1074:
53:
1786:
1839:
40:
1792:
1779:
1800:
246:
ventral flange of the frontal meeting the sphenethmoid; (7) an anterolaterally bulbous sphenethmoid; and (8) a massive epipterygoid. Like other ostodolepids, it has a wedge-shaped skull, a pointed snout, and a temporal emargination. The skull is highly ossified in order to resist anteroposteriorly directed forces, as indicated by the markedly interdigitated sutures.
245:
is the smallest known ostodolepid and is diagnosed by: (1) the high subtemporal recess that separates the postorbital from the squamosal; (2) four premaxillary tooth positions; (3) 12 maxillary tooth positions; circumorbital walls formed by medial laminae of the prefrontal, lacrimal, and jugal; (5) a
1037:
Huttenlocker, Adam K.; Pardo, Jason D.; Small, Bryan J.; Anderson, Jason S. (2013). "Cranial morphology of recumbirostrans (Lepospondyli) from the
Permian of Kansas and Nebraska, and early morphological evolution inferred by micro-computed tomography".
262:, or burrowing, lifestyle. More specifically, the animal would likely have used its recumbent snout to push forward into the substrate, facilitated by increased epaxial musculature that inserted into the broad, sloping occiput, as in
270:
was known from only one specimen, it has been suggested that it was either a naturally rare component of the assemblage or that its inferred burrowing ecology did not lend itself well to preservation in the fissure fill system.
1958:
1904:
228:
1891:
1968:
1107:
1010:, a new ostodolepid 'microsaur' (Tetrapoda, Lepospondyli, Recumbirostra) from the Early Permian of Richards Spur (Ft. Sill), Oklahoma".
204:
was named in 2009 by
Canadian paleontologists Jason Anderson, Diane Scott, and Robert Reisz. It was known from only the
52:
1100:
876:
1179:
1785:
1206:
893:
1859:
1791:
760:
1963:
1930:
1778:
1757:
1799:
1842:
1093:
135:
1544:
1079:
1338:
1713:
1553:
1303:
1295:
719:
47:
1935:
1917:
1909:
1697:
1055:
1922:
1686:
1571:
1288:
1047:
1019:
187:
1164:
843:
826:
227:. The specific name is given for Mark McKinzie, who found and donated the specimen to the
1832:
1817:
1671:
1580:
1526:
1505:
1191:
396:
1952:
1827:
1734:
1678:
1646:
1612:
1496:
1432:
435:
217:
209:
190:
183:
110:
30:
1822:
1745:
1664:
1601:
1560:
1421:
1373:
1324:
413:
308:
263:
234:
In 2021 a second specimen was referred, ROMVP 86541, a skull with right lower jaw.
161:
39:
1085:
1051:
1727:
1635:
1628:
1533:
1470:
1463:
1441:
1414:
1364:
1352:
1317:
1310:
1273:
1249:
1218:
1117:
802:
662:
570:
480:
341:
97:
1882:
1706:
1655:
1621:
1587:
1485:
1477:
1449:
1398:
1345:
1069:
743:
679:
587:
537:
221:
1059:
1387:
365:
259:
224:
64:
1876:
1720:
1594:
1519:
1512:
1407:
1380:
1331:
1148:
1142:
1023:
520:
463:
213:
205:
84:
1896:
1136:
74:
1853:
179:
1857:
1129:
1089:
216:. The holotype, a well preserved skull, was found in
1866:
1744:
1696:
1654:
1645:
1611:
1570:
1543:
1495:
1431:
1397:
1363:
1270:
1217:
1190:
1163:
1006:Anderson, J.S.; Scott, D.; Reisz, R.R. (2009). "
254:The high degree of ossification in the skull of
283:in the analysis of Huttenlocker et al. (2013):
