1492:
292:
419:
and
Uzbekistan institutes as well as officials of Tajikistan) has attempted to start a breeding program for the Amu Darya sturgeon, but as of 2021 it has not succeeded (in 2020 Khorezm Mamun Academy in Khiva (Khorezm province branch of Academy of Sciences of Uzbekistan) - the main partner in the program) has temporary stopped shovelnose conservation project because of lack of financial support) although some progress was made (the eggs were incubated in the field in 2012 but the embryos died).
1499:
31:
1505:
92:
50:
311:, reaching up to 2 kg (4.4 lb) in weight and 75 cm (30 in) in total length, excluding tail filament. It has a long, thin tail filament. For example, two individuals that both had a length of about 55 cm (22 in) excluding tail filament were 77 cm (30.5 in) and 85.5 cm (33.5 in) including tail filament. The species occurs in two
337:
May when the water is 14 to 16 °C (57 to 61 °F). During a season, a female of the large morph lays about 3,100–36,500 eggs and a female of the small morph about 1,000–2,000 eggs, but it is possible that they only do this every 4–5-year. The species can reach an age of more than 14 years. Historically, it was known to
831:
Bernard R. Kuhajda (University of
Alabama, USA), Richard L. Mayden (St.Louis University, USA), Vladimir. B. Sal’nikov (National Institute of Deserts, Flora and Fauna, Turkmenistan). Evolutionary and Conservation Studies of Imperiled Sturgeon Species of the Genus Pseudoscaphirhynchus in Central Asia.
418:
the wild populations. In the 1980s there were attempts of doing this with the Amu Darya sturgeon. Although some progress was made, captive breeding ultimately failed because the embryos died. Since 2010, Eurasian
Regional Association of Zoos and Aquariums (EARAZA) together with various Russian
336:
Studies in the 1960s and 1970s showed that Amu Darya sturgeon reach maturity when 5–8 years old (slightly later in females than males), but in the 1990s the youngest mature individuals only were 4 years old, possibly due to environmental changes in their habitat. The breeding season is from March to
815:
V.B. Sal’nikov, N.V. Akimova, G.I. Ruban, R.L. Mayden, B.R. Kuhajda. Reproductive System of Amu-Dar
Shovelnoses - Big Shovelnose Pseudoscaphirhynchus kaufmanni and Little Shovelnose P. hermanni (Acipenseridae). Journal of Ichthyology, Vol. 43, No. 7, 2003, pp. 516–527. Translated from Voprosy
838:
C. B. Dillman, R. M. Wood, B. R. Kuhajda, J. M. Ray, V. B. Salnikov, R. L. Mayden. Molecular systematics of
Scaphirhynchinae: an assessment of North American and Central Asian Freshwater Sturgeon Species. Journal of Applied Ichthyology, 23 (2007), pp. 290–296:
824:
V.B. Sal’nikov, R.L. Mayden, B.R. Kuhajda. Tagging of the Big Amu-Dar
Shovelnose Pseudoscaphirhynchus kaufmanni (Acipenseridae). Journal of Ichthyology, Vol. 44, No. 9, 2004, pp. 778–782. Translated from Voprosy Ikhtiologii, Vol. 44, No. 6, 2004, pp. 832–836:
410:
since 1998. This restricts international trade in the species, but two were legally exported from
Uzbekistan to Russia for zoological purposes in 2002–2012, and in 2013 Uzbekistan set the export quota to twenty individuals.
2057:
369:(modern name is Turkmenabad) in Turkmenistan and also lower in Kyzylrabat area (Uzbekistan), also in more lower Amu Darya River (between Pitnak and Urganch in Khorezm province, Uzbekistan) and in the lower
696:
Salnikov, V.B.; V.J. Birstein; and R.L. Mayden (1996). The contemporary status of the two Amu Darya River shovelnose sturgeons, Pseudoscaphirhynchus kaufmanni and P. hermanni. The
Sturgeon Quarterly 4(3):
1491:
727:
2254:
728:
Review of species selected on the basis of new or increased CITES export quotas in 2013: Pseudoscaphirhynchus kaufmanni from
Uzbekistan & update to the 2013 quota analysis
2135:
315:: a relatively large, light-coloured and late maturing morph, and a small, dark-coloured and early maturing morph. The small morph is roughly of similar size to
2174:
791:
2284:
597:
403:
1540:
645:
Birstein, V.J.; J.R. Waldman; and W.E. Bemis, editors (1997). Sturgeon biodiversity and conservation, pp. 377–378, 381–382. Kluwer
Academic Publishing.
