39:
46:
2591:
2349:
Land surrounding the area is either wooded or used for agriculture. Cattle ranching is important, as are hundreds of pineapple and sugar cane plantations. Other crops are bananas,
2551:
38:
2707:"Primer reporte de un árbol con nidos activos de guacamayo rojo (Ara macao) y guacamayo verde mayor (Ara ambiguus) en bosque muy húmedo tropical de Centroamérica"
2881:
2510:
248:, thereby allowing animals to move between the regions. This region is now known as the San Juan-La Selva Biological Corridor, located between the
2871:
88:
2876:
338:
210:
2886:
1836:
229:
233:
1234:
349:
It features a very wet, non-seasonal, evergreen rainforest environment with a yearly precipitation of 3,000-3,600mm. 70.21% is
225:
1391:
1415:
249:
1385:
969:
Fifty meters from the lakes the following plants have been recorded as food plants for the great green macaw: the palms
318:
in this area, in order to help link other existing neighbouring areas in Costa Rica and
Nicaragua, and thus protect the
260:
72:
1793:
1675:
1379:
1269:
241:
237:
221:
1397:
770:
365:
2060:
1816:
1421:
552:
379:
ecosystems. The lake complex consists of at least four major lakes: Canacas, Colpachi, Maquenque and
Tamborcito.
540:
2343:
2210:
2118:
2044:
1688:
1409:
1178:
1139:
853:
528:
217:
2812:
2705:
Chassot, Olivier; Monge Arias, Guisselle; Alemán Zelaya, Indalecio; González Téllez, Adolfo (November 2011).
758:
609:
2228:
2160:
2076:
2068:
1556:
1547:
1451:
1403:
1368:
1172:
1151:
858:
661:
576:
454:
1333:
1166:
1133:
1080:
1039:
956:
888:
546:
2840:
2264:
2110:
2019:
1947:
1880:
1860:
1562:
1491:
1373:
1264:
821:
650:
485:
464:
329:
The concept of this area as a biological corridor was already proposed in 1985 by the first revolutionary
2565:
1228:
1021:
722:
598:
404:
2545:
2511:"CONNECTIVITY CONSERVATION OF THE GREAT GREEN MACAW'S LANDSCAPE IN COSTA RICA AND NICARAGUA (1994-2012)"
2324:
1909:
1885:
1725:
1700:
1275:
1201:
1098:
962:
776:
716:
622:
592:
582:
491:
330:
1618:
1600:
1352:
746:
558:
1847:
1613:
1594:
1568:
827:
734:
728:
645:
628:
570:
564:
497:
2484:
2318:
2269:
2036:
1788:
1606:
1588:
1577:
1522:
1321:
1255:
989:
914:
673:
515:
942:
919:
764:
740:
2192:
2186:
2180:
2174:
2145:
2134:
2099:
2093:
2025:
1874:
1457:
1339:
1315:
1299:
1287:
1248:
1243:
1195:
1108:
833:
791:
604:
424:
1310:
687:
2246:
2234:
2222:
1990:
1960:
1810:
1720:
1706:
1694:
1664:
1654:
1534:
1517:
1507:
1496:
1222:
1207:
1092:
971:
751:
470:
1190:
The following amphibians, all considered threatened in Costa Rica, occur here (as of 2001): the
253:
2747:
353:, 6.17% is secondary forest or scrubland, and 20.13% is devoted to agriculture. There are many
2718:
2706:
2461:
2306:
2301:
2281:
2105:
1985:
1897:
1782:
1773:
1747:
1741:
1735:
1632:
1583:
1528:
1445:
1436:
1363:
1293:
1160:
1122:
616:
479:
439:
434:
350:
334:
319:
264:
153:
76:
2380:
1662:
which is reasonably common here. Other parrots considered locally threatened which occur are
275:
The refuge was created in 2005 from existing reserves such as El JardĂn Hill Forest
Reserve (
2645:
2525:
2312:
2290:
2273:
2258:
2240:
2216:
2084:
1968:
1930:
1902:
1767:
1669:
1475:
1358:
1345:
1328:
1282:
1239:
1145:
1086:
1074:
797:
667:
534:
418:
394:
300:
292:
284:
276:
198:
2529:
2330:
2252:
1637:
1463:
1216:
1117:
995:
843:
785:
781:
509:
503:
448:
399:
323:
2354:
2296:
2165:
2151:
2139:
2052:
2002:
1822:
696:
2865:
2411:
2350:
2286:
2277:
2204:
2198:
1855:
1831:
1801:
1711:
1659:
1541:
1485:
333:
as the "Sistema
Internacional de Areas Protegidas para la Paz" (SI-A-PAZ). It became
315:
311:
2010:
1914:
1753:
361:
2361:. Due to the wildlife refuge status, development is restricted for the residents.
1998:
1922:
1865:
1731:
1469:
1976:
1938:
1758:
1512:
1260:
1212:
817:
443:
376:
360:
The reserve included wetlands and protected an example of the virgin tropical
206:
62:
2722:
2670:
103:
90:
2032:
1952:
1480:
1305:
1191:
702:
245:
1642:
1128:
354:
1072:
There have been 80 fish species registered as of 2008. The bobo mullet (
2358:
2156:
1827:
1156:
1113:
524:
474:. Intermediate succession lowland woodlands are home to the threatened
2818:(Report) (in Spanish). Centro CientĂfico Tropical. 2009. pp. 2–12
1137:, found at a range of altitudes here. Threatened fish species include
1893:
1778:
1650:
1441:
1845:. The only bird endemic to Costa Rica found here is the hummingbird
2456:
1434:
Crocodilians are represented by the endangered
American crocodile (
1573:
1552:
475:
428:
borders lakes in dense stands. Dry land contains large amounts of
307:), which were deprecated and assimilated into the current refuge.
