790:"The Evolution of Complexity in the Visual Systems of Stomatopods: Insights from Transcriptomics" 2682: 2622: 2568: 2508: 2457: 2190: 1666: 1643: 1097: 722: 521: 501:
About 520 species of mantis shrimp have been discovered worldwide; all living species are in the
149: 4156: 3876:"The evolutionary history of Stomatopoda (Crustacea: Malacostraca) inferred from molecular data" 2355:"Filtering and polychromatic vision in mantis shrimps: themes in visible and ultraviolet vision" 1728:"The evolutionary history of Stomatopoda (Crustacea: Malacostraca) inferred from molecular data" 1407: 772: 6192: 3792: 1284: 540: 6174: 6047: 5839: 5550: 5536: 5487: 5473: 5438: 5291: 5284: 5162: 4962: 4906: 4857: 4837: 4814: 4652: 4627: 4622: 4607: 4528: 4423: 4130: 4076:
Hoplocarida: Stomatopoda fact sheet—Guide to the marine zooplankton of south eastern Australia
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Cronin, Thomas W.; Bok, Michael J.; Marshall, N. Justin; Caldwell, Roy L. (19 February 2014).
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about more specifics on species, better if combined with some list of fisheries. For example,
930: 904: 432: 304: 5986: 3781: 616:: A primitive appendage that only a few species have. This body plan is largely unresearched. 6215: 6117: 5610: 5596: 5589: 5543: 5452: 5377: 5359: 5167: 5157: 5124: 4988: 4896: 4754: 4739: 4734: 4365: 4090: 4034: 3995: 3945: 3905: 3887: 3730: 3713: 3606: 3567: 3502: 3492: 3419: 3411: 3322: 3305: 3275: 3265: 3224: 3214: 3142: 3066: 2997: 2927: 2867: 2850: 2801: 2734: 2718: 2666: 2606: 2558: 2550: 2492: 2449: 2374: 2366: 2316: 2308: 2237: 2174: 2157: 1749: 1739: 1553: 1529: 1349: 1289: 1230: 1137: 995: 963: 815: 760: 637: 289: 279: 264: 6179: 1418:(359-300 million years ago). Stem-group mantis shrimp are assigned to two major groups the 5810: 5773: 5749: 5696: 5617: 5445: 5403: 5345: 5323: 5312: 5305: 5246: 5172: 5152: 4827: 4452: 4370: 4097: 3799: 3788: 3669: 3125: 2961: 2910: 2475:
Cronin, Thomas W. (2001). "Sensory adaptation: Tunable colour vision in a mantis shrimp".
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Daly, Ilse M.; How, Martin J.; Partridge, Julian C.; Roberts, Nicholas W. (May 16, 2018).
1825: 1622: 1507: 1491: 1377: 975: 850: 470: 387: 331: 326: 321: 284: 247: 112: 1214: 849:(area of eye that absorbs light from a single direction) is divided into three different 3726: 3398:
Daly, Ilse M.; How, Martin J.; Partridge, Julian C.; Roberts, Nicholas W. (2018-05-16).
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Please expand the article to include this information. Further details may exist on the
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Close-up of a mantis shrimp showing the structure of the eyes. The three dark spots are
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Paul F. Clark & Frederick R. Schram (2009). "Raymond B. Manning: an appreciation".
3910: 3875: 3424: 3399: 3229: 3182: 3024: 2739: 2379: 2354: 2321: 2296: 1818: 1754: 1727: 1613: 1606: 1455: 1440: 1333:, some mantis shrimp have grown unusually large in the contaminated water of the Grand 1274: 645: 575: 361: 351: 274: 269: 252: 3682: 1726:
Van Der Wal, Cara; Ahyong, Shane T.; Ho, Simon Y. W.; Lo, Nathan (21 September 2017).
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Morrison, Jessica (23 January 2014). "Mantis shrimp's super colour vision debunked".
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New type of optical material discovered in the secret language of the mantis shrimp
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A large number of mantis shrimp species were first scientifically described by one
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The huge diversity seen in mantis shrimp photoreceptors likely comes from ancient
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appendages that are used to attack and kill prey either by spearing, stunning, or
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Mantis shrimps are also abundant along Vietnam's coast, known in Vietnamese as
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Van Der Wal, Cara; Ahyong, Shane T.; Ho, Simon Y.W.; Lo, Nathan (2017-09-21).
3610: 3147: 3120: 2932: 2906:"Biological Sunscreens Tune Polychromatic Ultraviolet Vision in Mantis Shrimp" 2905: 2610: 2554: 1813: 1569: 1411: 1195: 1038:
Although stomatopods typically display the standard types of movement seen in
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The live rock with mantis shrimp burrows is considered useful by some in the
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covers only the rear part of the head and the first four segments of the
440: 294: 85: 80: 65: 60: 50: 6039: 4110: 2461: 5340: 5108: 5103: 5098: 5028: 4852: 4847: 4832: 4799: 4794: 4709: 4676: 4672: 4592: 4433: 4416: 3892: 3507: 3497: 3480: 3270: 3253: 3078: 2722: 1981:"How the Mantis Shrimp Packs its Punch | Biomechanics in the Wild" 1744: 1338: 1258: 1164: 1130: 1043: 891: 846: 826: 741: 631: 467: 460: 452: 100: 75: 3999: 3053:"Parallel processing and image analysis in the eyes of mantis shrimps" 2707:"Colour vision in stomatopod crustaceans: more questions than answers" 2359:
Philosophical Transactions of the Royal Society B: Biological Sciences
2242: 2223: 732:, the species with the most monotonous natural photic environment. In 6104: 5784: 5779: 5767: 5424: 5239: 5225: 5188: 5093: 5003: 4998: 4993: 4891: 4744: 4719: 4632: 4567: 4542: 4381: 4230: 3734: 2563: 2496: 1912: 1820:
Extraordinary Animals: An Encyclopedia of Curious and Unusual Animals
1395: 1330: 1262: 1238: 1203: 1039: 978:
and photoreceptor activation comparison/barcode analysis is present.
858: 726:
species-specific. Cheroske et al. did not observe spectral tuning in
607: 486: 444: 424: 407: 176: 6091: 5942: 4497: 3070: 2178: 6078: 2526:
Cheroske, Alexander G.; Barber, Paul H.; Cronin, Thomas W. (2006).
6200: 5862: 5396: 5230: 4689: 4457: 3201: 2978:
Stephenson, 1967 (Crustacea: Stomatopoda) for detecting objects".
2871: 2266:"Mantis shrimp inspires next generation of ultra-strong materials" 1389: 1388: 1348: 1208: 1160: 1116: 989: 914: 866: 783: 771: 745: 698: 669: 582: 569: 539: 4080: 869:, greatly reducing the analytical requirements at higher levels. 5566: 2027:"How mantis shrimp evolved many shapes with same powerful punch" 948: 780:, indicating the ommatidia that are pointing towards the camera. 681: 665: 5946: 5192: 4501: 4190: 1650:
scanning of their surroundings for potential prey or threats.
3183:"The secret world of shrimps: polarisation vision at its best" 2904:
Michael Bok; Megan Porter; Allen Place; Thomas Cronin (2014).