1101:
8:
229:Sam Noble Oklahoma Museum of Natural History
1854:
1651:
1280:
1267:
1196:
1169:
1160:
1126:
1108:
1094:
1086:
38:
20:
1001:
999:
995:
1959:Cisuralian amphibians of North America
7:
868:
818:
794:
735:
711:
704:
654:
630:
562:
512:
505:
455:
448:
441:
388:
381:
357:
333:
300:
293:
286:
1040:Journal of Vertebrate Paleontology
1012:Journal of Vertebrate Paleontology
14:
208:specimen, which was found at the
1838:
1837:
1798:
1790:
1784:
1777:
1072:
51:
220:-aged fissure fill deposits in
258:is seen as an adaptation to a
1:
1969:Fossil taxa described in 2009
1052:10.1080/02724634.2013.728998
16:Extinct genus of amphibians
1985:
279:Below is the placement of
1812:
1774:
1283:
1266:
1245:
1199:
1172:
1159:
1125:
890:
877:Cardiocephalus sternbergi
873:
866:
840:
823:
816:
799:
792:
757:
740:
733:
716:
709:
702:
676:
659:
652:
635:
628:
584:
567:
560:
534:
517:
510:
503:
477:
460:
453:
446:
439:
410:
393:
386:
379:
362:
355:
338:
331:
305:
298:
291:
147:
142:
48:Scientific classification
46:
37:
23:
894:Cardiocephalus peabodyi
761:Huskerpeton englehorni
1931:Paleobiology Database
1024:10.1671/039.029.0222
1008:Nannaroter mckinziei
186:tetrapod within the
1080:Paleontology portal
720:Proxilodon bonneri
1946:
1945:
1918:Open Tree of Life
1860:Taxon identifiers
1851:
1850:
1808:
1807:
1772:
1771:
1768:
1767:
1698:Brachystelechidae
1262:
1261:
1258:
1257:
1241:
1240:
1237:
1236:
986:
985:
977:
976:
968:
967:
959:
958:
950:
949:
941:
940:
932:
931:
923:
922:
914:
913:
905:
904:
855:
854:
781:
780:
772:
771:
691:
690:
617:
616:
608:
607:
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598:
549:
548:
492:
491:
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320:
319:
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165:
138:
1976:
1939:
1938:
1926:
1925:
1913:
1912:
1900:
1899:
1887:
1886:
1885:
1855:
1841:
1840:
1803:
1802:
1795:
1794:
1788:
1782:
1781:
1652:
1572:Hapsidopareiidae
1289:Altenglanerpeton
1281:
1268:
1197:
1170:
1161:
1154:
1153:
1127:
1110:
1103:
1096:
1087:
1082:
1077:
1076:
1075:
1064:
1063:
1034:
1028:
1027:
1003:
869:
819:
795:
736:
712:
705:
655:
631:
563:
513:
506:
456:
449:
442:
389:
382:
358:
334:
301:
294:
287:
197:History of study
155:
129:
122:
109:
96:
56:
55:
42:
29:Temporal range:
21:
1984:
1983:
1979:
1978:
1977:
1975:
1974:
1973:
1949:
1948:
1947:
1942:
1934:
1929:
1921:
1916:
1908:
1903:
1895:
1890:
1881:
1880:
1875:
1862:
1852:
1847:
1804:
1797:
1789:
1776:
1764:
1740:
1692:
1641:
1607:
1566:
1539:
1491:
1433:Ostodolepididae
1427:
1393:
1359:
1277:
1254:
1233:
1213:
1186:
1180:Tetrapodomorpha
1165:Tetrapodomorpha
1155:
1132:
1131:
1121:
1114:
1078:
1073:
1071:
1068:
1067:
1036:
1035:
1031:
1005:
1004:
997:
992:
987:
978:
969:
960:
951:
942:
933:
924:
915:
906:
856:
844:Euryodus primus
827:Euryodus dalyae
782:
773:
692:
618:
609:
600:
550:
493:
426:
321:
277:
252:
240:
199:
128:
120:
107:
94:
50:
33:
27:
17:
12:
11:
5:
1982:
1980:
1972:
1971:
1966:
1961:
1951:
1950:
1944:
1943:
1941:
1940:
1927:
1914:
1901:
1888:
1872:
1870:
1864:
1863:
1858:
1849:
1848:
1846:
1845:
1835:
1833:Tuditanomorpha
1830:
1825:
1820:
1818:Batrachomorpha
1813:
1810:
1809:
1806:
1805:
1775:
1773:
1770:
1769:
1766:
1765:
1763:
1762:
1761:
1760:
1751:
1749:
1742:
1741:
1739:
1738:
1731:
1724:
1717:
1710:
1702:
1700:
1694:
1693:
1691:
1690:
1683:
1675:
1672:Infernovenator
1668:
1660:
1658:
1649:
1643:
1642:
1640:
1639:
1632:
1625:
1617:
1615:
1609:
1608:
1606:
1605:
1598:
1591:
1584:
1581:Hapsidopareion
1576:
1574:
1568:
1567:
1565:
1564:
1557:
1549:
1547:
1545:Odonterpetidae
1541:
1540:
1538:
1537:
1530:
1527:Leiocephalikon
1523:
1516:
1509:
1506:Cardiocephalus
1501:
1499:
1493:
1492:
1490:
1489:
1482:
1474:
1467:
1460:
1453:
1446:
1437:
1435:
1429:
1428:
1426:
1425:
1418:
1411:
1403:
1401:
1395:
1394:
1392:
1391:
1384:
1377:
1369:
1367:
1361:
1360:
1358:
1357:
1349:
1342:
1335:
1328:
1321:
1314:
1307:
1300:
1292:
1284:
1278:
1271:
1264:
1263:
1260:
1259:
1256:
1255:
1246:
1243:
1242:
1239:
1238:
1235:
1234:
1232:
1231:
1230:
1229:
1223:
1221:
1215:
1214:
1212:
1211:
1210:
1209:
1207:Reptiliomorpha
1200:
1194:
1192:Reptiliomorpha
1188:
1187:
1185:
1184:
1183:
1182:
1173:
1167:
1157:
1156:
1152:
1151:
1145:
1139:
1130:
1123:
1122:
1115:
1113:
1112:
1105:
1098:
1090:
1084:
1083:
1066:
1065:
1046:(3): 540–552.
1029:
1018:(2): 379–388.
994:
993:
991:
988:
984:
983:
980:
979:
975:
974:
971:
970:
966:
965:
962:
961:
957:
956:
953:
952:
948:
947:
944:
943:
939:
938:
935:
934:
930:
929:
926:
925:
921:
920:
917:
916:
912:
911:
908:
907:
903:
902:
899:
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872:
867:
865:
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839:
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835:
832:
831:
822:
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815:
812:
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583:
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485:
476:
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472:
469:
468:
459:
454:
452:
447:
445:
440:
438:
432:
431:
428:
427:
423:
422:
419:
418:
409:
406:
405:
402:
401:
397:Hapsidopareion
392:
387:
385:
380:
378:
375:
374:
371:
370:
361:
356:
354:
351:
350:
347:
346:
337:
332:
330:
327:
326:
323:
322:
318:
317:
314:
313:
304:
299:
297:
292:
290:
285:
276:
273:
264:amphisbaenians
251:
248:
239:
236:
198:
195:
184:Recumbirostran
178:is an extinct
169:
168:
167:
166:
145:
144:
140:
139:
118:
114:
113:
105:
101:
100:
92:
88:
87:
82:
78:
77:
72:
68:
67:
62:
58:
57:
44:
43:
35:
34:
28:
15:
13:
10:
9:
6:
4:
3:
2:
1981:
1970:
1967:
1965:
1962:
1960:
1957:
1956:
1954:
1937:
1932:
1928:
1924:
1919:
1915:
1911:
1906:
1902:
1898:
1893:
1889:
1884:
1878:
1874:
1873:
1871:
1869:
1865:
1861:
1856:
1844:
1836:
1834:
1831:
1829:
1828:Recumbirostra
1826:
1824:
1821:
1819:
1815:
1814:
1811:
1801:
1796:
1793:
1787:
1780:
1759:
1755:
1754:
1753:
1752:
1750:
1747:
1743:
1737:
1736:
1735:Quasicaecilia
1732:
1730:
1729:
1725:
1723:
1722:
1718:
1716:
1715:
1711:
1709:
1708:
1704:
1703:
1701:
1699:
1695:
1689:
1688:
1684:
1681:
1680:
1679:Megamolgophis
1676:
1674:
1673:
1669:
1667:
1666:
1662:
1661:
1659:
1657:
1653:
1650:
1648:
1647:Chthonosauria
1644:
1638:
1637:
1633:
1631:
1630:
1626:
1624:
1623:
1619:
1618:
1616:
1614:
1613:Rhynchonkidae
1610:
1604:
1603:
1599:
1597:
1596:
1592:
1590:
1589:
1585:
1583:
1582:
1578:
1577:
1575:
1573:
1569:
1563:
1562:
1558:
1556:
1555:
1551:
1550:
1548:
1546:
1542:
1536:
1535:
1531:
1529:
1528:
1524:
1522:
1521:
1517:
1515:
1514:
1510:
1508:
1507:
1503:
1502:
1500:
1498:
1497:Gymnarthridae
1494:
1488:
1487:
1483:
1480:
1479:
1475:
1473:
1472:
1468:
1466:
1465:
1461:
1459:
1458:
1454:
1452:
1451:
1447:
1444:
1443:
1439:
1438:
1436:
1434:
1430:
1424:
1423:
1419:
1417:
1416:
1412:
1410:
1409:
1405:
1404:
1402:
1400:
1396:
1390:
1389:
1385:
1383:
1382:
1378:
1376:
1375:
1371:
1370:
1368:
1366:
1362:
1355:
1354:
1350:
1348:
1347:
1343:
1341:
1340:
1336:
1334:
1333:
1329:
1327:
1326:
1322:
1320:
1319:
1315:
1313:
1312:
1308:
1306:
1305:
1301:
1298:
1297:
1293:
1291:
1290:
1286:
1285:
1282:
1279:
1275:
1269:
1265:
1253:
1251:
1244:
1227:
1226:
1225:
1224:
1222:
1220:
1216:
1208:
1204:
1203:
1202:
1201:
1198:
1195:
1193:
1189:
1181:
1177:
1176:
1175:
1174:
1171:
1168:
1166:
1162:
1158:
1150:
1146:
1144:
1140:
1138:
1134:
1133:
1128:
1124:
1119:
1111:
1106:
1104:
1099:
1097:
1092:
1091:
1088:
1081:
1070:
1061:
1057:
1053:
1049:
1045:
1041:
1033:
1030:
1025:
1021:
1017:
1013:
1009:
1002:
1000:
996:
989:
982:
981:
973:
972:
964:
963:
955:
954:
946:
945:
937:
936:
928:
927:
919:
918:
910:
909:
901:
900:
897:
896:
895:
888:
887:
884:
883:
880:
879:
878:
871:
870:
864:
863:
860:
859:
851:
850:
847:
846:
845:
838:
837:
834:
833:
830:
829:
828:
821:
820:
814:
813:
810:
809:
806:
805:
804:
797:
796:
790:
789:
786:
785:
777:
776:
768:
767:
764:
763:
762:
755:
754:
751:
750:
747:
746:
745:
738:
737:
731:
730:
727:
726:
723:
722:
721:
714:
713:
707:
706:
700:
699:
696:
695:
687:
686:
683:
682:
681:
674:
673:
670:
669:
666:
665:
664:
657:
656:
650:
649:
646:
645:
642:
641:
640:
633:
632:
626:
625:
622:
621:
613:
612:
604:
603:
595:
594:
591:
590:
589:
582:
581:
578:
577:
574:
573:
572:
565:
564:
558:
557:
554:
553:
545:
544:
541:
540:
539:
532:
531:
528:
527:
524:
523:
522:
515:
514:
508:
507:
501:
500:
497:
496:
488:
487:
484:
483:
482:
475:
474:
471:
470:
467:
466:
465:
458:
457:
451:
450:
444:
443:
437:
436:Recumbirostra
434:
433:
430:
429:
421:
420:
417:
416:
415:
408:
407:
404:
403:
400:
399:
398:
391:
390:
384:
383:
377:
376:
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372:
369:
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367:
360:
359:
353:
352:
349:
348:
345:
344:
343:
336:
335:
329:
328:
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324:
316:
315:
312:
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303:
302:
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295:
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275:Relationships
274:
272:
269:
265:
261:
257:
249:
247:
244:
237:
235:
232:
230:
226:
223:
219:
218:early Permian
215:
211:
210:Richards Spur
207:
203:
196:
194:
192:
191:Ostodolepidae
189:
185:
181:
177:
176:
163:
159:
154:
153:
149:
148:
146:
141:
137:
133:
127:
126:
119:
116:
115:
112:
111:Ostodolepidae
106:
103:
102:
99:
93:
90:
89:
86:
83:
80:
79:
76:
73:
70:
69:
66:
63:
60:
59:
54:
49:
45:
41:
36:
32:
31:Early Permian
26:
22:
19:
1964:Ostodolepids
1867:
1823:Lepospondyli
1783:
1758:Lissamphibia
1746:Lissamphibia
1733:
1726:
1719:
1712:
1705:
1685:
1677:
1670:
1665:Brachydectes
1663:
1634:
1627:
1620:
1602:Saxonerpeton
1600:
1593:
1586:
1579:
1561:Odonterpeton
1559:
1554:Joermungandr
1552:
1532:
1525:
1518:
1511:
1504:
1484:
1476:
1469:
1462:
1456:
1455:
1448:
1440:
1422:Trachystegos
1420:
1413:
1406:
1386:
1379:
1372:
1351:
1344:
1339:Steenerpeton
1337:
1330:
1325:Microbrachis
1323:
1316:
1309:
1302:
1294:
1287:
1247:
1147:Superclass:
1043:
1039:
1032:
1015:
1011:
1007:
892:
891:
875:
874:
842:
841:
825:
824:
801:
800:
759:
758:
742:
741:
718:
717:
678:
677:
661:
660:
638:
637:
636:
586:
585:
569:
568:
536:
535:
519:
518:
479:
478:
462:
461:
414:Saxonerpeton
412:
411:
395:
394:
364:
363:
340:
339:
309:Microbrachis
307:
306:
280:
278:
267:
255:
253:
242:
241:
233:
212:locality in
201:
200:
174:
173:
172:
157:
152:N. mckinziei
151:
150:
131:
124:
123:
24:
18:
1728:Diabloroter
1714:Bromerpeton
1636:Rhynchonkos
1629:Dvellecanus
1534:Pariotichus
1471:Pelodosotis
1464:Ostodolepis
1442:Huskerpeton
1415:Stegotretus
1365:Tuditanidae
1353:Utaherpeton
1318:Kirktonecta
1311:Hyloplesion
1304:Cymatorhiza
1296:Archerpeton
1274:Microsauria
1250:Microsauria
1219:Microsauria
1118:Microsauria
803:Pariotichus
663:Pelodosotis
571:Rhynchonkos
481:Stegotretus
342:Asaphestera
98:Microsauria
1953:Categories
1868:Nannaroter
1816:See also:
1707:Batropetes
1656:Lysorophia
1622:Aletrimyti
1588:Llistrofus
1486:Tambaroter
1478:Proxilodon
1457:Nannaroter
1450:Micraroter
1399:Pantylidae
1346:Trihecaton
1228:see below↓
990:References
744:Tambaroter
680:Micraroter
639:Nannaroter
588:Eocaecilia
538:Batropetes
281:Nannaroter
268:Nannaroter
256:Nannaroter
243:Nannaroter
222:Ordovician
202:Nannaroter
175:Nannaroter
125:Nannaroter
25:Nannaroter
1388:Tuditanus
1149:Tetrapoda
1135:Kingdom:
1060:0272-4634
366:Tuditanus
260:fossorial
225:limestone
156:Anderson
130:Anderson
71:Kingdom:
65:Eukaryota
1883:Q6963817
1877:Wikidata
1843:Category
1721:Carrolla
1595:Ricnodon
1520:Euryodus
1513:Elfridia
1408:Pantylus
1381:Crinodon
1332:Sparodus
1143:Chordata
1141:Phylum:
1137:Animalia
521:Carrolla
464:Pantylus
214:Oklahoma
206:holotype
160:, 2009 (
143:Species
104:Family:
85:Chordata
81:Phylum:
75:Animalia
61:Domain:
1923:3621350
1910:1475464
1897:4964736
250:Ecology
238:Anatomy
117:Genus:
91:Order:
1936:150626
1687:Nagini
1058:
188:family
158:et al.
132:et al.
1905:IRMNG
266:. As
180:genus
1892:GBIF
1756:see
1374:Boii
1205:see
1178:see
1056:ISSN
162:type
136:2009
1048:doi
1020:doi
182:of
1955::
1933::
1920::
1907::
1894::
1879::
1054:.
1044:33
1042:.
1016:29
1014:.
998:^
231:.
193:.
134:,
1748:?
1682:?
1481:?
1445:?
1356:?
1299:?
1276:"
1272:"
1252:"
1248:"
1120:"
1116:"
1109:e
1102:t
1095:v
1062:.
1050::
1026:.
1022::
164:)
121:†
108:†
95:†
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