140:
2410:
Chassot, Olivier; Monge Arias, Guisselle; Powell, George (2007).
2651:(Report) (in Spanish). Centro CientĂfico Tropical. pp. 1–34
1539:, all endemics to Costa Rica. Lizards include the green iguana (
392:
The permanently inundated swamps are home to stands of the palm
372:
2748:"Boca Tapada, Northern Lowlands - Costa Rica 2019 Travel Guide"
1344:
and those tree frogs with no particular elevation preferences:
2342:
The nearby town of Boca Tapada in Pital, has an abundance of
2590:
Arias, Guisselle Monge; Chassott, Olivier (November 2002).
2035:
birds which occur here and are rare for Costa Rica are the
402:. Other plants associated with this habitat are the trees
322:
population. Instead, the Costa Rican government created a
45:
2671:"Salvemos la Reserva Indio MaĂz, pulmĂłn de CentroamĂ©rica"
2509:
Chassot, Olivier; Monge Arias, Guisselle (1 March 2012).
398:
and are surrounded by thickets of species in the family
2813:
Humedales de Ramsar (FIR) – Anexo #2 Biodiversidad 2009
1901:. Locally threatened birds of prey include the falcons
878:
Lake
Tamborcito has in its surroundings the tall trees
501:. Non-native trees in secondary woodland are primarily
1274:
both endemics found only in the higher elevations and
1455:) and the locally endangered common snapping turtle (
1280:
in the canopy at all elevations, and the salamanders
357:, vines and climbing plants, and a dense understory.
2592:"Great Green Macaw: Flagship Species of Costa Rica"
1449:. Turtles include the Central American mud turtle (
1238:restricted to higher elevations, the low elevation
720:surrounding it, interspersed with smaller trees of
176:
168:
160:
151:
147:
135:
127:
119:
82:
68:
58:
18:
2328:and the uncommon Caribbean white tent-making bat (
2178:), the locally reduced Hoffmann's two-toed sloth (
337:in 2010. The area is administratively part of the
2550:: CS1 maint: DOI inactive as of September 2024 (
1096:) are quite common. The endangered tropical gar (
2807:
2805:
2803:
2801:
2799:
2797:
2795:
2793:
2791:
2789:
364:that once blanketed the territory alongside the
213:, it was created in 2005 by decree 32405-MINAE.
2787:
2785:
2783:
2781:
2779:
2777:
2775:
2773:
2771:
2769:
1545:), the endemic Yautepec tropical night lizard (
1384:of mid to higher elevations, the low elevation
1362:). There are particularly many rare species of
2700:
2698:
2696:
2694:
2692:
1402:from low to mid elevations, the low elevation
2835:
2833:
2646:Humedales de Ramsar (FIR) – Versión 2006-2008
2639:
2637:
2635:
2405:
2403:
2401:
422:grows in this habitat in more inland swamps.
24:
8:
2633:
2631:
2629:
2627:
2625:
2623:
2621:
2619:
2617:
2615:
256:and other areas near the Nicaraguan border.
2585:
2583:
635:Lake Canaca is surrounded by tall trees of
2375:
2373:
2276:. Monkeys included the locally endangered
1426:with no particular elevation preferences.
299:) and the Tamborcito Lacustrine Wetlands (
15:
2741:
2739:
2381:"Refugio de Vida Silvestre Maquenque Map"
1739:, the threatened South American bittern (
812:Lake Maquenque is fringed with shrubs of
2451:
2449:
2447:
2445:
2443:
2441:
123:518.61 km (200.24 sq mi)
2369:
2133:As of 1997 mammals included endangered
1372:with low to mid elevation preferences,
1149:. Other fish include the common snook (
139:National System of Conservation Areas (
2543:
2310:, and only at low elevations the rare
1808:) at the lowest altitudes, the motmot
1686:). Other birds include dusky antbird (
442:, which may grow to 60m high. Another
2530:10.2305/IUCN.CH.2012.PARKS-18-1.OC.en
2485:"Refugio de Vida Silvestre Maquenque"
2190:), the locally threatened armadillo (
2125:) usually found at higher altitudes.
1814:, the locally threatened green ibis (
1106:, occurs here. The large bull shark (
339:Arenal Huetar Norte Conservation Area
211:Arenal Huetar Norte Conservation Area
175:
167:
159:
150:
7:
924:, the smallish pioneer tree species
57:
2882:Protected areas established in 2005
801:in former clearings, and shrubs of
259:The nearest towns and villages are
203:Refugio de Vida Silvestre Maquenque
26:Refugio de Vida Silvestre Maquenque
2644:Mauricio Salas Varga (July 2008).