2705:
Streets, Amy; England, Hayley; Marshall, Justin (2022-03-15).
2438:
Milius, Susan (2012). "Mantis shrimp flub color vision test".
1068: 636:(102,000 m/s or 335,000 ft/s) and speeds of 23  6065: 4172:
Dana Point Fish Company—Top and Bottom Views of Mantis Shrimp
982:
be able to produce big, bright, colourful polarised signals.
821:
Mantis shrimp can perceive wavelengths of light ranging from
3759:(in Vietnamese). Dinh dưỡng. October 1, 2009. Archived from 1950: 1948: 1946: 1944: 1942: 1940: 1361:
Some saltwater aquarists keep stomatopods in captivity. The
688:, which their barbed claws can more easily slice and snag. 718:
complex front-end for any visual system ever discovered.
626:
Both types strike by rapidly unfolding and swinging their
3533:(Press release). University of Bristol. 25 October 2009. 3342:"A crustacean eye that rivals the best optical equipment" 3121:"Circular polarization vision in a stomatopod crustacean" 1136:
The mantis shrimp is eaten by a variety of cultures. In
3404:
Proceedings of the Royal Society B: Biological Sciences
2301:
Proceedings of the Royal Society B: Biological Sciences
4111:
Caldwell Lab at the University of California, Berkeley
3531:"Mantis shrimps could show us the way to a better DVD" 2148:
S. N. Patek, W. L. Korff & R. L. Caldwell (2004).
4116:
Patek Lab at the University of Massachusetts, Amherst
4091:
Research on Stomatopods at the University of Maryland
2413:"Mantis shrimp have the world's best eyes – but why?" 1873:"Order Stomatopoda: Mantis shrimp or thumb splitters" 2964:. Latimes.com (2014-07-05). Retrieved on 2015-10-21. 1229:, the mantis shrimp is known as "urinating shrimp" ( 713:
The eyes of the mantis shrimp are mounted on mobile
5955: 5819: 5758: 5565: 5339: 5224: 5117: 4976: 4950: 4866: 4813: 4671: 4535: 4442: 4356: 4327: 4318: 4288: 4251: 3176: 3174: 2981:Journal of Experimental Marine Biology and Ecology 1876: 1817: 1365:is especially colourful and desired in the trade. 4466:(shrimp, prawns, crayfish, lobsters, crabs, etc.) 4021:Marshall, N. Justin; O'Carroll, David C. (2005). 3634:"Parallels between Shrimp and Human Color Vision" 3020:"Mantis shrimp use 'nature's sunblock' to see UV" 1688:An Updated Classification of the Recent Crustacea 1314:, and is cooked and eaten like any other shrimp. 2955:Mantis shrimp wear tinted shades to see UV light 1182:. In regions such as Nha Trang, they are called 810:in one eye). In other words, each eye possesses 4081:The Lurker's Guide to Stomatopods—mantis shrimp 1587: 943:During mating rituals, mantis shrimps actively 3013: 3011: 2143: 2141: 1581: 1501:, zebra mantis shrimp or striped mantis shrimp 1152: 620:Spike Smashers (hammers or primitive smashers) 5204: 4513: 4483:The three most speciose orders are marked in 4202: 4058:: CS1 maint: DOI inactive as of April 2024 ( 3683:"Sharing the job: monogamy and parental care" 3400:"Complex gaze stabilization in mantis shrimp" 2297:"Complex gaze stabilization in mantis shrimp" 1782:"Large shrimp thriving in Ala Wai Canal muck" 1685:Joel W. Martin & George E. Davis (2001). 861:seems to be processed into numerous parallel 610:, hence the common name of these crustaceans. 8: 3051:Cronin, Thomas W.; Marshall, Justin (2001). 2899: 2897: 2406: 2404: 2402: 2400: 2398: 2150:"Deadly strike mechanism of a mantis shrimp" 2059:"Roy's List of Stomatopods for the Aquarium" 1696:Natural History Museum of Los Angeles County 1094:are commonly mentioned in SE Asian contexts. 508:, which arose around 250 million years ago. 5943: 5211: 5197: 5189: 4520: 4506: 4498: 4324: 4209: 4195: 4187: 3377:""Weird beastie" shrimp have super-vision" 2642: 2640: 2638: 2636: 1879:Seashore Life of Florida and the Caribbean 600:to cut prey while the mantis shrimp swims. 130: 111: 31: 3949: 3909: 3891: 3506: 3496: 3423: 3281:1983/1f1c982f-9a88-4184-b59a-2cebd73ec818 3279: 3269: 3228: 3218: 3200: 3181:Kleinlogel, Sonja; White, Andrew (2009). 3146: 2931: 2805: 2738: 2562: 2411:Franklin, Amanda M. (September 4, 2013). 2378: 2320: 2241: 2222:S. N. Patek & R. L. Caldwell (2005). 1753: 1743: 1399:, a primitive Carboniferous mantis shrimp 2348: 2346: 2344: 2342: 2340: 1775: 1773: 1253: 3782:A Load of Learnin' About Mantis Shrimps 3757:"Tôm tít – Đặc sản miền sông nước" 3672:. Bristol University (17 February 2016) 2764: 2762: 2760: 2758: 1677: 1278:is a common seafood, especially on the 4086:Mantis shrimp—colourful and aggressive 4051: 3839:"Shrimp spring into shattering action" 3632:Macknik, Stephen L. (March 20, 2014). 1794:from the original on November 11, 2020 1329:are abundant and are eaten boiled. In 3931: 3929: 3869: 3867: 3865: 2700: 2698: 2696: 2053: 2051: 2021: 2019: 1957:Chemical Communication in Crustaceans 919:Close-up of the trinocular vision of 911:Suggested advantages of visual system 7: 6206:F8A80A5A-D82D-5F30-B9E6-4C946A3F55CD 6167:846c3eb1-de28-4560-9d50-722c1ee770ac 3837:April Holladay (September 1, 2006). 