1666:
60:
31:
1144:
601:
Park, Tae-Yoon S.; Nielsen, Morten Lunde; Parry, Luke A.; Sørensen, Martin
Vinther; Lee, Mirinae; Kihm, Ji-Hoon; Ahn, Inhye; Park, Changkun; de Vivo, Giacinto; Smith, M. Paul; Harper, David A. T.; Nielsen, Arne T.; Vinther, Jakob (2024-01-05).
342:(i.e. streamlined body; small head with ovoid sclerites; well-developed swimming flaps; a pair of caudal furcae). Another distinctive features only known in amplectobeluid genera were pairs of gnathobase-like structures (known by
705:
1665:
1745:
1145:"A new radiodontan oral cone with a unique combination of anatomical features from the early Cambrian Guanshan Lagerstätte, eastern Yunnan, South China"
1732:
895:
Cong P, Ma X, Hou X, Edgecombe GD, Strausfeld NJ (September 2014). "Brain structure resolves the segmental affinity of anomalocaridid appendages".
1804:
1313:
1095:"New radiodonts with gnathobase-like structures from the Cambrian Chengjiang biota and implications for the systematics of Radiodonta"
324:-like structure that presumably better suited for a grasping function. Complete body fossils of amplectobeluids are only known by
741:
59:
659:, a complete radiodont from the early Cambrian Chengjiang Lagerstätte and its implications for the phylogeny of Radiodonta"
839:
Chen JY, Ramsköld L, Zhou GQ (May 1994). "Evidence for monophyly and arthropod affinity of
Cambrian giant predators".
320:
Amplectobeluids could be recognized by frontal appendages with well-developed first distal endite, which forming a
1306:
1590:
1046:"A new hurdiid radiodont from the Burgess Shale evinces the exploitation of Cambrian infaunal food sources"
1685:
1771:
1723:
1241:"The soft-bodied biota of the Cambrian Series 2 Parker Quarry Lagerstätte of northwestern Vermont, USA"
1094:
417:-weighted phylogenetic analysis in 2014 found the following relationships within the Amplectobeluidae:
995:"The functional head of the Cambrian radiodontan (stem-group Euarthropoda) Amplectobelua symbrachiata"
402:
1252:
1156:
904:
848:
760:
1799:
1646:
1299:
1278:
1180:
1125:
928:
872:
784:
686:
54:
1776:
1758:
1270:
1221:
1172:
1117:
1075:
1026:
972:
920:
864:
776:
678:
625:
352:), or an oral cone with combination of tetraradial arrangement and scale-like nodes (known by
1763:
1750:
1630:
1508:
1425:
1260:
1211:
1164:
1109:
1065:
1057:
1016:
1006:
962:
912:
856:
819:
768:
717:
670:
633:
615:
30:
1414:
951:"Origin of raptorial feeding in juvenile euarthropods revealed by a Cambrian radiodontan"
1256:
1198:
Jiao DG, Pates S, Lerosey-Aubril R, Ortega-Hernández J, Yang J, Lan T, Zhang XG (2021).
1160:
908:
852:
764:
638:
1400:
1070:
1045:
1021:
994:
448:
1793:
1611:
1483:
1476:
1384:
1346:
1282:
1184:
788:
690:
603:
546:
529:
481:
390:
374:
326:
321:
290:
263:
258:
240:
169:
159:
39:
1129:
808:
from the
Cambrian Emu Bay Shale and a reassessment of its inferred predatory habits"
1639:
1618:
1597:
1562:
1555:
1524:
1517:
1434:
1392:
932:
876:
338:
302:
296:
104:
860:
1708:
1717:
1604:
1548:
1497:
1449:
1407:
949:
Liu J, Lerosey-Aubril R, Steiner M, Dunlop JA, Shu D, Paterson JR (2018-11-01).
348:
276:, but that genus has become problematic in terms of its taxonomic placement.