2568:. Centro CientĂfico Tropical. 2017
2268:, and both the locally endangered
1396:from mid to very high elevations,
694:Lake Colpachi has the large trees
574:. Rare plants furthermore include
230:Barra del Colorado Wildlife Refuge
19:Maquenque National Wildlife Refuge
14:
523:). Endangered plants include the
438:), a favoured food source of the
2412:"BiologĂa de la ConservaciĂłn de
2304:, Bats included the vampire bat
2244:), the locally endangered otter
1420:from mid to high elevations and
1356:and the spiny-headed tree frog (
281:Reserva Forestal Cerro El JardĂn
44:
37:
2065:Xiphocolaptes promeropirhynchus
1751:), the vulnerable agami heron (
1235:Hyalinobatrachium vireovittatum
580:, and threatened lowland trees
126:
67:
2872:Geography of Alajuela Province
2675:Salvemos la Reserva Indio MaĂz
1826:), and the locally threatened
1378:(mid elevations), the endemic
1304:at low elevations. Threatened
556:. Rare endemic plants include
452:. Other common trees here are
291:), Laguna Maquenque Wetlands (
1:
2877:Nature reserves in Costa Rica
1631:Endangered birds include the
1440:) and in reduced numbers the
1326:(endemic) and the endangered
771:Stryphnodendron microstachyum
305:Humedal Lacustrino Tamborcito
234:Indio-MaĂz Biological Reserve
2109:at lower altitudes; and the
1858:include the solitary eagle (
1489:). Other boas are the small
1414:from low to mid elevations,
2316:, the rare disk-winged bat
1794:green-and-rufous kingfisher
1676:rufous-vented ground cuckoo
1308:include the high elevation
805:. Invasive species include
681:is dominant, and the palms
553:Hymenolobium mesoamericanum
242:Cerro Silva Natural Reserve
238:Punta Gorda Natural Reserve
222:La Selva Biological Station
2903:
2887:Ramsar sites in Costa Rica
2155:, the nationally uncommon
2123:Cephalopterus glabricollis
1692:), rufescent tiger-Heron (
1674:. An uncommon bird is the
1473:, the endemic earth snake
541:Sclerolobium costarricense
283:), Cureña Forest Reserve (
205:), is a protected area in
2841:"Boca Tapada, Costa Rica"
2464:Sites Information Service
2416:en Costa Rica, 1994-2006"
2294:and the locally uncommon
2184:), brown-throated sloth (
2164:, the locally endangered
2061:strong-billed woodcreeper
1965:Leucopternis semiplumbeus
1820:) and roseate spoonbill (
1817:Mesembrinibis cayennensis
1792:at higher altitudes, the
1232:, the endemic glass frog
1140:Cynodonichthys isthmensis
1084:, knife livebearer fish (
898:Eschweilera costaricensis
854:Eschweilera costaricensis
261:Puerto Viejo de SarapiquĂ
195:Maquenque Wildlife Refuge
188:
184:
73:Puerto Viejo de SarapiquĂ
32:
23:
2338:Socioeconomic activities
2211:Herpailurus yagouaroundi
2119:bare-necked umbrellabird
2115:Procnias tricarunculatus
2073:Campylorhamphus pusillus
2045:tawny-chested flycatcher
1981:Geranospiza caerulescens
1689:Cercomacroides tyrannina
1179:Hypsophrys nicaraguensis
529:Podocarpus guatemalensis
314:proposed creating a new
297:Humedal Laguna Maquenque
228:in Costa Rica, with the
218:Tortuguero National Park
2229:Bassariscus sumichrasti
2170:Myrmecophaga tridactyla
2161:Metachirus nudicaudatus
2077:long-tailed woodcreeper
2069:brown-billed scythebill
2049:Aphanotriccus capitalis
2001:include the vulnerable
1957:Accipiter superciliosus
1935:Chondrohierax uncinatus
1698:), keel-billed toucan (
1557:Thecadactylus rapicauda
1548:Lepidophyma reticulatum
1452:Kinosternon angustipons
1369:Eleutherodactylus altae
1173:Parachromis managuensis
1152:Centropomus undecimalis
906:Ferdinandusa panamensis
759:Ferdinandusa panamensis
657:Calophyllum brasiliense
655:, and smaller trees of
577:Aspidosperma spruceanum
455:Calophyllum brasiliense
289:Reserva Forestal Cureña
2265:Odocoileus virginianus
2111:three-wattled bellbird
2081:Deconychura longicauda
2020:Odontophorus melanotis
1983:) and the hawk-eagles
1973:Buteogallus urubitinga
1948:Busarellus nigricollis
1881:Spizastur melanoleucus
1861:Buteogallus solitarius
1563:Ptychoglossus plicatus
1513:Tropical ground snakes
1492:Ungaliophis panamensis
1265:Bolitoglossa alvaradoi
1031:Pentaclethra macroloba
1009:Cespedesia macrophylla
954:. Rare plants include
884:Pentaclethra macroloba
869:Pentaclethra macroloba
841:), the smallish trees
708:Pentaclethra macroloba
662:Cespedesia macrophylla
637:Pentaclethra macroloba
626:and the very uncommon
465:Pentaclethra macroloba
410:Pentaclethra macroloba
304:
296:
288:
280:
202:
25:
2847:. Pacific Trade Winds
2532:(inactive 2024-09-12)
2325:Micronycteris daviesi
2067:); the mid-elevation
2015:Penelope purpurascens
1943:Rostrhamus sociabilis
1910:Micrastur mirandollei
1886:red-throated caracara
1838:Crypturellus boucardi
1745:) and least bittern (
1701:Ramphastos sulfuratus
1202:Gymnopis multiplicata
1167:Parachromis loisellei
1134:Priapichtys annectens
1099:Atractosteus tropicus
1081:Brachyrhaphis olomina
1040:Sacoglottis tricogyna
993:, the emergent trees
957:Elaeoluma glabrescens
889:Swartzia maquenqueana
863:, and large trees of
583:Handroanthus guayacan
547:Platymiscium pinnatum
492:Minquartia guianensis
331:Sandinista government