3537:from the original on 31 October 2020 3460:from the original on 6 February 2017 1925:from the original on October 2, 2022 1721: 1719: 929:in the eye and could easily lead to 664:Smashers use this ability to attack 435:off from other members of the class 4487:; obelisks (†) mark extinct orders. 4106:. The Aquarium Wiki. April 7, 2006. 4027:Integrative and Comparative Biology 3258:The Journal of Experimental Biology 2794:Integrative and Comparative Biology 1995:"Why are Mantis Shrimp so Awesome?" 802:is made up of tens of thousands of 532:between eastern Africa and Hawaii. 4023:"Visual Ecology of Mantis Shrimps" 3687:University of California, Berkeley 3356:from the original on April 5, 2012 3032:from the original on 25 April 2023 1883:. Dover pictorial archive series. 1780:James Gonser (February 15, 2003). 1373:than the fishkeeper would prefer. 25: 3644:from the original on May 25, 2020 2419:from the original on July 5, 2018 2276:from the original on May 24, 2021 1213:Drying mantis shrimp at Gò Công, 1163:topping, and occasionally raw as 721:Compared with the three types of 579:, showing the spearing appendages 4143:Cressey, Daniel (May 14, 2008). 3638:Scientific American Blog Network 1959:. Chile: Springer. p. 219. 1660: 1303:, the mantis shrimp is known as 1073: 153: 3966:from the original on 2023-06-06 3851:from the original on 2012-06-30 3693:from the original on 2009-09-14 3485:Journal of Experimental Biology 3340:Lee, Chris (November 1, 2009). 3101:from the original on 2020-06-19 2831:Journal of Experimental Biology 2711:Journal of Experimental Biology 2579:from the original on 2024-01-04 2229:Journal of Experimental Biology 2127:from the original on 2023-04-18 2098:from the original on 2022-01-31 2069:from the original on 2022-08-23 2037:from the original on 2022-07-21 2005:from the original on 2022-08-10 661:can be enough to stun or kill. 3018:DuRant, Hassan (3 July 2014). 1999:California Academy of Sciences 1243: 994:Drawing of a mantis shrimp by 951:; the ability to perceive the 1: 4603:Pork knuckles and ginger stew 4126:"The superior eyes of shrimp" 3987:Journal of Crustacean Biology 3375:Minard, Anne (May 19, 2008). 1698:. p. 132. Archived from 1272:countries, the mantis shrimp 3220:10.1371/journal.pone.0002190 1140:, the mantis shrimp species 5828:Austromegabalanus psittacus 4310:Stomatopoda (mantis shrimp) 3381:National Geographic Society 3002:10.1016/j.jembe.2005.09.016 1588: 1015:, long-term relationships. 431: 'foot'). Stomatopods 6288: 5893:Paralithodes camtschaticus 5178:History of Chinese cuisine 2976:Hemisquilla californiensis 1885:Courier Dover Publications 1547:, pink-eared mantis shrimp 1544:Odontodactylus latirostris 1483:Hemisquilla californiensis 1414:mantis shrimp date to the 709:, showing the stalked eyes 546:Odontodactylus latirostris 417: 414: 'mouth' and 400: 238:Superfamilies and families 5931: 5278:Macrobrachium rosenbergii 5143: 4478: 4225: 3791:, by James Fatherree, in 3611:10.1038/nature.2014.14578 3148:10.1016/j.cub.2008.02.066 2933:10.1016/j.cub.2014.05.071 2611:10.1017/S0952523809090129 2555:10.1007/s00227-006-0313-5 1629:Heterosquilla tricarinata 1582: 1514:Nannosquilla decemspinosa 1469:Hemisquilla australiensis 1234: 1153: 1049:Nannosquilla decemspinosa 729:Neogonodactylus oerstedii 234: 229: 150:Scientific classification 148: 138: 129: 119: 110: 34: 5723:Portunus trituberculatus 5418:Metanephrops challengeri 5372:California spiny lobster 3951:10.3140/bull.geosci.1864 3327:10.1038/nphoton.2009.189 1918:Tree of Life Web Project 1871:Gilbert L. Voss (2002). 1536:Odontodactylus scyllarus 1476:Hemisquilla braziliensis 1121:Mantis shrimp caught at 839:linearly polarised light 466:Called "sea locusts" by 122:Odontodactylus scyllarus 5874:Lysiosquillina maculata 4942:White sugar sponge cake 4434:Cumacea (hooded shrimp) 3942:Bulletin of Geosciences 3572:10.1126/science.1089803 3058:The Biological Bulletin 2671:10.1126/science.1245824 2454:10.1002/scin.5591820609 1787:The Honolulu Advertiser 1539:, peacock mantis shrimp 1498:Lysiosquillina maculata 1321:, mantis shrimp called 1159:, is eaten boiled as a 972:Haptosquilla trispinosa 706:Lysiosquillina maculata 141:Lysiosquillina maculata 6262:Commercial crustaceans 5704:Ovalipes australiensis 5530:Scyllarides herklotsii 5502:Panulirus penicillatus 5391:Japanese spiny lobster 5254:Cryphiops caementarius 4558:Cantonese seafood soup 4145:"Shrimp's super sight" 4138:on September 12, 2009. 3416:10.1098/rspb.2018.0594 2371:10.1098/rstb.2013.0032 2313:10.1098/rspb.2018.0594 1667:Crustaceans portal 1563:, common mantis shrimp 1400: 1358: 1305:tatampal, hipong-dapa, 1222: 1133: 1084:is missing information 999: 925: 886:formed from stacks of 795: 781: 744:binding pocket of the 710: 588: 580: 554: 551:Wakatobi National Park 144:(Zebra Mantis Shrimp) 6188:Paleobiology Database 5886:Oratosquilla oratoria 5271:Litopenaeus setiferus 5024:Garland chrysanthemum 5004:Fermented black beans 4951:Condiments and spices 4159:on September 22, 2020 4041:(inactive 2024-04-02) 3816:. London: Andromeda. 