252:
200:
401:. Indeterminate frontal appendages assignable to this group are known from the
1370:
1355:
1352:
1322:
1200:"The endemic radiodonts of the Cambrian Stage 4 Guanshan biota of South China"
1011:
235:
117:
1274:
1225:
1176:
1121:
976:
682:
629:
1576:
1535:
1490:
1340:
505:
384:
332:
267:
246:
190:
91:
71:
45:
1216:
1199:
1079:
1061:
1030:
924:
868:
780:
620:
1702:
1456:
1441:
1265:
1240:
967:
950:
745:
272:
232:
210:
179:
1168:
916:
802:
Daley AC, Paterson JR, Edgecombe GD, GarcĂa-Bellido DC, Jago JB (2013).
772:
721:
674:
1737:
1583:
824:
803:
1113:
654:
1569:
1334:
706:"A large pelagic lobopodian from the Cambrian Pioche Shale of Nevada"
308:
81:
1679:
1239:
Pari, Giovanni; Briggs, Derek E.G.; Gaines, Robert R. (2022-02-16).
228:
1093:
Cong PY, Edgecombe GD, Daley AC, Guo J, Pates S, Hou XG (2018).
1683:
1295:
1291:
746:"A suspension-feeding anomalocarid from the Early Cambrian"
262:
and a currently unnamed genus from the lower
Cambrian aged
993:
Cong P, Daley AC, Edgecombe GD, Hou X (August 2017).
653:
Zeng, Han; Zhao, Fangchen; Zhu, Maoyan (2022-09-07).
336:, both showing combination of characters resembling
206:
196:
186:
175:
165:
155:
1692:
1629:
1534:
1507:
1466:
1424:
1368:
388:ignored that assessment and created a genus within
288:was defined as the most inclusive clade including
238:. It currently includes five definitive genera,
382:. Later that year, however, the discoverers of
1307:
735:
733:
731:
8:
1680:
1540:
1376:
1365:
1314:
1300:
1292:
890:
888:
886:
29:
20:
1264:
1215:
1069:
1020:
1010:
966:
823:
637:
619:
270:. There is also a potential fifth genus,
593:
704:McCall, Christian (13 December 2023).
1143:Zeng H, Zhao F, Yin Z, Zhu M (2018).
7:
988:
986:
944:
942:
1044:Moysiuk J, Caron JB (August 2019).
521:
497:
473:
427:
420:
16:Extinct clade of Cambrian organisms
14:
740:Vinther J, Stein M, Longrich NR,
663:Journal of the Geological Society
1664:
1050:Proceedings. Biological Sciences
604:"A giant stem-group chaetognath"
58:
804:"New anatomical information on
1805:Prehistoric arthropod families
1:
1204:Acta Palaeontologica Polonica
861:10.1126/science.264.5163.1304
372:was tentatively assigned to
482:"Anomalocaris" kunmingensis
370:"Anomalocaris" kunmingensis
1821:
547:Amplectobelua stephenensis
530:Amplectobelua symbrachiata
291:Amplectobelua symbrachiata
1662:
1543:
1379:
1364:
1329:
1012:10.1186/s12862-017-1049-1
543:
526:
519:
502:
495:
478:
471:
445:
432:
425:
152:
147:
55:Scientific classification
53:
37:
28:
23:
999:BMC Evolutionary Biology
1245:Journal of Paleontology
1149:Journal of Paleontology
1102:Papers in Palaeontology
955:National Science Review
710:Journal of Paleontology
297:Anomalocaris canadensis
1217:10.4202/app.00870.2020
1062:10.1098/rspb.2019.1079
621:10.1126/sciadv.adi6678
1772:Paleobiology Database
506:Lyrarapax unguispinus
385:Lyrarapax unguispinus
303:Tamisiocaris borealis
1266:10.1017/jpa.2021.125
449:"Anomalocaris" saron
1647:Pseudoangustidontus
1257:2022JPal...96..770P
1169:10.1017/jpa.2017.77
1161:2018JPal...92...40Z
917:10.1038/nature13486
909:2014Natur.513..538C
853:1994Sci...264.1304C
773:10.1038/nature13010
765:2014Natur.507..496V
722:10.1017/jpa.2023.63
675:10.1144/jgs2021-164
1056:(1908): 20191079.
