42:
547:
70:
1265:
564:
1275:
581:
186:
384:
The
Melolonthinae life cycle has the four stages of egg, larva, pupa and adult, similar to other beetles. Eggs are laid in soil, which is also where the larval and pupal stages occur. Adults occur above ground. The larval stage is long and may last up to two years, whereas the adult stage is short
584:
588:
587:
583:
582:
589:
586:
367:
Larvae are C-shaped with a white/cream body and a darker, well-sclerotised head. They can be distinguished from other scarabaeid larvae by the galea and lacinia either partly fused proximally or fitting tightly together, the
1286:
585:
1587:
1471:
356:, each with 7 to 10 segments, of which the last 3 to 7 segments are elongate and form a club, and the antennal bases are usually concealed from above. The
740:
1229:
1574:
1287:
https://web.archive.org/web/20161025112710/http://bug.tamu.edu/research/collection/hallan/test/Arthropoda/Insects/Coleoptera/Family/Scarabaeidae.txt
1445:
340:
Melolonthinae adults range from 3 to 58 mm in length and are usually brown or black in colour. Some species are shiny, while many are covered in
364:. The margins of the elytra are straight instead of strongly curved. Each leg ends in a pair of tarsal claws that are often toothed or double.
41:
488:
In feeding species, adults gather on trees and this helps them find each other to mate. In non-feeding species, virgin females emit a
376:
areas, the apical segment of the antenna about as wide as its penultimate segment, and the anal cleft usually Y-shaped or angulate.
1450:
1256:
Arrow (Coleoptera: Scarabaeidae): redescription and removal from the
Aclopinae, with systematic notes. Zootaxa, 1916: 57–62.
1600:
1458:
69:
1646:
1344:
369:
349:
1411:
1553:
357:
1566:
1476:
1424:
1279:
1120:
Cave, R.D. & Ratcliffe, B.C. (2008). Scarab beetles (Coleoptera: Scarabaeoidea). In J.F. Capinera (Ed.),
1641:
1297:
510:
345:
191:
1507:
959:
227:
Scarabaeidae). It is a very diverse group; distributed over most of the world, it contains over 11,000
1486:
1393:
876:
642:
634:
573:
247:
51:
700:
868:
726:
683:
652:
1058:
1042:
935:
546:
236:
1335:
1269:
1000:
927:
885:
783:
708:
626:
551:
470:
441:
435:
166:
64:
1094:
1066:
1050:
1008:
672:
617:
447:
429:
411:
313:
1512:
1100:
902:
757:
1613:
1494:
1380:
1076:
1018:
982:
353:
263:
259:
203:
200:
1618:
1592:
1499:
1082:
453:
224:
601:
309:
266:
is particularly pronounced. Many species have striking – though rarely brilliant or
240:
262:, while those of the females are smaller and somewhat knobby. In the Melolonthinae, this
1561:
1548:
1419:
1406:
975:
774:
692:
352:, a structure on the head above the mouthparts, is not armed with teeth. There are two
321:
1635:
1349:
1198:
489:
273:
Being often quite sizeable and swarming in numbers at certain times, for example the
255:
969:
Several genera are of unclear relations; they are not yet firmly placed in a tribe:
600:
According to various authors, the living
Melolonthinae are divided into about 20–30
308:– feature widely in folklore. Some Melolonthinae are economically significant
943:
791:
474:
417:
373:
325:
305:
217:
196:
152:
131:
56:
563:
1539:
1385:
1605:
1463:
1432:
1329:
1088:
955:
893:
852:
837:
822:
815:
801:
765:
717:
317:
292:
281:
275:
1264:
1034:
992:
845:
829:
662:
568:
482:
478:
301:
297:
287:
267:
232:
1320:
17:
1163:
860:
735:
535:
423:
406:
258:
in outward appearance. Like in many
Scarabaeidae, males have large fingered
251:
213:
101:
81:
1274:
1533:
1367:
1314:
1026:
919:
910:
808:
749:
556:
458:
329:
1359:
1230:"Guide to New World Scarab Beetles-Scarabaeidae-Melolonthinae overview"
609:
527:
505:
398:
361:
228:
1372:
1134:
1437:
963:
605:
402:
220:
121:
111:
91:
1579:
1291:
185:
579:
562:
545:
523:
394:
184:
1398:
341:
231:
in over 750 genera. Some authors include the scarab subfamilies
1295:
312:. Other than the Melolonthini, the most diverse tribes are the
457:). Adults that feed do so on the leaves of trees, such as
477:, but the flower- and pollen-feeding species are often
1135:"Subfamily Melolonthinae - May Beetles and Junebugs"
1523:
1304:
1252:Ocampo, F.C., Vaz-de-Mello, F.Z. (2008) The genus
534:spp. can sometimes cause complete defoliation of
648:– including Stethaspini and Xylonichini
393:Larval melolonthines feed on plant roots and
8:
522:is another sugarcane pest and also attacks
1292:
250:is usually not bizarre. They resemble the
40:
31:
270:– hues and bold patterns of hairs.