2771:Ophthalmic Physiology 1576:Oratosquilla oratoria 1560:Pseudosquilla ciliata 1392: 1355:Peacock mantis shrimp 1352: 1212: 1143:Oratosquilla oratoria 1120: 993: 922:Pseudosquilla ciliata 918: 791:Oratosquilla oratoria 787: 775: 702: 586: 573: 543: 473:, "prawn killers" in 5916:Tasmanian giant crab 5671:Hypothalassia acerba 5557:Tristan rock lobster 5516:Parribacus japonicus 5509:Panulirus versicolor 5495:Panulirus pascuensis 5330:Xiphopenaeus kroyeri 5148:Cantonese restaurant 5130:Dried shredded squid 4578:Dragon tiger phoenix 4573:Crispy fried chicken 4563:Chinese steamed eggs 4458:Euphausiacea (krill) 4039:10.1093/icb/45.4.734 2415:. The Conversation. 1860:on January 21, 2021. 1600:Rissoides desmaresti 1521:Platysquilla eusebia 1462:Hemisquilla ensigera 1447:Gonodactylus smithii 1385:Evolutionary history 1092:Oratosquilla anomala 1088:Harpiosquilla harpax 1025:and some species in 900:Gonodactylus smithii 125:(Red Mantis Shrimp) 41:Carboniferous–Recent 27:Order of crustaceans 5798:Procambarus clarkii 5737:Scylla paramamosain 5659:Gecarcinus ruricola 5631:Chinese mitten crab 5625:Chaceon quinquedens 5583:Callinectes similis 5576:Callinectes sapidus 5460:Panulirus echinatus 5432:Nephrops norvegicus 5366:Arctides guineensis 4958:Fermented bean curd 4790:Water chestnut cake 4663:Yangzhou fried rice 4648:White boiled shrimp 4643:Sweet and sour pork 4588:Eight treasure duck 3814:The Marine Aquarium 3812:Nick Dakin (2004). 3727:1979Natur.282...71C 3319:2009NaPho...3..641R 3211:2008PLoSO...3.2190K 3139:2008CBio...18..429C 2994:2006JEMBE.330..528C 2924:2014CBio...24.1636B 2864:1999Natur.401..873M 2663:2014Sci...343..411T 2599:Visual Neuroscience 2547:2006MarBi.150..213C 2489:2001Natur.411..547C 2171:2004Natur.428..819P 1921:. January 1, 2002. 1424:Archaeostomatopodea 300:Erythrosquilloidea 6267:Edible crustaceans 5805:Orconectes virilis 5716:Portunus pelagicus 5653:Florida stone crab 5643:Declawing of crabs 5481:Panulirus longipes 5467:Panulirus guttatus 5299:Penaeus esculentus 5219:Edible crustaceans 5074:Shrimp roe noodles 4618:Snake bite chicken 4305:Archaeostomatopoda 4096:2010-05-30 at the 3893:10.7717/peerj.3844 3798:2014-02-19 at the 3787:2011-07-15 at the 3763:on August 16, 2012 3668:2016-03-07 at the 3498:10.1242/jeb.107581 3410:(1878): 20180594. 3271:10.1242/jeb.162941 2960:2014-11-22 at the 2807:10.1093/icb/ict060 2723:10.1242/jeb.243699 2365:(1636): 20130032. 2307:(1878): 20180594. 2117:"h_californiensis" 1745:10.7717/peerj.3844 1644:Raymond B. Manning 1420:Palaeostomatopodea 1401: 1393:Reconstruction of 1359: 1223: 1134: 1000: 926: 829:(720 nm) and 796: 782: 723:photoreceptor cell 711: 589: 581: 555: 6257:Cantonese cuisine 6239: 6238: 6175:Open Tree of Life 5949:Taxon identifiers 5940: 5939: 5840:Galathea strigosa 5551:Thymops birsteini 5537:Scyllarides latus 5488:Panulirus ornatus 5474:Panulirus homarus 5439:Palinurus elephas 5425:Thenus orientalis 5292:Pandalus borealis 5285:Palaemon serratus 5186: 5185: 5163:Hong Kong cuisine 4963:Five-spice powder 4907:Malay sponge cake 4858:White cut chicken 4838:Orange cuttlefish 4653:White cut chicken 4628:Steam minced pork 4623:Soy sauce chicken 4608:Seafood birdsnest 4529:Cantonese cuisine 4495: 4494: 4488: 4474: 4473: 4424:Pygocephalomorpha 4000:10.1651/09-3158.1 3823:978-1-902389-67-7 3491:(19): 3425–3431. 3264:(18): 3222–3230. 2858:(6756): 873–874. 2657:(6169): 411–413. 2243:10.1242/jeb.01831 2236:(19): 3655–3664. 2165:(6985): 819–820. 2121:ucmp.berkeley.edu 2092:ucmp.berkeley.edu 2063:ucmp.berkeley.edu 1898:978-0-486-42068-4 1854:Queensland Museum 1835:978-0-313-33922-6 1227:Cantonese cuisine 1115: 1114: 953:phase of the moon 931:natural selection 905:Stokes parameters 894:Disc technology. 865:leading into the 825:(300 nm) to 812:trinocular vision 451:in many shallow, 371: 370: 305:Erythrosquillidae 243:Bathysquilloidea 239: 225: 16:(Redirected from 6279: 6272:Japanese seafood 6232: 6231: 6219: 6218: 6209: 6208: 6196: 6195: 6183: 6182: 6170: 6169: 6160: 6159: 6147: 6146: 6144:NHMSYS0021050823 6134: 6133: 6121: 6120: 6108: 6107: 6095: 6094: 6082: 6081: 6069: 6068: 6056: 6055: 6043: 6042: 6030: 6029: 6017: 6016: 6004: 6003: 5991: 5990: 5989: 5976: 5975: 5974: 5944: 5611:Cancer productus 5597:Cancer bellianus 5590:Cancer irroratus 5544:Scyllarus arctus 5453:Panulirus cygnus 5378:Homarus gammarus 5360:American lobster 5213: 5206: 5199: 5190: 5168:Macanese cuisine 5158:Shanghai cuisine 5125:Chinese herb tea 4989:Black bean paste 4897:Ginger milk curd 4755:Steamed meatball 4740:Rice noodle roll 4735:Ox-tongue pastry 4553:Buddha's delight 4548:Bird's nest soup 4522: 4515: 4508: 4499: 4482: 4385:(opossum shrimp) 4366:Spelaeogriphacea 4325: 4211: 4204: 4197: 4188: 4168: 4166: 4164: 4155:. Archived from 4149:The Great Beyond 4139: 4134:. Archived from 4124:(May 24, 2008). 4107: 4064: 4063: 4057: 4049: 4047: 4046: 4018: 4012: 4011: 3981: 3975: 3974: 3972: 3971: 3953: 3933: 3924: 3923: 3913: 3895: 3871: 3860: 3859: 3857: 3856: 3834: 3828: 3827: 3809: 3803: 3802:online magazine. 