2109:
381:. Despite its threatened status, strong decline and range contraction, it can be locally fairly common and overall it is the least rare species of
2148:
666:
2187:
867:
786:
2279:
796:
840:
833:
2083:
797:
Large Amu Darya Shovelnose Sturgeon (Pseudoscaphirhynchus kaufmanni) in the lower Vakhsh River (an Amu Darya tributary) in Tadjikistan
650:
2022:
291:
2264:
2226:
741:
Kovalev, Konstantin; Balashov, Dmitry; Cherniak, Alexey; Lebedeva, Elena; Vasil’eva, Ekaterina; Vasil’ev, Victor (30 June 2014).
398:
In general the status of the Amu Darya sturgeon is poorly known and further surveys are necessary. A part of its range is in the
91:
2289:
946:
711:
Mikodina, E.; and A. Novosadova (2015). Rare Acipenserids in Russian Aquaculture. ВОПРОСЫ РЫБНОГО ХОЗЯЙСТВА БЕЛАРУСИ 31: 38–51.
1533:
2153:
808:
2269:
2259:
399:
54:
373:(an Amu Darya tributary) in Tajikistan. It has declined because of habitat loss, pollution and poaching, and is rated as
2274:
1504:
1498:
803:
2031:
817:
2179:
1526:
860:
809:
Large Amu Darya Shovelnose Sturgeon (Pseudoscaphirhynchus kaufmanni) in the middle Amu Darya River (Turkmenistan)
608:
804:
Large Amu Darya Shovelnose Sturgeon (Pseudoscaphirhynchus kaufmanni) in the middle Amu Darya River (Uzbekistan)
792:
Large Amu Darya Shovelnose Sturgeon (Pseudoscaphirhynchus kaufmanni) in the lower Amu Darya River (Uzbekistan)
203:
1984:
826:
329:
The Amu Darya sturgeon feeds on small fish and aquatic insect larvae, with large individuals being mostly
312:
2200:
2036:
1769:
374:
257:
186:
30:
2231:
2070:
1826:
1460:
853:
781:
307:
163:
1426:
1416:
1386:
1165:
841:
http://life-on-earth.ru/images/Lopatonosy-Turkmenistan/Molecular_systematics_of_Scaphirhynchinae.pdf
834:
http://life-on-earth.ru/images/Lopatonosy-Turkmenistan/Salnikov-Amudarya_Shovelnose_Sturgeon_Eng.pdf
1934:
1888:
1634:
1453:
1356:
440:
39:
1038:
521:
406:
is widespread and fisheries management in the region is very poor. The species has been listed on
2294:
1958:
1837:
1625:
1376:
1283:
1130:
1070:
1060:
387:
212:
86:
2062:
1116:
1049:
2205:
2140:
1297:
1269:
1158:
2044:
1811:
1715:
1688:
1679:
1670:
1616:
1137:
1092:
1081:
646:
407:
1897:
1751:
1724:
1697:
1652:
1550:
1181:
1151:
758:
516:
448:
338:
261:
143:
2166:
1879:
1760:
1198:
1518:
569:
1870:
1846:
1778:
1661:
1467:
1366:
1339:
1308:
1260:
1246:
1223:
1209:
1107:
931:
906:
782:
Large Amu Darya Shovelnose Sturgeon (Pseudoscaphirhynchus kaufmanni) digital morphology
317:
133:
2248:
2161:
1915:
1802:
1742:
1706:
1643:
1443:
1349:
662:
507:
415:
414:
Many sturgeon species have been bred in captivity and this often forms the basis for
402:. Although the Amu Darya sturgeon is a fully protected species throughout its range,
264:
59:
498:
345:, with the offspring having features that are intermediate between the two species.