968:10.1093/nsr/nwy057
825:10.1111/pala.12029
1787:
1786:
1759:Open Tree of Life
1686:Taxon identifiers
1677:
1676:
1673:
1672:
1660:
1659:
1656:
1655:
1114:10.1002/spp2.1219
585:
584:
576:
575:
567:
566:
558:
557:
460:
459:
220:
219:
143:
24:Amplectobeluidae
1812:
1780:
1779:
1767:
1766:
1754:
1753:
1741:
1740:
1728:
1727:
1726:
1724:Amplectobeluidae
1713:
1712:
1711:
1694:Amplectobeluidae
1681:
1668:
1631:Aegirocassisinae
1541:
1509:Tamisiocarididae
1468:Amplectobeluidae
1426:Anomalocarididae
1377:
1366:
1316:
1309:
1302:
1293:
1287:
1286:
1268:
1236:
1230:
1229:
1219:
1195:
1189:
1188:
1140:
1134:
1133:
1099:
1090:
1084:
1083:
1073:
1041:
1035:
1034:
1024:
1014:
990:
981:
980:
970:
946:
937:
936:
903:(7519): 538–42.
892:
881:
880:
847:(5163): 1304–8.
836:
830:
829:
827:
799:
793:
792:
750:
737:
726:
725:
716:(5): 1009–1024.
701:
695:
694:
650:
644:
643:
641:
623:
608:Science Advances
598:
522:
498:
474:
428:
421:
403:Parker Formation
286:Amplectobeluidae
224:Amplectobeluidae
208:
198:
188:
177:
167:
157:
137:
134:Amplectobeluidae
132:
116:
103:
63:
62:
33:
21:
1820:
1819:
1815:
1814:
1813:
1811:
1810:
1809:
1790:
1789:
1788:
1783:
1775:
1770:
1762:
1757:
1749:
1744:
1736:
1731:
1722:
1721:
1716:
1707:
1706:
1701:
1688:
1678:
1669:
1652:
1625:
1530:
1503:
1462:
1420:
1415:Paranomalocaris
1373:
1360:
1325:
1320:
1290:
1238:
1237:
1233:
1197:
1196:
1192:
1142:
1141:
1137:
1097:
1092:
1091:
1087:
1043:
1042:
1038:
992:
991:
984:
948:
947:
940:
894:
893:
884:
838:
837:
833:
801:
800:
796:
759:(7493): 496–9.
748:
739:
738:
729:
703:
702:
698:
652:
651:
647:
600:
599:
595:
591:
586:
577:
568:
559:
461:
411:
368:Early in 2014,
366:
318:
309:Hurdia victoria
282:
136:
130:
114:
101:
57:
17:
12:
11:
5:
1818:
1816:
1808:
1807:
1802:
1792:
1791:
1785:
1784:
1782:
1781:
1768:
1755:
1742:
1729:
1714:
1698:
1696:
1690:
1689:
1684:
1675:
1674:
1671:
1670:
1663:
1661:
1658:
1657:
1654:
1653:
1651:
1650:
1643:
1635:
1633:
1627:
1626:
1624:
1623:
1615:
1608:
1601:
1594:
1591:Schinderhannes
1587:
1580:
1573:
1566:
1559:
1552:
1544:
1538:
1532:
1531:
1529:
1528:
1521:
1513:
1511:
1505:
1504:
1502:
1501:
1494:
1487:
1480:
1472:
1470:
1464:
1463:
1461:
1460:
1453:
1446:
1438:
1430:
1428:
1422:
1421:
1419:
1418:
1411:
1404:
1401:Innovatiocaris
1397:
1389:
1380:
1374:
1369:
1362:
1361:
1359:
1358:
1349:
1343:
1337:
1330:
1327:
1326:
1321:
1319:
1318:
1311:
1304:
1296:
1289:
1288:
1251:(4): 770–790.