3779: 3773: 3772: 3770: 3768: 3753: 3747: 3746: 3735:10.1038/282071a0 3708: 3702: 3701: 3699: 3698: 3679: 3673: 3660: 3654: 3653: 3651: 3649: 3629: 3623: 3622: 3598: 3592: 3591: 3553: 3547: 3546: 3544: 3542: 3527: 3521: 3520: 3510: 3500: 3476: 3470: 3469: 3467: 3465: 3456:. 14 July 2016. 3444: 3438: 3437: 3427: 3395: 3389: 3388: 3387:on May 27, 2008. 3383:. 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Archived from 1846: 1840: 1839: 1823: 1810: 1804: 1803: 1801: 1799: 1777: 1768: 1767: 1757: 1747: 1723: 1714: 1713: 1711: 1710: 1704: 1693: 1682: 1665: 1664: 1595: 1593: 1585: 1584: 1554:Pseudosquillidae 1530:Odontodactylidae 1255: 1245: 1236: 1158: 1156: 1155: 1138:Japanese cuisine 1110: 1107: 1101: 1077: 1069: 996:Richard Lydekker 964:opponent process 851:pigmented layers 823:deep ultraviolet 816:depth perception 814:, and therefore 761:gene duplication 428: 421: 411: 404: 357:Parasquilloidea 347:Eurysquilloidea 311:Lysiosquilloidea 290:Pseudosquillidae 280:Odontodactylidae 265:Alainosquillidae 237: 220: 158: 157: 134: 115: 105: 42: 38:Temporal range: 32: 21: 6287: 6286: 6282: 6281: 6280: 6278: 6277: 6276: 6242: 6241: 6240: 6235: 6227: 6222: 6214: 6212: 6204: 6199: 6191: 6186: 6178: 6173: 6165: 6163: 6155: 6150: 6142: 6137: 6129: 6124: 6116: 6111: 6103: 6098: 6090: 6085: 6077: 6072: 6064: 6059: 6051: 6046: 6038: 6033: 6025: 6020: 6012: 6007: 5999: 5994: 5985: 5984: 5979: 5970: 5969: 5964: 5951: 5941: 5936: 5927: 5815: 5811:Signal crayfish 5774:Astacus astacus 5754: 5750:Soft-shell crab 5697:Orithyia sinica 5618:Chaceon fenneri 5561: 5446:Panulirus argus 5404:Jasus edwardsii 5348: 5343: 5335: 5324:Whiteleg shrimp 5313:Processa edulis 5306:Penaeus monodon 5247:Crangon crangon 5229: 5220: 5217: 5187: 5182: 5173:Chinese cuisine 5153:Beijing cuisine 5139: 5113: 4972: 4946: 4862: 4828:Chinese sausage 4809: 4667: 4531: 4526: 4496: 4491: 4470: 4438: 4371:Thermosbaenacea 4352: 4348:Palaeocaridacea 4314: 4284: 4247: 4221: 4215: 4182:Deep Look (PBS) 4162: 4160: 4142: 4120: 4104:"Mantis shrimp" 4102: 4098:Wayback Machine 4072: 4067: 4050: 4044: 4042: 4020: 4019: 4015: 3983: 3982: 3978: 3969: 3967: 3935: 3934: 3927: 3873: 3872: 3863: 3854: 3852: 3836: 3835: 3831: 3824: 3811: 3810: 3806: 3800:Wayback Machine 3789:Wayback Machine 3780: 3776: 3766: 3764: 3755: 3754: 3750: 3721:(5734): 71–73. 3710: 3709: 3705: 3696: 3694: 3681: 3680: 3676: 3670:Wayback Machine 3661: 3657: 3647: 3645: 3631: 3630: 3626: 3600: 3599: 3595: 3555: 3554: 3550: 3540: 3538: 3529: 3528: 3524: 3478: 3477: 3473: 3463: 3461: 3446: 3445: 3441: 3397: 3396: 3392: 3374: 3373: 3369: 3359: 3357: 3339: 3338: 3334: 3313:(11): 641–644. 3302: 3301: 3297: 3251: 3250: 3246: 3180: 3179: 3172: 3126:Current Biology 3118: 3117: 3113: 3104: 3102: 3071:10.2307/1543312 3050: 3049: 3045: 3035: 3033: 3017: 3016: 3009: 2973: 2972: 2968: 2962:Wayback Machine 2953: 2949: 2918:(14): 1636–42. 2911:Current Biology 2903: 2902: 2895: 2847: 2846: 2842: 2828: 2827: 2823: 2787: 2786: 2782: 2768: 2767: 2756: 2704: 2703: 2694: 2646: 2645: 2634: 2596: 2595: 2591: 2582: 2580: 2530: 2525: 2524: 2520: 2483:(6837): 547–8. 2474: 2473: 2469: 2437: 2436: 2432: 2422: 2420: 2410: 2409: 2396: 2352: 2351: 2338: 2294: 2293: 2289: 2279: 2277: 2264: 2263: 2259: 2221: 2220: 2216: 2207: 2205: 2201: 2179:10.1038/428819a 2152: 2147: 2146: 2139: 2130: 2128: 2115: 2114: 2110: 2101: 2099: 2086: 2085: 2081: 2072: 2070: 2057: 2056: 2049: 2040: 2038: 2025: 2024: 2017: 2008: 2006: 1993: 1992: 1988: 1979: 1978: 1974: 1967: 1954: 1953: 1938: 1928: 1926: 1911: 1910: 1906: 1899: 1870: 1869: 1865: 1850:"Mantis shrimp" 1848: 1847: 1843: 1836: 1826:Greenwood Press 1812: 1811: 1807: 1797: 1795: 1779: 1778: 1771: 1725: 1724: 1717: 1708: 1706: 1702: 1691: 1684: 1683: 1679: 1675: 1659: 1656: 1623:Tetrasquillidae 1579: 1508:Nannosquillidae 1492:Lysiosquillidae 1436: 1434:Example species 1387: 1378:marine aquarium 1357: 1347: 1312:alupihang-dagat 1254:laai6 niu6 haa1 1150: 1111: 1105: 1102: 1095: 1078: 1067: 1054:N. decemspinosa 1046:, one species, 988: 976:color opponency 913: 879: 877:Polarized light 831:polarised light 770: 697: 608:praying mantids 568: 538: 499: 367: 366: 332:Tetrasquillidae 327:Nannosquillidae 322:Lysiosquillidae 285:Protosquillidae 259:Gonodactyloidea 248:Bathysquillidae 240: 219: 152: 106: 104: 103: 98: 93: 88: 83: 78: 73: 68: 63: 58: 53: 48: 40: 39: 36: 28: 23: 22: 15: 12: 11: 5: 6285: 6283: 6275: 6274: 6269: 6264: 6259: 6254: 6244: 6243: 6237: 6236: 6234: 6233: 6220: 6210: 6197: 6184: 6171: 6161: 6148: 6135: 6122: 6109: 6096: 6083: 6070: 6057: 6044: 6031: 6018: 6005: 5992: 5977: 5961: 5959: 5953: 5952: 5947: 5938: 5937: 5932: 5929: 5928: 5926: 5925: 5918: 5913: 5910:Squilla mantis 5906: 5901: 5896: 5889: 5882: 5877: 5870: 5865: 5860: 5855: 5853:Goose barnacle 5850: 5847:Glyptolithodes 5843: 5836: 5831: 5823: 5821: 5817: 5816: 5814: 5813: 5808: 5801: 5794: 5787: 5782: 5777: 5770: 5764: 5762: 5756: 5755: 5753: 5752: 5747: 5744:Scylla serrata 5740: 