2319:Thyroptera discifera
2270:white-lipped peccary
2037:yellow-tailed oriole
1789:Pharomachrus mocinno
1716:Patagioenas speciosa
1680:Neomorphus geoffroyi
1578:Polychrus gutturosus
1332:, the low elevation
1256:Phyllobates lugubris
1211:, the low elevation
1102:), known locally as
896:, the smaller trees
865:Terminalia amazonica
837:, the walking palm (
822:Maranthes panamensis
782:pioneer tree species
652:Palicourea tomentosa
516:Terminalia ivorensis
486:Terminalia amazonica
310:In 1998 a number of
209:, managed under the
2457:"Humedal Maquenque"
2193:Cabassous centralis
2187:Bradypus variegatus
2181:Choloepus hoffmanni
2175:Cyclopes didactylus
2172:), silky anteater (
2146:Dasyprocta punctata
2100:Carpodectes nitidus
2094:Piprites griseiceps
2089:Laniocera rufescens
2026:Rhynchortyx cinctus
1927:Lophostrix cristata
1875:Morphnus guianensis
1854:Locally endangered
1483:-eating mussurana (
1458:Chelydra serpentina
1244:Dendrobates auratus
1229:Centrolenella magna
1196:Dermophis parviceps
1127:can also occur. An
1109:Carcharhinus leucas
1057:Vochysia ferruginea
1022:Guarea rhopalocarpa
952:Mouriri gleasoniana
934:Tapirira guianensis
894:Vochysia ferruginea
873:Vochysia ferruginea
834:Mouriri gleasoniana
792:Tapirira guianensis
723:Alibertia atlantica
712:Vochysia ferruginea
641:Vochysia ferruginea
605:Dalbergia glomerata
599:Copaifera aromatica
460:Vochysia ferruginea
405:Vochysia ferruginea
136:Governing body
100: /
2307:Vampyrum spectrum
2247:Lontra longicaudis
2235:Bassaricyon gabbii
2223:Leopardus pardalis
2009:), the threatened
1961:semiplumbeous hawk
1890:Ibycter americanus
1811:Hylomanes momotula
1783:Trogon clathratus
1721:Geotrygon violacea
1707:Psittacara finschi
1695:Tigrisoma lineatum
1665:Amazona autumnalis
1655:Electron carinatum
1572:(endemic) and the
1518:Trimetopon gracile
1508:Epicrates cenchria
1498:Corallus annulatus
1461:). Snakes include
1223:Teratohyla spinosa
1208:Bufo melanochloris
1093:Astyanax fasciatus
1045:Vantanea barbourii
1013:Croton schiedeanus
987:, the large shrub
981:Socratea exorrhiza
972:Iriartea deltoidea
963:Vantanea barbourii
938:Socratea exorrhiza
839:Socratea exorrhiza
777:Vismia macrophylla
717:Pouteria durlandii
683:Socratea exorrhiza
623:Terminalia oblonga
593:Sideroxylon capiri
519:(locally known as
471:Socratea exorrhiza
414:Socratea exorrhiza
226:Cordillera Central
177:Reference no.
2302:White-nosed coati
2282:Alouatta palliata
2106:Lanio leucothorax
2041:Icterus mesomelas
2017:) and the quails
1986:Spizaetus ornatus
1919:Sarcoramphus papa
1898:Falco deiroleucus
1878:, the hawk-eagle
1806:Jacamerops aureus
1798:Chloroceryle inda
1774:Nomonyx dominicus
1765:), Muscovy duck (
1748:Ixobrychus exilis
1742:Botaurus pinnatus
1736:Lophornis helenae
1633:great green macaw
1584:Norops carpenteri
1505:and rainbow boa (
1495:, and the larger
1446:Caiman crocodilus
1437:Crocodylus acutus
1353:Hyla microcephala
1271:B. arborescandens
1240:poison dart frogs
1161:Parachromis dovii
1090:) and the tetra (
1001:Dipteryx oleifera
940:, and the shrubs
747:Croton smithianus
617:Prioria copaifera
559:Quararibea pumila
480:Carapa guianensis
440:great green macaw
435:Dipteryx oleifera
320:great green macaw
192:
191:
164:Humedal Maquenque
69:Nearest city
2894:
2857:
2856:
2854:
2852:
2845:Enter Costa Rica
2837:
2828:
2827:
2825:
2823:
2817:
2809:
2764:
2763:
2761:
2759:
2743:
2734:
2733:
2731:
2729:
2702:
2687:
2686:
2684:
2682:
2667:
2661:
2660:
2658:
2656:
2650:
2641:
2610:
2609:
2607:
2606:
2596:
2587:
2578:
2577:
2575:
2573:
2562:
2556:
2555:
2549:
2541:
2539:
2537:
2515:
2506:
2500:
2499:
2497:
2495:
2489:Protected Planet
2481:
2475:
2474:
2472:
2470:
2453:
2436:
2435:
2433:
2431:
2407:
2396:
2395:
2393:
2391:
2377:
2322:, the very rare
2313:Cyttarops alecto
2291:Ateles geoffroyi
2274:collared peccary
2259:Mazama americana
2241:Galictis vittata
2217:Leopardus wiedii
2085:speckled mourner
2057:Cotinga amabilis
1969:great black hawk
1945:), fishing hawk
1931:hook-billed kite
1848:Orcha cupreiceps
1771:), masked duck (
1768:Cairina moschata
1647:Heliornis fulica
1614:Dactyloa frenata
1569:Celestus hylaeus
1476:Geophis ruthveni
1359:Anotheca spinosa
1329:Agalychnis annae
1283:Oedipina alfaroi
1146:Dajaus monticola
1131:fish species is
1087:Alfaro cultratus
1075:Joturus pichardi
1035:Qualea paraensis
1017:Dialum guianense
1005:Byrsonima crispa
977:Raphia taedigera
948:Miconia punctata
926:Jacaranda copaia
849:Byrsonima crispa
828:Miconia punctata
798:Jacaranda copaia
735:Casearia arborea
729:Byrsonima crispa
679:Raphia taedigera
668:Couma macrocarpa
646:Dialum guianense
629:Cynometra retusa
611:D. melanocardium
571:Vochysia allenii
565:Trichilia adolfi
535:Cedrela fissilis
498:Qualea paraensis
419:Pachira aquatica
395:Raphia taedigera
335:Ramsar Site 1918
250:San Carlos River
115:
114:
112:
111:
110:
105:
101:
98:
97:
96:
93:
48:
47:
41:
28:
16:
2902:
2901:
2897:
2896:
2895:
2893:
2892:
2891:
2862:
2861:
2860:
2850:
2848:
2839:
2838:
2831:
2821:
2819:
2815:
2811:
2810:
2767:
2757:
2755:
2754:. Anywhere, Inc
2746:Andrea (2019).