2049:
1581:
1329:
1290:
1276:
1232:
818:
http://life-on-earth.ru/images/Salnikov-Amudarya_Shovelnose_Reproductive_System.pdf
452:
370:
268:
455:. Further karyological research on the Amu Darya sturgeon may help in determining
2218:
2122:
2016:
1587:
1006:
984:
958:
915:
876:
787:
Large Amu Darya Shovelnose Sturgeon (Pseudoscaphirhynchus kaufmanni) at FishBase
330:
280:
2007:
1575:
1317:
1144:
1123:
435:; it is classified as a low-chromosome group acipenserid fish. Studies of its
432:
299:
276:
272:
2088:
1949:
1607:
1406:
1239:
763:
742:
456:
428:
366:
354:
103:
543:
2096:
2001:
1925:
1597:
1569:
1397:
1027:
900:
675:
574:
436:
358:
153:
123:
2114:
2101:
1733:
827:
http://life-on-earth.ru/images/Salnikov-Amudarya_Shovelnose_Tagging.PDF
305:
The Amu Darya sturgeon is a small sturgeon, but the largest species of
2127:
283:. It inhabits quite shallow flowing waters that are turbid and muddy.
1563:
894:
113:
1978:
2213:
362:
290:
76:
70:
2075:
2192:
1793:
1433:
378:
1982:
1522:
887:
849:
492:
845:
490:
488:
486:
484:
482:
480:
478:
476:
474:
472:
361:, today it is found in the middle Amu Darya River (between
431:
of the Amu Darya sturgeon consists of roughly 118–120
439:
and karyotype show clear differences compared to the
1449:
1439:
1422:
1412:
1382:
1372:
1362:
1345:
1335:
1325:
1257:
1220:
1205:
1194:
1104:
1088:
1077:
1066:
1056:
1045:
1034:
1023:
447:), indicating that the usually recognized subfamily
353:
The Amu Darya sturgeon once occurred throughout the
1991:
1947:
1923:
1914:
1868:
1824:
1791:
1605:
1596:
1396:
1316:
1307:
1256:
1219:
1179:
1103:
1004:
957:
930:
832:Report, 2007. Copyright 2016, All Rights Reserved:
747:(Actinopterygii: Acipenseriformes: Acipenseridae)"
816:Ikhtiologii, Vol. 43, No. 4, 2003, pp. 499–510:
707:
705:
703:
692:
690:
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686:
641:
639:
637:
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633:
631:
629:
1534:
861:
522:10.2305/IUCN.UK.2010-1.RLTS.T18601A8498207.en
8:
2255:IUCN Red List critically endangered species
1979:
1920:
1602:
1541:
1527:
1519:
1313:
1187:
1016:
1012:
1001:
936:
927:
884:
868:
854:
846:
743:"The karyotype of the Amu Darya sturgeon,
721:
719:
717:
48:
29:
20:
762:
520:
357:river basin, including its delta in the
321:, a species that lacks a tail filament.
468:
395:only survives in very small numbers).
219:
7:
2285:Critically endangered fauna of Asia
508:IUCN Red List of Threatened Species
14:
1503:
1497:
1490:
751:Acta Ichthyologica et Piscatoria
605:Deeplyfish – fishes of the world
90:
459:evolution in acipenserid fish.
2063:pseudoscaphirhynchus-kaufmanni
2023:Pseudoscaphirhynchus kaufmanni
1993:Pseudoscaphirhynchus kaufmanni
745:Pseudoscaphirhynchus kaufmanni
669:Pseudoscaphirhynchus kaufmanni
568:Froese, R.; Pauly, D. (2017).