5733: 5726: 5719: 5712: 5710:Pie crust crab 5707: 5700: 5693: 5686: 5679: 5674: 5667: 5665:Horsehair crab 5662: 5655: 5650: 5648:Dungeness crab 5645: 5640: 5633: 5628: 5621: 5614: 5607: 5604:Cancer pagurus 5600: 5593: 5586: 5579: 5571: 5569: 5563: 5562: 5560: 5559: 5554: 5547: 5540: 5533: 5526: 5519: 5512: 5505: 5498: 5491: 5484: 5477: 5470: 5463: 5456: 5449: 5442: 5435: 5428: 5421: 5414: 5411:Jasus lalandii 5407: 5400: 5393: 5388: 5385:Ibacus peronii 5381: 5374: 5369: 5362: 5356: 5354: 5337: 5336: 5334: 5333: 5326: 5321: 5316: 5309: 5302: 5295: 5288: 5281: 5274: 5267: 5262: 5257: 5250: 5243: 5235: 5233: 5222: 5221: 5218: 5216: 5215: 5208: 5201: 5193: 5184: 5183: 5181: 5180: 5175: 5170: 5165: 5160: 5155: 5150: 5144: 5141: 5140: 5138: 5137: 5132: 5127: 5121: 5119: 5115: 5114: 5112: 5111: 5106: 5101: 5096: 5091: 5086: 5081: 5076: 5071: 5066: 5061: 5056: 5051: 5046: 5044:Pig blood curd 5041: 5036: 5031: 5026: 5021: 5016: 5011: 5006: 5001: 4996: 4991: 4986: 4980: 4978: 4974: 4973: 4971: 4970: 4965: 4960: 4954: 4952: 4948: 4947: 4945: 4944: 4939: 4934: 4929: 4924: 4919: 4914: 4909: 4904: 4899: 4894: 4889: 4887:Deuk Deuk Tong 4884: 4879: 4877:Almond biscuit 4873: 4871: 4864: 4863: 4861: 4860: 4855: 4850: 4845: 4840: 4835: 4830: 4825: 4819: 4817: 4811: 4810: 4808: 4807: 4802: 4797: 4792: 4787: 4782: 4780:Tofu skin roll 4777: 4772: 4767: 4762: 4757: 4752: 4747: 4742: 4737: 4732: 4727: 4722: 4717: 4712: 4707: 4702: 4697: 4692: 4687: 4681: 4679: 4669: 4668: 4666: 4665: 4660: 4658:Wonton noodles 4655: 4650: 4645: 4640: 4635: 4630: 4625: 4620: 4615: 4613:Shark fin soup 4610: 4605: 4600: 4595: 4590: 4585: 4580: 4575: 4570: 4565: 4560: 4555: 4550: 4545: 4539: 4537: 4533: 4532: 4527: 4525: 4524: 4517: 4510: 4502: 4493: 4492: 4490: 4489: 4479: 4476: 4475: 4472: 4471: 4469: 4468: 4460: 4455: 4453:Angustidontida 4448: 4446: 4440: 4439: 4437: 4436: 4431: 4426: 4420: 4413: 4408: 4401: 4396: 4391: 4386: 4378: 4373: 4368: 4362: 4360: 4354: 4353: 4351: 4350: 4344: 4339: 4333: 4331: 4322: 4320:Eumalacostraca 4316: 4315: 4313: 4312: 4307: 4301: 4299:Aeschronectida 4294: 4292: 4286: 4285: 4283: 4282: 4275: 4270: 4264: 4257: 4255: 4249: 4248: 4246: 4245: 4239: 4233: 4226: 4223: 4222: 4216: 4214: 4213: 4206: 4199: 4191: 4185: 4184: 4179: 4174: 4169: 4140: 4118: 4113: 4108: 4100: 4088: 4083: 4078: 4071: 4070:External links 4068: 4066: 4065: 4033:(4): 734–742. 4013: 3994:(4): 431–457. 3976: 3925: 3861: 3829: 3822: 3804: 3774: 3748: 3703: 3674: 3655: 3624: 3593: 3548: 3522: 3471: 3439: 3390: 3367: 3332: 3295: 3244: 3170: 3133:(6): 429–434. 3111: 3065:(2): 177–183. 3043: 3025:sciencemag.org 3007: 2988:(2): 528–534. 2966: 2947: 2893: 2840: 2821: 2780: 2754: 2692: 2632: 2605:(3): 255–265. 2589: 2541:(2): 213–220. 2535:Marine Biology 2518: 2467: 2430: 2394: 2336: 2287: 2257: 2214: 2137: 2108: 2079: 2047: 2015: 1986: 1972: 1965: 1936: 1904: 1897: 1863: 1841: 1834: 1805: 1769: 1715: 1676: 1674: 1671: 1670: 1669: 1655: 1652: 1636: 1635: 1634: 1633: 1619: 1618: 1617: 1614:Squilla mantis 1610: 1607:Squilla empusa 1603: 1596: 1566: 1565: 1564: 1550: 1549: 1548: 1540: 1526: 1525: 1524: 1517: 1504: 1503: 1502: 1488: 1487: 1486: 1479: 1472: 1465: 1456:Hemisquillidae 1452: 1451: 1450: 1441:Gonodactylidae 1435: 1432: 1386: 1383: 1363:peacock mantis 1353: 1346: 1343: 1275:Squilla mantis 1113: 1112: 1081: 1079: 1072: 1066: 1063: 987: 984: 912: 909: 878: 875: 769: 766: 696: 693: 624: 623: 617: 611: 601: 576:Squilla mantis 567: 564: 537: 534: 530:Pacific Oceans 498: 495: 369: 368: 365: 364: 362:Parasquillidae 355: 354: 352:Eurysquillidae 345: 344: 335: 334: 329: 324: 319: 308: 307: 298: 297: 292: 287: 282: 277: 275:Gonodactylidae 272: 270:Hemisquillidae 267: 256: 255: 253:Indosquillidae 250: 241: 236: 235: 232: 231: 227: 226: 214: 210: 209: 204: 200: 199: 194: 190: 189: 184: 180: 179: 174: 170: 169: 164: 160: 159: 146: 145: 136: 135: 127: 126: 117: 116: 108: 107: 99: 94: 89: 84: 79: 74: 69: 64: 59: 54: 49: 44: 43: 37: 26: 24: 18:Mantis shrimps 14: 13: 10: 9: 6: 4: 3: 2: 6284: 6273: 6270: 6268: 6265: 6263: 6260: 6258: 6255: 6253: 6250: 6249: 6247: 6230: 6225: 6221: 6217: 6211: 6207: 6202: 6198: 6194: 6189: 6185: 6181: 6176: 6172: 6168: 6162: 6158: 6153: 6149: 6145: 6140: 6136: 6132: 6127: 6123: 6119: 6114: 6110: 6106: 6101: 6097: 6093: 6088: 6084: 6080: 6075: 6071: 6067: 6062: 6058: 6054: 6049: 6045: 6041: 6036: 6032: 6028: 6023: 6019: 6015: 6010: 6006: 6002: 5997: 5993: 5988: 5982: 5978: 5973: 5967: 5963: 5962: 5960: 5958: 5954: 5950: 5945: 5935: 5930: 5924: 5923: 5919: 5917: 5914: 5912: 5911: 5907: 5905: 5904:Squat lobster 5902: 5900: 5899:Red king crab 5897: 5895: 5894: 5890: 5888: 5887: 5883: 5881: 5880:Mantis shrimp 5878: 5876: 5875: 5871: 5869: 5866: 5864: 5861: 5859: 5856: 5854: 5851: 5849: 5848: 5844: 5842: 5841: 5837: 5835: 5832: 5830: 5829: 5825: 5824: 5822: 5818: 5812: 5809: 5807: 5806: 5802: 5800: 5799: 5795: 5793: 5792: 5788: 5786: 5783: 5781: 5778: 5776: 5775: 5771: 5769: 5766: 5765: 5763: 5761: 5757: 5751: 5748: 5746: 5745: 