2745:
2744:
2737:
2727:
2725:
2704:
2703:
2690:
2680:
2678:
2669:
2668:
2664:
2654:
2652:
2648:
2643:
2642:
2613:
2604:
2602:
2594:
2589:
2588:
2581:
2571:
2569:
2564:
2563:
2559:
2542:
2535:
2533:
2513:
2508:
2507:
2503:
2493:
2491:
2483:
2482:
2478:
2468:
2466:
2455:
2454:
2439:
2429:
2427:
2409:
2408:
2399:
2389:
2387:
2379:
2378:
2371:
2367:
2340:
2331:Ectophylla alba
2253:Tapirus bairdii
2226:), cacomistle (
2208:), jaguarundi (
2149:, the squirrel
2131:
1763:Eurypyga helias
1638:Jabiru mycteria
1629:
1464:Boa constrictor
1432:
1393:E. fleischmanni
1217:Sachatamia ilex
1188:
1118:Pristis pristis
1070:
1065:
1053:Virola sebifera
1049:Virola koschnyi
996:Balizia elegans
990:Solanum rugosum
930:Simarouba amara
915:Licania affinis
844:Virola sebifera
807:Gmelina arborea
786:Simarouba amara
691:grow here too.
674:Virola koschnyi
588:Ceiba pentandra
510:Tectona grandis
504:Gmelina arborea
449:Ceiba pentandra
400:Melastomataceae
390:
385:
347:
324:wildlife refuge
273:
254:SarapiquĂ River
156:
108:
106:
104:10.70°N 84.15°W
102:
99:
94:
91:
89:
87:
86:
54:
53:
52:
51:
50:
49:
12:
11:
5:
2900:
2898:
2890:
2889:
2884:
2879:
2874:
2864:
2863:
2859:
2858:
2829:
2765:
2735:
2717:(1–2): 72–79.
2713:(in Spanish).
2688:
2662:
2611:
2579:
2557:
2501:
2476:
2437:
2422:(in Spanish).
2397:
2368:
2366:
2363:
2339:
2336:
2297:Cebus imitator
2166:giant anteater
2152:Sciurus deppei
2140:Cuniculus paca
2130:
2127:
2053:lovely cotinga
2003:great curassow
1870:Harpia harpyja
1823:Platalea ajaja
1786:, the quetzal
1727:G. veraguensis
1628:
1625:
1467:, the endemic
1431:
1428:
1417:E. podiciferus
1187:
1184:
1069:
1066:
1064:
1061:
1027:Laetia procera
943:Hirtella media
910:Laetia procera
902:|E. panamensis
880:Brosimum utile
765:Laetia procera
753:C. schiedeanus
741:Cordia bicolor
697:Brosimum utile
532:and the trees
389:
386:
384:
381:
371:It also has a
366:San Juan River
351:primary forest
346:
343:
312:ornithologists
272:
269:
190:
189:
186:
185:
182:
181:
178:
174:
173:
170:
166:
165:
162:
158:
157:
154:Ramsar Wetland
152:
149:
148:
145:
144:
137:
133:
132:
129:
125:
124:
121:
117:
116:
84:
80:
79:
70:
66:
65:
60:
56:
55:
43:
42:
36:
35:
34:
33:
30:
29:
21:
20:
13:
10:
9:
6:
4:
3:
2:
2899:
2888:
2885:
2883:
2880:
2878:
2875:
2873:
2870:
2869:
2867:
2846:
2842:
2836:
2834:
2830:
2814:
2808:
2806:
2804:
2802:
2800:
2798:
2796:
2794:
2792:
2790:
2788:
2786:
2784:
2782:
2780:
2778:
2776:
2774:
2772:
2770:
2766:
2753:
2749:
2742:
2740:
2736:
2724:
2720:
2716:
2712:
2708:
2701:
2699:
2697:
2695:
2693:
2689:
2676:
2672:
2666:
2663:
2647:
2640:
2638:
2636:
2634:
2632:
2630:
2628:
2626:
2624:
2622:
2620:
2618:
2616:
2612:
2600:
2593:
2586:
2584:
2580:
2567:
2566:"Green Macaw"
2561:
2558:
2553:
2547:
2531:
2527:
2523:
2519:
2512:
2505:
2502:
2490:
2486:
2480:
2477:
2465:
2463:
2458:
2452:
2450:
2448:
2446:
2444:
2442:
2438:
2425:
2421:
2420:Mesoamericana
2417:
2415:
2406:
2404:
2402:
2398:
2386:
2382:
2376:
2374:
2370:
2364:
2362:
2360:
2356:
2352:
2351:heart of palm
2347:
2345:
2337:
2335:
2333:
2332:
2327:
2326:
2321:
2320:
2315:
2314:
2309:
2308:
2303:
2299:
2298:
2293:
2292:
2288:
2287:spider monkey
2284:
2283:
2279:
2278:howler monkey
2275:
2271:
2267:
2266:
2261:
2260:
2255:
2254:
2249:
2248:
2243:
2242:
2237:
2236:
2231:
2230:
2225:
2224:
2219:
2218:
2213:
2212:
2207:
2206:
2205:Puma concolor
2201:
2200:
2199:Panthera onca
2195:
2194:
2189:
2188:
2183:
2182:
2177:
2176:
2171:
2167:
2163:
2162:
2158:
2154:
2153:
2148:
2147:
2143:, the agouti
2142:
2141:
2136:
2128:
2126:
2124:
2120:
2116:
2112:
2108:
2107:
2102:
2101:
2096:
2095:
2090:
2086:
2082:
2078:
2074:
2070:
2066:
2062:
2058:
2054:
2050:
2046:
2042:
2038:
2034:
2030:
2028:
2027:
2022:
2021:
2016:
2012:
2008:
2004:
2000:
1996:
1994:
1993:
1988:
1987:
1982:
1978:
1974:
1970:
1966:
1962:
1958:
1954:
1950:
1949:
1944:
1940:
1936:
1932:
1928:
1924:
1920:
1916:
1912:
1911:
1906:
1905:
1904:F. peregrinus
1900:
1899:
1895:
1891:
1887:
1883:
1882:
1877:
1876:
1872:), the eagle
1871:
1867:
1863:
1862:
1857:
1856:birds of prey
1852:
1850:
1849:
1844:
1843:
1842:costaricensis
1839:
1834:
1833:
1832:Tinamus major
1829:
1825:
1824:
1819:
1818:
1813:
1812:
1807:
1803:
1802:great jacamar
1799:
1795:
1791:
1790:
1785:
1784:
1780:
1776:
1775:
1770:
1769:
1764:
1760:
1756:
1755:
1750:
1749:
1744:
1743:
1738:
1737:
1733:
1729:
1728:
1723:
1722:
1718:), the doves
1717:
1713:
1712:scaled pigeon
1709:
1708:
1703:
1702:
1697:
1696:
1691:
1690:
1685:
1681:
1677:
1673:
1672:
1667:
1666:
1661:
1660:scarlet macaw
1657:
1656:
1652:
1648:
1644:
1640:
1639:
1634:
1626:
1624:
1622:
1621:
1616:
1615:
1610:
1609:
1604:
1603:
1602:N. pentaprion
1598:
1597:
1592:
1591:
1586:
1585:
1580:
1579:
1575:
1571:
1570:
1565:
1564:
1559:
1558:
1554:
1550:
1549:
1544:
1543:
1542:Iguana iguana
1538:
1537:
1532:
1531:
1526:
1525:
1520:
1519:
1514:
1510:
1509:
1504:
1503:
1499:
1494:
1493:
1488:
1487:
1486:Clelia clelia
1482:
1478:
1477:
1472:
1471:
1466:
1465:
1460:
1459:
1454:
1453:
1448:
1447:
1443:
1439:
1438:
1429:
1427:
1425:
1424:
1419:
1418:
1413:
1412:
1407:
1406:
1401:
1400:
1395:
1394:
1389:
1388:
1387:E. biporcatus
1383:
1382:
1377:
1376:
1371:
1370:
1365:
1361:
1360:
1355:
1354:
1349:
1348:
1343:
1342:
1337:
1336:
1335:A. calcarifer
1331:
1330:
1325:
1324:
1323:H. rufioculis
1319:
1318:
1313:
1312:
1307:
1303:
1302:
1301:O. cyclocauda
1297:
1296:
1291:
1290:
1289:O. carablanca
1285:
1284:
1279:
1278:
1273:
1272:
1267:
1266:
1262:
1258:
1257:
1252:
1251:
1246:
1245:
1241:
1237:
1236:
1231:
1230:
1225:
1224:
1219:
1218:
1214:
1210:
1209:
1204:
1203:
1198:
1197:
1193:
1185:
1183:
1181:
1180:
1175:
1174:
1169:
1168:
1163:
1162:
1158:
1154:
1153:
1148:
1147:
1142:
1141:
1136:
1135:
1130:
1126:
1125:
1120:
1119:
1115:
1111:
1110:
1105:
1101:
1100:
1095:
1094:
1089:
1088:
1083:
1082:
1077:
1076:
1067:
1062:
1060:
1058:
1054:
1050:
1046:
1042:
1041:
1036:
1032:
1028:
1024:
1023:
1018:
1014:
1010:
1006:
1002:
998:
997:
992:
991:
986:
982:
978:
974:
973:
967:
965:
964:
959:
958:
953:
949:
945:
944:
939:
935:
931:
927:
923:
922:
917:
916:
911:
907:
903:
899:
895:
891:
890:
885:
881:
876:
874:
870:
866:
862:
861:
860:E. panamensis
856:
855:
850:
846:
845:
840:
836:
835:
830:
829:
825:, the shrubs
824:
823:
819:
815:
814:Zygia confusa
810:
808:
804:
803:Zygia confusa
800:
799:
794:
793:
788:
787:
783:
779:
778:
773:
772:
767:
766:
761:
760:
755:
754:
749:
748:
743:
742:
737:
736:
731:
730:
725:
724:
719:
718:
713:
709:
705:
704:
699:
698:
692:
690:
689:
684:
680:
676:
675:
670:
669:
664:
663:
658:
654:
653:
648:
647:
642:
638:
633:
631:
630:
625:
624:
619:
618:
613:
612:
607:
606:
601:
600:
595:
594:
589:
585:
584:
579:
578:
573:
572:
567:
566:
561:
560:
555:
554:
549:
548:
543:
542:
537:
536:
531:
530:
526:
522:
518:
517:
512:
511:
506:
505:
500:
499:
494:
493:
488:
487:
482:
481:
477:
473:
472:
468:and the palm
467:
466:
461:
457:
456:
451:
450:
445:
444:emergent tree
441:
437:
436:
431:
427:
426:
425:Zygia confusa
421:
420:
415:
412:and the palm
411:
407:
406:
401:
397:
396:
387:
382:
380:
378:
374:
369:
367:
363:
358:
356:
352:
344:
342:
340:
336:
332:
327:
325:
321:
317:
316:national park
313:
308:
306:
302:
298:
294:
290:
286:
282:
278:
270:
268:
266:
262:
257:
255:
251:
247:
243:
239:
235:
231:
227:
223:
219:
214:
212:
208:
204:
200:
196:
187:
183:
179:
171:
163:
161:Official name
155:
146:
142:
138:
134:
130:
122:
118:
113:
109:10.70; -84.15
85:
81:
78:
74:
71:
64:
61:
40:
31:
27:
22:
17:
2849:. Retrieved
2844:
2820:. Retrieved
2756:. Retrieved
2751:
2726:. Retrieved
2714:
2710:
2679:. Retrieved
2674:
2665:
2653:. Retrieved
2603:. Retrieved
2598:
2570:. Retrieved
2560:
2546:cite journal
2534:. Retrieved
2521:
2517:
2504:
2492:. Retrieved
2488:
2479:
2467:. Retrieved
2460:
2428:. Retrieved
2423:
2419:
2414:Ara ambiguus
2413:
2388:. Retrieved
2384:
2348:
2341:
2329:
2323:
2317:
2311:
2305:
2295:
2289:
2280:
2263:
2257:
2256:), the deer
2251:
2245:
2239:
2233:
2227:
2221:
2215:
2209:
2203:
2197:
2191:
2185:
2179:
2173:
2169:
2159:
2150:
2144:
2138:
2132:
2122:
2114:
2104:
2098:
2092:
2088:
2080:
2072:
2064:
2056:
2048:
2040:
2031:
2024:
2018:
2014:
2011:crested guan
2006:
1997:
1991:
1984:
1980:
1972:
1964:
1956:
1946:
1942:
1934:
1926:
1918:
1915:king vulture
1908:
1903:
1896:
1889:
1879:
1873:
1869:
1859:
1853:
1846:
1841:
1837:
1830:
1821:
1815:
1809:
1805:
1797:
1787:
1781:
1772:
1766:
1762:
1754:Agamia agami
1752:
1746:
1740:
1734:
1726:
1719:
1715:
1705:
1699:
1693:
1687:
1683:
1679:
1670:
1663:
1653:
1646:
1636:
1635:, the stork
1630:
1619:
1612:
1607:
1601:
1596:N. lemurinus
1595:
1589:
1582:
1576:
1567:
1561:
1555:
1546:
1540:
1535:
1529:
1524:T. pliolepis
1523:
1516:
1506:
1501:
1497:
1490:
1484:
1474:
1468:
1462:
1456:
1450:
1444:
1435:
1433:
1422:
1416:
1410:
1404:
1398:
1392:
1386:
1381:E. angelicus
1380:
1374:
1367:
1357:
1351:
1346:
1340:
1334:
1327:
1322:
1316:
1311:Hyla colymba
1309:
1300:
1294:
1288:
1281:
1276:
1270:
1263:
1254:
1249:
1242:
1233:
1227:
1221:
1215:
1206:
1200:
1194:
1189:
1177:
1171:
1165:
1159:
1150:
1144:
1138:
1132:
1124:P. pectinata
1123:
1116:
1107:
1103:
1097:
1091:
1085:
1079:
1073:
1071:
1056:
1052:
1048:
1044:
1038:
1034:
1030:
1026:
1020:
1016:
1012:
1008:
1004:
1003:, the trees
1000:
994:
988:
985:Welfia regia
984:
980:
976:
970:
968:
961:
955:
951:
947:
941:
937:
933:
929:
925:
920:
913:
909:
905:
901:
897:
893:
887:
883:
879:
877:
872:
868:
864:
859:
852:
848:
842:
838:
832:
826:
820:
816:, the large
813:
811:
806:
802:
796:
790:
784:
775:
769:
763:
757:
752:
745:
739:
733:
727:
721:
715:
711:
707:
701:
695:
693:
688:Welfia regia
686:
682:
678:
672:
666:
660:
656:
651:
644:
640:
636:
634:
627:
621:
615:
610:
603:
597:
591:
587:
581:
575:
569:
563:
557:
551:
545:
539:
533:
527:
520:
514:
508:
502:
496:
490:
484:
478:
469:
463:
459:
453:
447:
433:
429:
423:
417:
413:
409:
403:
393:
391:
375:complex and
370:
362:moist forest
359:
348:
328:
309:
274:
258:
216:It connects
215:
194:
193:
2851:6 September
2758:6 September
2599:PsittaScene
2238:), grison (
2232:), olingo (
2220:), ocelot (
2214:), margay (
2196:), jaguar (
2137:, the paca
1999:Galliformes
1992:S. tyrannus
1923:crested owl
1866:harpy eagle
1732:hummingbird
1710:parakeets,
1671:A. farinosa
1620:D. insignis
1608:N. sericeus
1590:N. fungosus
1470:Sibon argus
1399:E. gollmeri
1341:A. saltator
1317:H. miliaria
1295:O. collaris
1292:(endemic),
1286:(endemic),
1277:B. colonnea
1261:salamanders
1213:glass frogs
1205:, the toad
936:, the palm
677:. The palm
649:, the bush
172:22 May 2010
128:Established
107: /
83:Coordinates
2866:Categories
2605:2013-01-30
2426:(2): 43–49
2365:References
2007:Crax rubra
1977:crane hawk
1939:snail kite
1892:) and the
1800:) and the
1759:sunbittern
1536:T. viquezi
1423:E. rugosus
1366:including
1364:rain frogs
1306:tree frogs
1250:D. pumilio
1192:caecilians
1186:Amphibians
1155:) and the
1112:) and the
1104:pez gaspar
818:herbaceous
377:palustrine
207:Costa Rica
169:Designated