501:Pseudoscaphirhynchus kaufmanni
253:Pseudoscaphirhynchus kaufmanni
230:Pseudoscaphirhynchus rossikowi
196:Pseudoscaphirhynchus kaufmanni
1:
400:Amudarya State Nature Reserve
445:Scaphirhynchus platorhynchus
2311:
2280:Critically endangered fish
1558:
1487:
1190:
1019:
1015:
1000:
981:
939:
926:
883:
248:false shovelnose sturgeon
218:
211:
192:
185:
87:Scientific classification
85:
68:
46:
37:
28:
23:
1718:A. oxyrinchus oxyrinchus
665:; Pauly, Daniel (eds.).
391:is possibly extinct and
2265:Freshwater fish of Asia
1770:European sea sturgeon (
1551:sturgeon and paddlefish
764:10.3750/AIP2014.44.2.04
260:species of fish in the
222:Scaphirhynchus hermanni
2290:Fish described in 1877
302:
2201:Paleobiology Database
1935:American paddlefish (
1889:Shovelnose sturgeon (
1709:A. oxyrinchus desotoi
731:UNEP-WCMC, Cambridge.
375:critically endangered
294:
258:critically endangered
55:Critically Endangered
2270:Fish of Central Asia
2260:Pseudoscaphirhynchus
1959:Chinese paddlefish (
1856:Amu Darya sturgeon (
1838:Syr Darya sturgeon (
1626:Shortnose sturgeon (
1461:Pseudoscaphirhynchus
614:on 18 September 2017
544:"Appendices | CITES"
515:: e.T18601A8498207.
383:Pseudoscaphirhynchus
308:Pseudoscaphirhynchus
165:Pseudoscaphirhynchus
2275:Fauna of Tajikistan
1716:Atlantic sturgeon (
1689:Adriatic sturgeon (
1680:Japanese sturgeon (
1671:Sakhalin sturgeon (
1617:Siberian sturgeon (
1454:Protoscaphirhynchus
725:UNEP-WCMC. (2013).
679:. May 2019 version.
441:shovelnose sturgeon
349:Conservation status
298:on a 1996 stamp of
40:Conservation status
24:Amu Darya sturgeon
1898:Alabama sturgeon (
1752:Chinese sturgeon (
1725:Persian sturgeon (
1698:Bastard sturgeon (
1655:A. gueldenstaedtii
1653:Russian sturgeon (
1635:Dabry's sturgeon (
1071:Bobasatraniiformes
497:Mugue, N. (2010).
303:
244:Amu Darya sturgeon
202:Bogdanov 1874 ex.
2242:
2241:
1985:Taxon identifiers
1976:
1975:
1972:
1971:
1910:
1909:
1880:Pallid sturgeon (
1761:Starry sturgeon (
1516:
1515:
1512:
1511:
1485:
1484:
1481:
1480:
1477:
1476:
1175:
1174:
1093:Saurichthyiformes
1082:Guildayichthyidae
996:
995:
992:
991:
977:
976:
577:version (02/2017)
408:CITES Appendix II
267:. It is found in
240:
239:
234:
226:
178:P. kaufmanni
80:
63:
2302:
2235:
2234:
2222:
2221:
2209:
2208:
2196:
2195:
2183:
2182:
2170:
2169:
2157:
2156:
2144:
2143:
2131:
2130:
2118:
2117:
2105:
2104:
2092:
2091:
2079:
2078:
2066:
2065:
2053:
2052:
2040:
2039:
2027:
2026:
2025:
2012:
2011:
2010:
1980:
1921:
1847:Dwarf sturgeon (
1781:A. transmontanus
1779:White sturgeon (
1682:A. multiscutatus
1662:Green sturgeon (
1603:
1543:
1536:
1529:
1520:
1507:
1501:
1495:
1494:
1451:
1441:
1424:
1414:
1384:
1374:
1364:
1347:
1337:
1327:
1314:
1259:
1222:
1207:
1196:
1188:
1182:Acipenseriformes
1152:Plesiococcolepis
1106:
1090:
1079:
1068:
1058:
1047:
1036:
1025:
1017:
1013:
1002:
937:
928:
921:
920:
885:
870:
863:
856:
847:
769:
768:
766:
738:
732:
730:
723:
712:
709:
698:
694:
681:
680:
659:
653:
643:
624:
623:
621:
619:
613:
607:. Archived from
602:
594:
588:
587:
585:
583:
565:
559:
558:
556:
554:
540:
534:
533:
531:
529:
524:
494:
449:Scaphirhynchinae
233:Nikolskii 1900
232:
224:
198:
144:Acipenseriformes
95:
94:
74:
57:
52:
51:
33:
21:
2310:
2309:
2305:
2304:
2303:
2301:
2300:
2299:
2245:
2244:
2243:
2238:
2230:
2225:
2217:
2212:
2204:
2199:
2191:
2186:
2178:
2173:
2165:
2160:
2152:
2147:
2139:
2134:
2126:
2121:
2113:
2108:
2100:
2095:
2087:
2082:
2074:
2069:
2061:
2056:
2048:
2043:
2035:
2030:
2021:
2020:
2015:
2006:
2005:
2000:
1987:
1977:
1968:
1943:
1906:
1891:S. platorynchus
1864:
1840:P. fedtschenkoi
1828:
1820:
1787:
1743:Amur sturgeon (
1707:Gulf sturgeon (
1644:Lake sturgeon (
1628:A. brevirostrum
1592:
1554:
1547:
1517:
1508:
1489:
1473:
1392:
1303:
1252:
1215:
1199:Cylindracanthus
1184:
1171:
1099:
1009:
988:
973:
953:
922:
890:
889:
879:
874:
778:
773:
772:
740:
739:
735:
726:
724:
715:
710:
701:
695:
684:
661:
660:
656:
644:
627:
617:
615:
611:
600:
598:"Acipenseridae"
596:
595:
591:
581:
579:
570:"Acipenseridae"
567:
566:
562:
552:
550:
542:
541:
537:
527:
525:
496:
495:
470:
465:
425:
404:illegal fishing
388:P. fedtschenkoi
351:
327:
289:
207:
200:
194:
181:
89:
81:
64:
53:
49:
42:
17:
16:Species of fish
12:
11:
5:
2308:
2306:
2298:
2297:
2292:
2287:
2282:
2277:
2272:
2267:
2262:
2257:
2247:
2246:
2240:
2239:
2237:
2236:
2223:
2210:
2197:
2184:
2171:
2158:
2145:
2132:
2119:
2106:
2093:
2080:
2067:
2054:
2041:
2028:
2013:
1997:
1995:
1989:
1988:
1983:
1974:
1973:
1970:
1969:
1967:
1966:
1955:
1953:
1945:
1944:
1942:
1941:
1931:
1929:
1918:
1912:
1911:
1908:
1907:
1905:
1904:
1895:
1886:
1876:
1874:
1871:Scaphirhynchus
1866:
1865:
1863:
1862:
1853:
1844:
1834:
1832:
1829:scaphirhynchus
1822:
1821:
1819:
1818:
1809:
1799:
1797:
1789:
1788:
1786:
1785:
1776:
1767:
1758:
1749:
1740:
1731:
1722:
1713:
1704:
1700:A. nudiventris
1695:
1686:
1677:
1668:
1664:A. medirostris
1659:
1650:
1641:
1632:
1623:
1613:
1611:
1600:
1594:
1593:
1591:
1590:
1584:
1578:
1572:
1566:
1559:
1556:
1555:
1548:
1546:
1545:
1538:
1531:
1523:
1514:
1513:
1510:
1509:
1488:
1486:
1483:
1482:
1479:
1478:
1475:
1474:
1472:
1471:
1468:Scaphirhynchus
1464:
1457:
1447:
1437:
1430:
1427:Engdahlichthys
1420:
1417:Anchiacipenser
1410:
1402:
1400:
1394:
1393:
1391:
1390:
1387:Pugiopsephurus
1380:
1370:
1367:Protopsephurus
1360:
1353:
1343:
1340:Paleopsephurus
1333:
1322:
1320:
1311:
1309:Acipenseroidei
1305:
1304:
1302:
1301:
1294:
1287:
1280:
1273:
1265:
1263:
1261:Peipiaosteidae
1254:
1253:
1251:
1250:
1247:Strongylosteus
1243:
1236:
1228:
1226:
1224:Chondrosteidae
1217:
1216:
1214:
1213:
1210:Eochondrosteus
1203:
1191:
1185:
1180:
1177:
1176:
1173:
1172:
1170:
1169:
1166:Toarcocephalus
1162:
1155:
1148:
1141:
1134:
1127:
1120:
1112:
1110:
1108:Coccolepididae
1101:
1100:
1098:
1097:
1086:
1075:
1064:
1054:
1043:
1032:
1020:
1010:
1005:
998:
997:
994:
993:
990:
989:
982:
979:
978:
975:
974:
972:
971:
970:
969:
963:
961:
955:
954:
952:
951:
950:
949:
947:Actinopterygii
940:
934:
932:Actinopterygii
924:
923:
919:
918:
909:
907:Actinopterygii
903:
897:
888:
881:
880:
875:
873:
872:
865:
858:
850:
844:
843:
836:
829:
821:
820:
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811:
806:
800:
799:
794:
789:
784:
777:
776:External links
774:
771:
770:
757:(2): 111–116.
733:
713:
699:
682:
663:Froese, Rainer
654:
625:
589:
560:
535:
467:
466:
464:
461:
424:
421:
350:
347:
326:
323:
288:
285:
238:
237:
236:
235:
227:
225:Kessler 1877
216:
215:
209:
208:
201:
190:
189:
183:
182:
175:
173:
169:
168:
161:
157:
156:
151:
147:
146:
141:
137:
136:
134:Actinopterygii
131:
127:
126:
121:
117:
116:
111:
107:
106:
101:
97:
96:
83:
82:
69:
66:
65:
47:
44:
43:
38:
35:
34:
26:
25:
15:
13:
10:
9:
6:
4:
3:
2:
2307:
2296:
2293:
2291:
2288:
2286:
2283:
2281:
2278:
2276:
2273:
2271:
2268:
2266:
2263:
2261:
2258:
2256:
2253:
2252:
2250:
2233:
2228:
2224:
2220:
2215:
2211:
2207:
2202:
2198:
2194:
2189:
2185:
2181:
2176:
2172:
2168:
2163:
2159:
2155:
2150:
2146:
2142:
2137:
2133:
2129:
2124:
2120:
2116:
2111:
2107:
2103:
2098:
2094:
2090:
2085:
2081:
2077:
2072:
2068:
2064:
2059:
2055:
2051:
2046:
2042:
2038:
2033:
2029:
2024:
2018:
2014:
2009:
2003:
1999:
1998:
1996:
1994:
1990:
1986:
1981:
1964:
1962:
1957:
1956:
1954:
1952:
1951:
1946:
1940:
1938:
1933:
1932:
1930:
1928:
1927:
1922:
1919:
1917:
1916:Polyodontidae
1913:
1903:
1901:
1896:
1894:
1892:
1887:
1885:
1883:
1878:
1877:
1875:
1873:
1872:
1867:
1861:
1859:
1854:
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1845:
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1841:
1836:
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1833:
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1823:
1817:
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1806:
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1790:
1784:
1782:
1777:
1775:
1773:
1768:
1766:
1764:
1759:
1757:
1755:
1750:
1748:
1746:
1745:A. schrenckii
1741:
1739:
1737:
1732:
1730:
1728:
1723:
1721:
1719:
1714:
1712:
1710:
1705:
1703:
1701:
1696:
1694:
1692:
1687:
1685:
1683:
1678:
1676:
1674:
1669:
1667:
1665:
1660:
1658:
1656:
1651:
1649:
1647:
1646:A. fulvescens
1642:
1640:
1638:
1633:
1631:
1629:
1624:
1622:
1620:
1615:
1614:
1612:
1610:
1609:
1604:
1601:
1599:
1598:Acipenseridae
1595:
1589:
1585:
1583:
1579:
1577:
1573:
1571:
1567:
1565:
1561:
1560:
1557:
1552:
1544:
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1537:
1532:
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1525:
1524:
1521:
1506:
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1500:
1493:
1470:
1469:
1465:
1463:
1462:
1458:
1456:
1455:
1448:
1446:
1445:
1444:Priscosturion
1438:
1436:
1435:
1431:
1429:
1428:
1421:
1419:
1418:
1411:
1409:
1408:
1404:
1403:
1401:
1399:
1398:Acipenseridae
1395:
1389:
1388:
1381:
1379:
1378:
1371:
1369:
1368:
1361:
1359:
1358:
1354:
1352:
1351:
1350:Parapsephurus
1344:
1342:
1341:
1334:
1332:
1331:
1324:
1323:
1321:
1319:
1318:Polyodontidae
1315:
1312:
1310:
1306:
1300:
1299:
1295:
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1292:
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1285:
1281:
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1237:
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1234:
1230:
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1227:
1225:
1218:
1212:
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1200:
1193:
1192:
1189:
1186:
1183:
1178:
1168:
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1163:
1161:
1160:
1156:
1154:
1153:
1149:
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1142:
1140:
1139:
1135:
1133:
1132:
1128:
1126:
1125:
1121:
1119:
1118:
1114:
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1111:
1109:
1102:
1095:
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1087:
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1076:
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1072:
1065:
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1055:
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1018:
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966:
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917:
913:
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886:
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871:
866:
864:
859:
857:
852:
851:
848:
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837:
835:
830:
828:
823:
822:
819:
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813:
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807:
805:
802:
801:
798:
795:
793:
790:
788:
785:
783:
780:
779:
775:
765:
760:
756:
752:
748:
746:
737:
734:
729:
722:
720:
718:
714:
708:
706:
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700:
693:
691:
689:
687:
683:
678:
677:
672:
670:
664:
658:
655:
652:
651:0-792-34517-7
648:
642:
640:
638:
636:
634:
632:
630:
626:
610:
606:
599:
593:
590:
578:
576:
571:
564:
561:
549:
545:
539:
536:
523:
518:
514:
510:
509:
504:
502:
493:
491:
489:
487:
485:
483:
481:
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477:
475:
473:
469:
462:
460:
458:
454:
450:
446:
442:
438:
434:
430:
422:
420:
417:
412:
409:
405:
401:
396:
394:
390:
389:
384:
380:
376:
372:
368:
364:
360:
356:
348:
346:
344:
340:
334:
332:
324:
322:
320:
319:
314:
310:
309:
301:
297:
293:
286:
284:
282:
278:
274:
270:
266:
265:Acipenseridae
263:
259:
255:
254:
249:
245:
231:
228:
223:
220:
217:
214:
210:
205:
199:
197:
191:
188:
187:Binomial name
184:
180:
179:
174:
171:
170:
167:
166:
162:
159:
158:
155:
154:Acipenseridae
152:
149:
148:
145:
142:
139:
138:
135:
132:
129:
128:
125:
122:
119:
118:
115:
112:
109:
108:
105:
102:
99:
98:
93:
88:
84:
78:
72:
67:
61:
56:
45:
41:
36:
32:
27:
22:
19:
1992:
1960:
1948:
1936:
1924:
1899:
1890:
1881:
1869:
1858:P. kaufmanni
1857:
1855:
1848:
1839:
1825:
1813:
1804:
1792:
1780:
1771:
1763:A. stellatus
1762:
1753:
1744:
1735:
1726:
1717:
1708:
1699:
1690:
1681:
1672:
1663:
1654:
1645:
1637:A. dabryanus
1636:
1627:
1618:
1606:
1582:Osteichthyes
1496:
1466:
1459:
1452:
1442:
1432:
1425:
1415:
1405:
1385:
1375:
1365:
1355:
1348:
1338:
1330:Crossopholis
1328:
1296:
1291:