5741: 5739: 5738: 5734: 5732: 5731: 5730:Ranina ranina 5727: 5725: 5724: 5720: 5718: 5717: 5713: 5711: 5708: 5706: 5705: 5701: 5699: 5698: 5694: 5692: 5691: 5690:Menippe adina 5687: 5685: 5684: 5683:Maja squinado 5680: 5678: 5675: 5673: 5672: 5668: 5666: 5663: 5661: 5660: 5656: 5654: 5651: 5649: 5646: 5644: 5641: 5639: 5638: 5634: 5632: 5629: 5627: 5626: 5622: 5620: 5619: 5615: 5613: 5612: 5608: 5606: 5605: 5601: 5599: 5598: 5594: 5592: 5591: 5587: 5585: 5584: 5580: 5578: 5577: 5573: 5572: 5570: 5568: 5564: 5558: 5555: 5553: 5552: 5548: 5546: 5545: 5541: 5539: 5538: 5534: 5532: 5531: 5527: 5525: 5524: 5520: 5518: 5517: 5513: 5511: 5510: 5506: 5504: 5503: 5499: 5497: 5496: 5492: 5490: 5489: 5485: 5483: 5482: 5478: 5476: 5475: 5471: 5469: 5468: 5464: 5462: 5461: 5457: 5455: 5454: 5450: 5448: 5447: 5443: 5441: 5440: 5436: 5434: 5433: 5429: 5427: 5426: 5422: 5420: 5419: 5415: 5413: 5412: 5408: 5406: 5405: 5401: 5399: 5398: 5394: 5392: 5389: 5387: 5386: 5382: 5380: 5379: 5375: 5373: 5370: 5368: 5367: 5363: 5361: 5358: 5357: 5355: 5352: 5347: 5342: 5338: 5332: 5331: 5327: 5325: 5322: 5320: 5317: 5315: 5314: 5310: 5308: 5307: 5303: 5301: 5300: 5296: 5294: 5293: 5289: 5287: 5286: 5282: 5280: 5279: 5275: 5273: 5272: 5268: 5266: 5263: 5261: 5258: 5256: 5255: 5251: 5249: 5248: 5244: 5242: 5241: 5237: 5236: 5234: 5232: 5227: 5223: 5214: 5209: 5207: 5202: 5200: 5195: 5194: 5191: 5179: 5176: 5174: 5171: 5169: 5166: 5164: 5161: 5159: 5156: 5154: 5151: 5149: 5146: 5145: 5142: 5136: 5133: 5131: 5128: 5126: 5123: 5122: 5120: 5116: 5110: 5107: 5105: 5102: 5100: 5097: 5095: 5092: 5090: 5087: 5085: 5082: 5080: 5077: 5075: 5072: 5070: 5067: 5065: 5064:Sea cucumbers 5062: 5060: 5057: 5055: 5052: 5050: 5047: 5045: 5042: 5040: 5037: 5035: 5034:Mantis shrimp 5032: 5030: 5027: 5025: 5022: 5020: 5017: 5015: 5012: 5010: 5007: 5005: 5002: 5000: 4997: 4995: 4992: 4990: 4987: 4985: 4982: 4981: 4979: 4975: 4969: 4966: 4964: 4961: 4959: 4956: 4955: 4953: 4949: 4943: 4940: 4938: 4935: 4933: 4932:Tang bu shuai 4930: 4928: 4925: 4923: 4922:Red bean soup 4920: 4918: 4917:Red bean cake 4915: 4913: 4910: 4908: 4905: 4903: 4900: 4898: 4895: 4893: 4890: 4888: 4885: 4883: 4880: 4878: 4875: 4874: 4872: 4870: 4867:Desserts and 4865: 4859: 4856: 4854: 4851: 4849: 4846: 4844: 4841: 4839: 4836: 4834: 4831: 4829: 4826: 4824: 4821: 4820: 4818: 4816: 4812: 4806: 4803: 4801: 4798: 4796: 4793: 4791: 4788: 4786: 4783: 4781: 4778: 4776: 4773: 4771: 4770:Taro dumpling 4768: 4766: 4763: 4761: 4758: 4756: 4753: 4751: 4748: 4746: 4743: 4741: 4738: 4736: 4733: 4731: 4728: 4726: 4723: 4721: 4718: 4716: 4713: 4711: 4708: 4706: 4703: 4701: 4698: 4696: 4693: 4691: 4688: 4686: 4683: 4682: 4680: 4678: 4674: 4670: 4664: 4661: 4659: 4656: 4654: 4651: 4649: 4646: 4644: 4641: 4639: 4636: 4634: 4631: 4629: 4626: 4624: 4621: 4619: 4616: 4614: 4611: 4609: 4606: 4604: 4601: 4599: 4598:Lemon chicken 4596: 4594: 4591: 4589: 4586: 4584: 4583:Egg foo young 4581: 4579: 4576: 4574: 4571: 4569: 4566: 4564: 4561: 4559: 4556: 4554: 4551: 4549: 4546: 4544: 4541: 4540: 4538: 4534: 4530: 4523: 4518: 4516: 4511: 4509: 4504: 4503: 4500: 4486: 4481: 4480: 4477: 4467: 4465: 4461: 4459: 4456: 4454: 4450: 4449: 4447: 4445: 4441: 4435: 4432: 4430: 4427: 4425: 4421: 4419: 4418: 4414: 4412: 4411:Ingolfiellida 4409: 4407: 4406: 4402: 4400: 4397: 4395: 4392: 4390: 4387: 4384: 4383: 4379: 4377: 4376:Lophogastrida 4374: 4372: 4369: 4367: 4364: 4363: 4361: 4359: 4355: 4349: 4345: 4343: 4342:Bathynellacea 4340: 4338: 4335: 4334: 4332: 4330: 4326: 4323: 4321: 4317: 4311: 4308: 4306: 4302: 4300: 4296: 4295: 4293: 4291: 4287: 4280: 4276: 4274: 4271: 4269: 4265: 4263: 4262:Archaeostraca 4259: 4258: 4256: 4254: 4250: 4244: 4240: 4238: 4234: 4232: 4228: 4227: 4224: 4220: 4212: 4207: 4205: 4200: 4198: 4193: 4192: 4189: 4183: 4180: 4178: 4175: 4173: 4170: 4158: 4154: 4150: 4146: 4141: 4137: 4133: 4132: 4127: 4123: 4119: 4117: 4114: 4112: 4109: 4105: 4101: 4099: 4095: 4092: 4089: 4087: 4084: 4082: 4079: 4077: 4074: 4073: 4069: 4061: 4055: 4040: 4036: 4032: 4028: 4024: 4017: 4014: 4009: 4005: 4001: 3997: 3993: 3989: 3988: 3980: 3977: 3965: 3961: 3957: 3952: 3947: 3943: 3939: 3932: 3930: 3926: 3921: 3917: 3912: 3907: 3903: 3899: 3894: 3889: 3885: 3881: 3877: 3870: 3868: 3866: 3862: 3850: 3846: 3845: 3840: 3833: 3830: 3825: 3819: 3815: 3808: 3805: 3801: 3797: 3794: 3790: 3786: 3783: 3778: 3775: 3762: 3758: 3752: 3749: 3744: 3740: 3736: 3732: 3728: 3724: 3720: 3716: 3715: 3707: 3704: 3692: 3688: 3684: 3678: 3675: 3671: 3667: 3664: 3659: 3656: 3643: 3639: 3635: 3628: 3625: 3620: 3616: 3612: 3608: 3604: 3597: 3594: 3589: 3585: 3581: 3577: 3573: 3569: 3565: 3561: 3560: 3552: 3549: 3536: 3532: 3526: 3523: 3518: 3514: 3509: 3504: 3499: 3494: 3490: 3486: 3482: 3475: 3472: 3459: 3455: 3454: 3449: 3443: 3440: 3435: 3431: 3426: 3421: 3417: 3413: 3409: 3405: 3401: 3394: 3391: 3386: 3382: 3378: 3371: 3368: 3355: 3351: 3347: 3343: 3336: 3333: 3328: 3324: 3320: 3316: 3312: 3308: 3307: 3299: 3296: 3291: 3287: 3282: 