1231:
1190:
1135:
1108:(4): 605–621.
1085:
1036:
982:
961:(6): 863–869.
938:
882:
831:
818:(5): 971–990.
794:
744:(March 2014).
727:
696:
657:Innovatiocaris
645:
592:
590:
587:
583:
582:
579:
578:
574:
573:
570:
569:
565:
564:
561:
560:
556:
555:
552:
551:
542:
539:
538:
535:
534:
525:
520:
518:
515:
514:
511:
510:
501:
496:
494:
491:
490:
487:
486:
477:
472:
470:
467:
466:
463:
462:
458:
457:
454:
453:
444:
441:
440:
437:
436:
431:
426:
424:
419:
410:
407:
365:
364:Classification
362:
317:
314:
281:
278:
218:
217:
216:
215:
204:
194:
184:
173:
163:
150:
149:
145:
144:
128:
121:
120:
112:
108:
107:
99:
95:
94:
89:
85:
84:
79:
75:
74:
69:
65:
64:
51:
50:
35:
34:
26:
25:
15:
13:
10:
9:
6:
4:
3:
2:
1817:
1806:
1803:
1801:
1798:
1797:
1795:
1778:
1773:
1769:
1765:
1760:
1756:
1752:
1747:
1743:
1739:
1734:
1730:
1725:
1719:
1715:
1710:
1704:
1700:
1699:
1697:
1695:
1691:
1687:
1682:
1667:
1649:
1648:
1644:
1642:
1641:
1637:
1636:
1634:
1632:
1628:
1621:
1620:
1616:
1614:
1613:
1612:Ursulinacaris
1609:
1607:
1606:
1602:
1600:
1599:
1595:
1593:
1592:
1588:
1586:
1585:
1581:
1579:
1578:
1574:
1572:
1571:
1567:
1565:
1564:
1560:
1558:
1557:
1553:
1551:
1550:
1546:
1545:
1542:
1539:
1537:
1533:
1527:
1526:
1522:
1520:
1519:
1515:
1514:
1512:
1510:
1506:
1500:
1499:
1495:
1493:
1492:
1488:
1486:
1485:
1484:Guanshancaris
1481:
1479:
1478:
1477:Amplectobelua
1474:
1473:
1471:
1469:
1465:
1459:
1458:
1454:
1452:
1451:
1447:
1444:
1443:
1439:
1437:
1436:
1432:
1431:
1429:
1427:
1423:
1417:
1416:
1412:
1410:
1409:
1405:
1403:
1402:
1398:
1395:
1394:
1390:
1387:
1386:
1385:Caryosyntrips
1382:
1381:
1378:
1375:
1372:
1367:
1363:
1357:
1354:
1350:
1348:
1347:Panarthropoda
1344:
1342:
1339:Superphylum:
1338:
1336:
1332:
1331:
1328:
1324:
1317:
1312:
1310:
1305:
1303:
1298:
1297:
1294:
1284:
1280:
1276:
1272:
1267:
1262:
1258:
1254:
1250:
1246:
1242:
1235:
1232:
1227:
1223:
1218:
1213:
1209:
1205:
1201:
1194:
1191:
1186:
1182:
1178:
1174:
1170:
1166:
1162:
1158:
1154:
1150:
1146:
1139:
1136:
1131:
1127:
1123:
1119:
1115:
1111:
1107:
1103:
1096:
1089:
1086:
1081:
1077:
1072:
1067:
1063:
1059:
1055:
1051:
1047:
1040:
1037:
1032:
1028:
1023:
1018:
1013:
1008:
1004:
1000:
996:
989:
987:
983:
978:
974:
969:
964:
960:
956:
952:
945:
943:
939:
934:
930:
926:
922:
918:
914:
910:
906:
902:
898:
891:
889:
887:
883:
878:
874:
870:
866:
862:
858:
854:
850:
846:
842:
835:
832:
826:
821:
817:
813:
812:Palaeontology
809:
807:
798:
795:
790:
786:
782:
778:
774:
770:
766:
762:
758:
754:
747:
743:
736:
734:
732:
728:
723:
719:
715:
711:
707:
700:
697:
692:
688:
684:
680:
676:
672:
668:
664:
660:
658:
649:
646:
640:
635:
631:
627:
622:
617:
613:
609:
605:
597:
594:
588:
581:
580:
572:
571:
563:
562:
554:
553:
550:
549:
548:
541:
540:
537:
536:
533:
532:
531:
524:
523:
517:
516:
513:
512:
509:
508:
507:
500:
499:
493:
492:
489:
488:
485:
484:
483:
476:
475:
469:
468:
465:
464:
456:
455:
452:
451:
450:
443:
442:
439:
438:
435:
430:
429:
423:
422:
418:
416:
408:
406:
404:
400:
396:
393:
392:
391:Amplectobelua
387:
386:
381:
377:
376:
375:Amplectobelua
371:
363:
361:
359:
358:Guanshancaris
355:
351:
350:
345:
344:Amplectobelua
341:
340:
335:
334:
329:
328:
327:Amplectobelua
323:
315:
313:
311:
310:
305:
304:
299:
298:
293:
292:
287:
279:
277:
275:
274:
269:
265:
264:Sirius Passet
261:
260:
259:Guanshancaris
255:
254:
249:
248:
243:
242:
241:Amplectobelua
237:
234:
230:
226:
225:
213:
212:
205:
203:
202:
195:
193:
192:
185:
182:
181:
174:
172:
171:
170:Guanshancaris
164:
162:
161:
160:Amplectobelua
154:
153:
151:
146:
141:
135:
129:
126:
123:
122:
119:
113:
110:
109:
106:
100:
97:
96:
93:
90:
87:
86:
83:
80:
77:
76:
73:
70:
67:
66:
61:
56:
52:
48:
47:
42:
41:
40:Amplectobelua
36:
32:
27:
22:
19:
1693:
1645:
1640:Aegirocassis
1638:
1619:Zhenghecaris
1617:
1610:
1603:
1598:Stanleycaris
1596:
1589:
1582:
1575:
1568:
1563:Cordaticaris
1561:
1556:Cambroraster
1554:
1547:
1525:Tamisiocaris
1523:
1518:Echidnacaris
1516:
1496:
1489:
1482:
1475:
1467:
1455:
1448:
1440:
1435:Anomalocaris
1433:
1413:
1406:
1399:
1393:Cucumericrus
1391:
1383:
1248:
1244:
1234:
1207:
1203:
1193:
1155:(1): 40–48.
1152:
1148:
1138:
1105:
1101:
1088:
1053:
1049:
1039:
1002:
998:
958:
954:
900:
896:
844:
840:
834:
815:
811:
806:Anomalocaris
805:
797:
756:
752:
713:
709:
699:
666:
662:
656:
648:
611:
607:
596:
545:
544:
528:
527:
504:
503:
480:
479:
447:
446:
434:NIGP 154565
433:
415:a posteriori
414:
412:
405:of Vermont.
398:
394:
389:
383:
379:
373:
369:
367:
357:
353:
347:
343:
339:Anomalocaris
337:
331:
325:
319:
307:
301:
295:
289:
285:
283:
271:
257:
251:
245:
239:
223:
222:
221:
209:
199:
189:
178:
168:
158:
139:
133:
124:
105:Dinocaridida
44:
38:
18:
1718:Wikispecies
1605:Titanokorys
1549:Buccaspinea
1498:Ramskoeldia
1450:Lenisicaris
1408:Laminacaris
378:by Vinther
349:Ramskoeldia
316:Description
253:Ramskoeldia
201:Ramskoeldia
43:(left) and
1800:Radiodonta
1794:Categories
1371:Radiodonta
1356:Arthropoda
1353:Stem-group
1323:Radiodonta
1005:(1): 208.