63:Costa Rica
2822:30 August
2728:26 August
2723:1659-0732
2711:Zeledonia
2681:21 August
2655:30 August
2572:21 August
2536:18 August
2430:25 August
2355:plantains
2344:ecolodges
2250:, tapir (
2202:), puma (
2033:Passerine
1953:tiny hawk
1530:T. simile
1502:annulatus
1481:pit viper
1411:E. noblei
921:L. belloi
703:Inga alba
355:epiphytes
326:in 2005.
246:Nicaragua
2752:Anywhere
2469:25 April
2272:and the
2135:manatees
1828:tinamous
1658:and the
1643:sungrebe
1515:include
1479:and the
1430:Reptiles
1405:E. mimus
1347:A. lemur
1157:chiclids
780:and the
430:almendro
345:Habitats
252:and the
240:and the
59:Location
2359:cassava
2157:opossum
2129:Mammals
2083:); the
1864:), the
1777:), the
1684:salvini
1649:), the
1551:), the
1375:E. andi
1129:endemic
1114:sawfish
525:conifer
507:, teak
432:trees (
383:Species
301:Spanish
293:Spanish
285:Spanish
277:Spanish
271:History
224:in the
199:Spanish
95:84°09′W
92:10°42′N
2721:
2677:. 2019
2462:Ramsar
2285:, the
2117:) and
2075:) and
1913:, the
1894:falcon
1779:trogon
1730:, the
1651:motmot
1574:anoles
1442:caiman
1259:, the
521:acacia
232:, the
2816:(PDF)
2649:(PDF)
2595:(PDF)
2524:(1).
2518:PARKS
2514:(PDF)
2494:6 May
2390:6 May
2385:SINAC
1840:ssp.
1682:ssp.
1627:Birds
1553:gecko
1500:ssp.
1063:Fauna
388:Flora
265:Pital
141:SINAC
77:Pital
2853:2019
2824:2019
2760:2019
2730:2019
2719:ISSN
2683:2019
2657:2019
2601:(53)
2574:2019
2552:link
2538:2019
2496:2020
2471:2018
2432:2019
2392:2020
2357:and
2262:and
2103:and
2023:and
1989:and
1907:and
1835:and
1724:and
1668:and
1617:and
1533:and
1338:and
1298:and
1268:and
1253:and
1226:and
1199:and
1176:and
1143:and
1121:and
1068:Fish
1055:and
999:and
983:and
960:and
950:and
932:and
918:and
892:and
871:and
857:and
831:and
795:and
714:and
685:and
671:and
643:and
568:and
550:and
513:and
495:and
476:taxa
408:and
373:lake
263:and
220:and
180:1918
131:2005
120:Area
2526:doi
2346:,.
2334:).
2091:),
2059:),
2051:),
2043:),
1975:),
1967:),
1959:),
1937:),
1929:),
1921:),
1757:),
1704:),
1511:).
1078:),
446:is
244:in
2868::
2843:.
2832:^
2768:^
2750:.
2738:^
2715:15
2709:.
2691:^
2673:.
2614:^
2597:.
2582:^
2548:}}
2544:{{
2522:18
2520:.
2516:.
2487:.
2459:.
2440:^
2424:11
2418:.
2400:^
2383:.
2372:^
2353:,
2300:,
2097:,
2029:.
1995:.
1951:,
1884:,
1851:.
1641:,
1623:.
1611:,
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1587:,
1581:,
1566:,
1560:,
1527:,
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1408:,
1390:,
1350:,
1320:,
1314:,
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1220:,
1182:.
1170:,
1164:,
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1033:,
1029:,
1025:,
1019:,
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912:,
908:,
904:,
900:,
886:,
882:,
875:.
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851:,
847:,
809:.
789:,
774:,
768:,
762:,
756:,
750:,
744:,
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732:,
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710:,
706:,
700:,
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632:.
620:,
614:,
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596:,
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562:,
544:,
538:,
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368:.
341:.
303::
295::
287::
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2528::
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2013:(
2005:(
1979:(
1971:(
1963:(
1955:(
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1933:(
1925:(
1917:(
1888:(
1868:(
1804:(
1796:(
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1678:(
1645:(
197:(
143:)
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