771:– including Allarini, Colpochilini
1113:
1086:", both used by Erichson in 1847, are
246:Unlike some of their relatives, their
1124:(pp. 3273–3286. Heidelberg. Springer.
7:
1567:b4ce37f7-e885-4fe0-ab40-aecbf6013375
1487:70817595-f753-46ac-8aef-c0e680240c3c
1425:e4ac69dd-3125-415e-9887-8c43b6eb6164
1224:
1222:
1220:
1218:
1193:
1191:
1189:
1187:
1185:
1183:
1158:
1156:
1154:
385:and lasts only a few days or weeks.
714:– sometimes in Macrodactylini
25:
723:– sometimes in Melolonthini
1273:
1263:
780:– including Dichelonyciini
159:
68:
797:– including Rhizotrogini
530:and various fruits. Adults of
360:is exposed at the base of the
1:
1199:"Australian Faunal Directory"
899:– sometimes in Hopliini
833:– cockchafers, May bugs
492:so that males can find them.
949:– includes Pachydemini
397:. Known host plants include
199:), male on top – note
1104:are of uncertain validity.
689:– including Comophini
623:– including Camentini
461:, or on flowers or pollen.
1663:
1122:Encyclopedia of Entomology
172:
165:
158:
151:
65:Scientific classification
63:
48:
39:
34:
1046:– Colymbomorphini?
1022:– Colymbomorphini?
596:in Laos, moving its head
516:Antitrogus and Lepidiota
445:) or may not feed (e.g.
508:pests, while larvae of
304:– all from tribe
612:are also listed here:
597:
577:
560:
511:Dermolepida albohirtum
243:in the Melolonthinae.
206:
192:Rhizotrogus marginipes
1508:Paleobiology Database
1282:at Wikimedia Commons
962:, is only known from
592:
566:
549:
518:are sugarcane pests.
188:
1562:Fauna Europaea (new)
1420:Fauna Europaea (new)
1062:– Liparetrini?
1054:– Liparetrini?
1038:– Liparetrini?
1030:– Liparetrini?
1004:– Pachydemini?
996:– Liparetrini?
27:Subfamily of beetles
1203:biodiversity.org.au
841:– May beetles
469:Adults are usually
409:. Adults may (e.g.
1647:Beetle subfamilies
598:
578:
561:
483:attracted to light
207:
201:sexually dimorphic
160:About 20–30 tribes
1629:
1628:
1614:Open Tree of Life
1495:Open Tree of Life
1298:Taxon identifiers
1278:Media related to
1234:unsm-ento.unl.edu
1014:
988:
960:Cretomelolonthini
948:
940:
932:
915:
914:Dalla Torre, 1912
907:
898:
881:
873:
865:
796:
788:
779:
770:
762:
754:
744:
731:
722:
713:
705:
697:
688:
678:
668:
658:
647:
639:
631:
622:
590:
481:. They are often
264:sexual dimorphism
183:
182:
147:
16:(Redirected from
1654:
1622:
1621:
1609:
1608:
1606:NHMSYS0020959392
1596:
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1489:
1480:
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1467:
1466:
1464:NHMSYS0001718975
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1428:
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1402:
1401:
1389:
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1363:
1362:
1353:
1352:
1340:
1339:
1338:
1325:
1324:
1323:
1293:
1277:
1268:Data related to
1267:
1257:
1250:
1244:
1243:
1241:
1240:
1226:
1213:
1212:
1210:
1209:
1195:
1178:
1177:
1175:
1174:
1160:
1149:
1148:
1146:
1145:
1131:
1125:
1118:
1012:
986:
946:
938:
931:Burmeister, 1855
930:
913:
905:
896:
879:
872:Burmeister, 1855
871:
863:
794:
787:Burmeister, 1855
786:
777:
769:Burmeister, 1855
768:
761:Burmeister, 1844
760:
752:
741:Guérin-Méneville
739:
729:
720:
711:
703:
695:
686:
676:
666:
656:
645:
638:Lacordaire, 1856
637:
629:
621:Burmeister, 1855
620:
591:
520:Lepidiota stigma
254:in being fairly
178:Systellopodinae
145:
73:
72:
44:
32:
21:
1662:
1661:
1657:
1656:
1655:
1653:
1652:
1651:
1632:
1631:
1630:
1625:
1617:
1612:
1604:
1599:
1591:
1586:
1578:
1573:
1565:
1560:
1552:
1547:
1538:
1537:
1532:
1519:
1511:
1506:
1498:
1493:
1485:
1483:
1475:
1470:
1462:
1457:
1449:
1444:
1436:
1431:
1423:
1418:
1410:
1405:
1397:
1392:
1384:
1379:
1371:
1366:
1358:
1356:
1348:
1343:
1334:
1333:
1328:
1319:
1318:
1313:
1300:
1272:at Wikispecies
1261:
1260:
1251:
1247:
1238:
1236:
1228:
1227:
1216:
1207:
1205:
1197:
1196:
1181:
1172:
1170:
1164:"Melolonthinae"
1162:
1161:
1152:
1143:
1141:
1133:
1132:
1128:
1119:
1115:
1110:
1072:
954:In addition, a
952:
877:Phyllotocidiini
753:Latreille, 1829
696:Burmeister 1855
677:Boisduval, 1835
657:Blanchard, 1850
646:Blanchard, 1850
643:Colymbomorphini
635:Chasmatopterini
604:. Some notable
580:
574:Colymbomorphini
544:
498:
467:
391:
382:
338:
177:
175:
144:
67:
52:Polyphylla alba
28:
23:
22:
15:
12:
11:
5:
1660:
1658:
1650:
1649:
1644:
1634:
1633:
1627:
1626:
1624:
1623:
1610:
1597:
1584:
1571:
1558:
1549:Fauna Europaea
1545:
1529:
1527:
1521:
1520:
1518:
1517:
1504:
1491:
1481:
1468:
1455:
1442:
1429:
1416:
1407:Fauna Europaea
1403:
1390:
1377:
1364:
1354:
1341:
1326:
1310:
1308:
1302:
1301:
1296:
1290:
1289:
1259:
1258:
1245:
1214:
1179:
1150:
1126:
1112:
1111:
1109:
1106:
1071:
1070:
1063:
1055:
1047:
1039:
1031:
1023:
1015:
1013:Westwood, 1842
1005:
997:
989:
979:
971:
951:
950:
947:Erichson, 1847
941:
933:
925:
924:
923:
908:
900:
891:
890:
889:
874:
866:
858:
857:
856:
849:
842:
834:
826:
819:
812:
805:
789:
781:
775:Macrodactylini
772:
763:
755:
747:
746:
745:
724:
715:
712:Medvedev, 1952
706:
701:Diphucephalini
698:
693:Dichelonychini
690:
681:
680:
679:
669:
659:
640:
632:
624:
614:
543:
540:
497:
494:
466:
463:
390:
387:
381:
378:
337:
334:
322:Macrodactylini
181:
180:
170:
169:
163:
162:
156:
155:
149:
148:
139:
135:
134:
129:
125:
124:
119:
115:
114:
109:
105:
104:
99:
95:
94:
89:
85:
84:
79:
75:
74:
61:
60:
46:
45:
37:
36:
35:Melolonthinae
26:
24:
14:
13:
10:
9:
6:
4:
3:
2:
1659:
1648:
1645:
1643:
1642:Melolonthinae
1640:
1639:
1637:
1620:
1615:
1611:
1607:
1602:
1598:
1594:
1589:
1585:
1581:
1576:
1572:
1568:
1563:
1559:
1555:
1550:
1546:
1541:
1535:
1531:
1530:
1528:
1526:
1525:Melolonthidae
1522:
1514:
1509:
1505:
1501:
1496:
1492:
1488:
1482:
1478:
1473:
1469:
1465:
1460:
1456:
1452:
1447:
1443:
1439:
1434:
1430:
1426:
1421:
1417:
1413:
1408:
1404:
1400:
1395:
1391:
1387:
1382:
1378:
1374:
1369:
1365:
1361:
1355:
1351:
1350:Melolonthinae
1346:
1342:
1337:
1336:Melolonthinae
1331:
1327:
1322:
1316:
1312:
1311:
1309:
1307:
1306:Melolonthinae
1303:
1299:
1294:
1288:
1285:
1284:
1283:
1281:
1280:Melolonthinae
1276:
1271:
1270:Melolonthinae
1266:
1255:
1249:
1246:
1235:
1231:
1225:
1223:
1221:
1219:
1215:
1204:
1200:
1194:
1192:
1190:
1188:
1186:
1184:
1180:
1169:
1165:
1159:
1157:
1155:
1151:
1140:
1136:
1130:
1127:
1123:
1117:
1114:
1107:
1105:
1103:
1102:
1097:
1096:
1091:
1090:
1085:
1084:
1079:
1078:
1069:
1068:
1064:
1061:
1060:
1056:
1053:
1052:
1048:
1045:
1044:
1040:
1037:
1036:
1032:
1029:
1028:
1024:
1021:
1020:
1016:
1011:
1010:
1006:
1003:
1002:
998:
995:
994:
990:
985:
984:
980:
978:
977:
973:
972:
970:
967:
965:
961:
957:
945:
942:
937:
934:
929:
926:
922:
921:
917:
916:
912:
909:
906:Britton, 1957
904:
901:
895:
892:
888:
887:
883:
882:
880:Britton, 1957
878:
875:
870:
869:Pachytrichini
867:
864:LeConte, 1861
862:
859:
855:
854:
850:
848:
847:
843:
840:
839:
835:
832:
831:
827:
825:
824:
820:
818:
817:
813:
811:
810:
806:
804:
803:
799:
798:
793:
790:
785:
782:
776:
773:
767:
764:
759:
756:
751:
748:
742:
738:
737:
733:
732:
730:Britton, 1957
728:
727:Heteronychini
725:
719:
716:
710:
707:
704:Britton, 1957
702:
699:
694:
691:
687:Britton, 1957
685:
684:Comophorinini
682:
675:
674:
670:
665:
664:
660:
655:
654:
653:Colymbomorpha
650:
649:
644:
641:
636:
633:
630:Britton, 1978
628:
625:
619:
616:
615:
613:
611:
607:
603:
595:
575:
571:
570:
565:
558:
554:
553:
548:
541:
539:
537:
533:
529:
525:
521:
517:
513:
512:
507:
503:
495:
493:
491:
490:sex pheromone
486:
484:
480:
476:
472:
464:
462:
460:
456:
455:
450:
449:
444:
443:
438:
437:
432:
431:
426:
425:
420:
419:
414:
413:
408:
404:
400:
396:
388:
386:
379:
377:
375:
371:
365:
363:
359:
355:
351:
347:
343:
335:
333:
331:
327:
323:
319:
315:
311:
307:
303:
300:
299:
294:
290:
289:
284:
283:
278:
277:
271:
269:
265:
261:
257:
256:plesiomorphic
253:
249:
244:
242:
238:
234:
230:
226:
222:
219:
215:
211:
210:Melolonthinae
205:
202:
198:
194:
193:
187:
179:
176:Melolonthidae
171:
168:
164:
161:
157:
154:
150:
143:
142:Melolonthinae
140:
137:
136:
133:
130:
127:
126:
123:
120:
117:
116:
113:
110:
107:
106:
103:
100:
97:
96:
93:
90:
87:
86:
83:
80:
77:
76:
71:
66:
62:
58:
54:
53:
47:
43:
38:
33:
30:
19:
18:Melolonthidae
1524:
1305:
1262:
1253:
1248:
1237:. Retrieved
1233:
1206:. Retrieved
1202:
1171:. Retrieved
1167:
1142:. Retrieved
1139:bugguide.net
1138:
1129:
1121:
1116:
1099:
1093:
1087:
1081:
1075:
1073:
1065:
1059:Sericospilus
1057:
1049:
1043:Psilodontria
1041:
1033:
1025:
1017:
1007:
999:
991:
987:Fuavel, 1903
981:
974:
968:
953:
944:Tanyproctini
936:Systellopini
918:
897:Howden, 1997
884:
851:
844:
836:
828:
821:
814:
807:
800:
792:Melolonthini
734:
671:
661:
651:
599:
594:Leucopholina
593:
567:
550:
531:
519:
515:
509:
501:
499:
487:
468:
452:
446:
440:
434:
428:
422:
418:Diphucephala
416:
410:
392:
383:
374:stridulatory
366:
339:
326:Tanyproctini
306:Melolonthini
296:
293:June beetles
286:
280:
274:
272:
245:
237:Pachypodinae
209:
208:
197:Melolonthini
190:
173:
141:
132:Scarabaeidae
57:Melolonthini
50:
29:
1433:iNaturalist
1330:Wikispecies
1089:nomina nuda
1001:Hemictenius
958:tribe, the
956:prehistoric
939:Sharp, 1877
928:Sericoidini
894:Podolasiini
886:Phyllotocus
853:Rhizotrogus
838:Phyllophaga
823:Leucopholis
816:Holotrichia
802:Amphimallon
795:Leach, 1819
784:Maechidiini
778:Kirby, 1837
766:Liparetrini
721:Kirby, 1837
718:Diplotaxini
709:Diphycerini
627:Automoliini
552:Phyllotocus
542:Systematics
532:Phyllophaga
502:Sericesthis
471:crepuscular
442:Sericesthis
436:Phyllotocus
336:Description
318:Liparetrini
302:cockchafers
282:Phyllophaga
276:Amphimallon
146:Leach, 1819
138:Subfamily:
1636:Categories
1254:Xenaclopus
1239:2023-01-18
1208:2023-01-18
1173:2023-01-18
1168:uk beetles
1144:2023-01-18
1108:References
1095:Holophylla
1067:Xenaclopus
1051:Scythrodes
1035:Prodontria
1009:Metascelis
993:Costelytra
846:Polyphylla
830:Melolontha
673:Xylonichus
667:Hope, 1837
663:Stethaspis
618:Ablaberini
569:Stethaspis
500:Larvae of
448:Antitrogus
430:Liparetrus
412:Automolius
380:Life cycle
314:Ablaberini
298:Melolontha
288:Polyphylla
268:iridescent
233:Euchirinae
122:Coleoptera
102:Arthropoda
1540:Q14852127
1101:Hoplorida
903:Scitalini
758:Lichniini
736:Heteronyx
536:deciduous
504:spp. are
475:nocturnal
465:Behaviour
459:eucalypts
424:Heteronyx
407:sugarcane
370:mandibles
358:scutellum
295:" or the
252:Rutelinae
214:subfamily
174:Hopliinae
153:Diversity
88:Kingdom:
82:Eukaryota
1534:Wikidata
1399:63741253
1368:BugGuide
1357:BioLib:
1315:Wikidata
1077:Anonetus
1027:Odontria
1019:Mycernus
983:Conebius
920:Maladera
911:Sericini
861:Oncerini
809:Brahmina
750:Hopliini
557:Sericini
372:lacking
354:antennae
330:Sericini
260:antennae
204:antennae
167:Synonyms
128:Family:
98:Phylum:
92:Animalia
78:Domain:
1619:7175370
1500:7175370
1321:Q785080
1083:Tryssus
1080:" and "
964:fossils
610:species
538:trees.
528:sorghum
506:pasture
479:diurnal
454:Rhopaea
399:grasses
350:clypeus
248:habitus
229:species
221:beetles
216:of the
189:Mating
118:Order:
112:Insecta
108:Class:
1593:117613
1484:NZOR:
1451:678508
1412:257338
743:, 1838
606:genera
602:tribes
403:clover
362:elytra
348:. The
346:scales
241:tribes
225:family
218:scarab
1588:IRMNG
1554:11130
1513:70051
1438:48200
976:Acoma
572:sp. (
555:sp. (
496:Pests
395:humus
342:setae
310:pests
212:is a
49:Male
1580:5593
1575:GBIF
1477:7059
1472:NCBI
1446:ITIS
1373:2951
1360:8080
1098:and
608:and
524:corn
451:and
439:and
405:and
389:Diet
328:and
285:and
235:and
1601:NBN
1459:NBN
1394:EoL
1386:JW6
1381:CoL
1345:AFD
473:or
344:or
239:as
1638::
1616::
1603::
1590::
1577::
1564::
1551::
1536::
1510::
1497::
1474::
1461::
1448::
1435::
1422::
1409::
1396::
1383::
1370::
1347::
1332::
1317::
1232:.
1217:^
1201:.
1182:^
1166:.
1153:^
1137:.
1092:.
966:.
526:,
514:,
485:.
433:,
427:,
421:,
415:,
401:,
332:.
324:,
320:,
316:,
279:,
59:)
1242:.
1211:.
1176:.
1147:.
1074:"
576:)
559:)
291:"
223:(
195:(
55:(
20:)
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