3277: 3272: 3267: 3263: 3259: 3255: 3248: 3245: 3240: 3236: 3231: 3226: 3221: 3216: 3212: 3208: 3203: 3198: 3194: 3190: 3189: 3184: 3177: 3175: 3171: 3166: 3162: 3158: 3154: 3149: 3144: 3140: 3136: 3132: 3128: 3127: 3122: 3115: 3112: 3100: 3096: 3092: 3088: 3084: 3080: 3076: 3072: 3068: 3064: 3060: 3059: 3054: 3047: 3044: 3031: 3027: 3026: 3021: 3014: 3012: 3008: 3003: 2999: 2995: 2991: 2987: 2983: 2982: 2977: 2970: 2967: 2963: 2959: 2956: 2951: 2948: 2943: 2939: 2934: 2929: 2925: 2921: 2917: 2913: 2912: 2907: 2900: 2898: 2894: 2889: 2885: 2881: 2877: 2873: 2872:10.1038/44751 2869: 2865: 2861: 2857: 2853: 2852: 2844: 2841: 2836: 2832: 2825: 2822: 2817: 2813: 2808: 2803: 2799: 2795: 2791: 2784: 2781: 2776: 2772: 2765: 2763: 2761: 2759: 2755: 2750: 2746: 2741: 2736: 2732: 2728: 2724: 2720: 2716: 2712: 2708: 2701: 2699: 2697: 2693: 2688: 2684: 2680: 2676: 2672: 2668: 2664: 2660: 2656: 2652: 2651: 2643: 2641: 2639: 2637: 2633: 2628: 2624: 2620: 2616: 2612: 2608: 2604: 2600: 2593: 2590: 2578: 2574: 2570: 2565: 2560: 2556: 2552: 2548: 2544: 2540: 2536: 2529: 2522: 2519: 2514: 2510: 2506: 2502: 2498: 2494: 2490: 2486: 2482: 2478: 2471: 2468: 2463: 2459: 2455: 2451: 2447: 2443: 2442: 2434: 2431: 2418: 2414: 2407: 2405: 2403: 2401: 2399: 2395: 2390: 2386: 2381: 2376: 2372: 2368: 2364: 2360: 2356: 2349: 2347: 2345: 2343: 2341: 2337: 2332: 2328: 2323: 2318: 2314: 2310: 2306: 2302: 2298: 2291: 2288: 2275: 2271: 2267: 2261: 2258: 2253: 2249: 2244: 2239: 2235: 2231: 2230: 2225: 2218: 2215: 2204:on 2021-01-26 2200: 2196: 2192: 2188: 2184: 2180: 2176: 2172: 2168: 2164: 2160: 2159: 2151: 2144: 2142: 2138: 2126: 2122: 2118: 2112: 2109: 2097: 2093: 2089: 2088:"a_derijardi" 2083: 2080: 2068: 2064: 2060: 2054: 2052: 2048: 2036: 2032: 2028: 2022: 2020: 2016: 2004: 2000: 1996: 1990: 1987: 1982: 1976: 1973: 1968: 1966:9780387771014 1962: 1958: 1951: 1949: 1947: 1945: 1943: 1941: 1937: 1924: 1920: 1919: 1914: 1913:"Stomatopoda" 1908: 1905: 1900: 1894: 1890: 1886: 1881: 1880: 1874: 1867: 1864: 1859: 1855: 1851: 1845: 1842: 1837: 1831: 1827: 1822: 1821: 1815: 1809: 1806: 1793: 1789: 1788: 1783: 1776: 1774: 1770: 1765: 1761: 1756: 1751: 1746: 1741: 1737: 1733: 1729: 1722: 1720: 1716: 1705:on 2013-05-12 1701: 1697: 1690: 1689: 1681: 1678: 1672: 1668: 1663: 1658: 1657: 1653: 1651: 1647: 1645: 1641: 1640:carcinologist 1632:, New Zealand 1631: 1630: 1626: 1625: 1624: 1620: 1616: 1615: 1611: 1609: 1608: 1604: 1602: 1601: 1597: 1592: 1591: 1578: 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Myers 3793:ReefKeeping 3508:11603/13393 2270:Space Daily 1887:. pp.  1308:pitik-pitik 1301:Philippines 1190:, salt and 1040:true shrimp 1004:fluorescent 949:tidal cycle 768:Description 738:N. wennerae 686:cephalopods 522:crepuscular 491:common name 489:" in their 457:subtropical 391:Stomatopoda 384:crustaceans 378:carnivorous 317:Coronididae 217:Stomatopoda 207:Hoplocarida 6246:Categories 5922:Thalassina 5868:Langostino 5677:Jonah crab 5079:Spare ribs 5059:Saang mein 4902:Lo mai chi 4760:Swiss wing 4725:Lo mai gai 4690:Beef tripe 4429:Tanaidacea 4399:Bochusacea 4358:Peracarida 4237:Arthropoda 4217:Orders of 4153:nature.com 4131:Pharyngula 4045:2024-03-16 3970:2023-05-23 3944:: 95–110. 3855:2017-09-04 3767:January 8, 3697:2009-11-13 3464:6 February 3105:2021-05-24 2583:2019-09-02 2208:2017-05-02 2131:2022-07-21 2102:2022-07-21 2073:2022-07-21 2041:2022-07-21 2009:2022-07-21 1929:August 26, 1814:Ross Piper 1709:2009-12-14 1673:References 1570:Squillidae 1412:stem-group 1408:Eopteridae 1327:Gilbertese 1288:) and the 1215:Tiền Giang 1196:fish sauce 1013:monogamous 888:microvilli 884:waveplates 835:circularly 734:N. bredini 659:shock wave 650:cavitation 644:; 51  506:Unipeltata 393:(from 342:Squillidae 203:Subclass: 187:Arthropoda 5858:King crab 5089:Tofu skin 5069:Shahe fen 5049:Pig's ear 5019:Frog legs 5009:Fish ball 4984:Beef ball 4765:Taro cake 4405:Amphipoda 4329:Syncarida 4243:Crustacea 3960:1802-8225 3902:2167-8359 3886:: e3844. 3844:USA Today 3619:191386729 3202:0804.2162 2731:0022-0949 2564:1912/1391 2513:205017718 2448:(6): 11. 1738:: e3844. 1371:live rock 1285:canocchia 1146:, called 1127:Thanh Hóa 1098:talk page 945:fluoresce 938:barracuda 804:ommatidia 640:(83  628:raptorial 559:sediments 518:nocturnal 479:raptorial 475:Australia 471:Assyrians 449:predators 222:Latreille 173:Kingdom: 167:Eukaryota 6035:BugGuide 5966:Wikidata 5934:Category 5760:Crayfish 5341:Lobsters 5084:Suan cai 5054:Rapeseed 5039:Ong choy 5014:Fishcake 4968:XO sauce 4937:Tong sui 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Index

Mantis shrimps
PreꞒ

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Pg
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Odontodactylus scyllarus

Lysiosquillina maculata
Scientific classification
Edit this classification
Eukaryota
Animalia
Arthropoda
Malacostraca
Hoplocarida
Stomatopoda
Latreille
Bathysquillidae
Indosquillidae
Gonodactyloidea
Alainosquillidae

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