589:References
280:Definition
236:radiodonts
118:Radiodonta
92:Arthropoda
1709:Q17409560
1577:Pahvantia
1536:Hurdiidae
1491:Lyrarapax
1341:Ecdysozoa
1333:Kingdom:
1283:246933116
1275:0022-3360
1226:0567-7920
1185:134157062
1177:0022-3360
1122:2056-2802
977:2095-5138
789:205237459
742:Harper DA
691:252147346
683:0016-7649
630:2375-2548
409:Phylogeny
354:Lyrarapax
333:Lyrarapax
284:In 2014,
268:Greenland
247:Lyrarapax
191:Lyrarapax
78:Kingdom:
72:Eukaryota
46:Lyrarapax
1703:Wikidata
1457:Shucaris
1442:Houcaris
1351:Phylum:
1335:Animalia
1130:90258934
1080:31362637
1031:28854872
925:25043032
869:17780848
781:24670770
639:10796117
397:Vinther
294:but not
273:Houcaris
266:site in
233:Cambrian
211:Shucaris
180:Houcaris
88:Phylum:
82:Animalia
68:Domain:
49:(right)
1764:6164272
1738:8607539
1584:Peytoia
1345:Clade:
1253:Bibcode
1157:Bibcode
1071:6710600
1022:5577670
933:4451239
905:Bibcode
877:1913482
849:Bibcode
841:Science
761:Bibcode
148:Genera
111:Order:
98:Class:
1777:290858
1751:122878
1570:Hurdia
1281:
1273:
1224:
1183:
1175:
1128:
1120:
1078:
1068:
1029:
1019:
975:
931:
923:
897:Nature
875:
867:
787:
779:
753:Nature
689:
681:
636:
628:
322:pincer
142:, 2019
140:et al.
138:Pates
1746:IRMNG
1279:S2CID
1181:S2CID
1126:S2CID
1098:(PDF)
929:S2CID
873:S2CID
785:S2CID
749:(PDF)
687:S2CID
614:(1).
399:et al
395:sensu
380:et al
306:, or
229:clade
227:is a
125:Clade
1733:GBIF
1271:ISSN
1222:ISSN
1173:ISSN
1118:ISSN
1076:PMID
1027:PMID
973:ISSN
921:PMID
865:PMID
777:PMID
679:ISSN
626:ISSN
356:and
346:and
330:and
1261:doi
1212:doi
1165:doi
1110:doi
1066:PMC
1058:doi
1054:286
1017:PMC
1007:doi
963:doi
913:doi
901:513
857:doi
845:264
820:doi
769:doi
757:507
718:doi
671:doi
667:180
634:PMC
616:doi
413:An
360:).
231:of
1796::
1774::
1761::
1748::
1735::
1720::
1705::
1277:.
1269:.
1259:.
1249:96
1247:.
1243:.
1220:.
1210:.
1208:66
1206:.
1202:.
1179:.
1171:.
1163:.
1153:92
1151:.
1147:.
1124:.
1116:.
1104:.
1100:.
1074:.
1064:.
1052:.
1048:.
1025:.
1015:.
1003:17
1001:.
997:.
985:^
971:.
957:.
953:.
941:^
927:.
919:.
911:.
899:.
885:^
871:.
863:.
855:.
843:.
816:56
814:.
810:.
783:.
775:.
767:.
755:.
751:.
730:^
714:97
712:.
708:.
685:.
677:.
669:.
665:.
661:.
632:.
624:.
612:10
610:.
606:.
312:.
300:,
256:,
250:,
244:,
127::
1622:?
1445:?
1396:?
1388:?
1315:e
1308:t
1301:v
1285:.
1263::
1255::
1228:.
1214::
1187:.
1167::
1159::
1132:.
1112::
1106:4
1082:.
1060::
1033:.
1009::
979:.
965::
959:5
935:.
915::
907::
879:.
859::
851::
828:.
822::
791:.
771::
763::
724:.
720::
693:.
673::
655:"
642:.
618::
214:?
207:â€
197:â€
187:â€
183:?
176:â€
166:â€
156:â€
131:â€
115:â€
102:â€
Text is available under the Creative Commons Attribution-ShareAlike License. Additional terms may apply.
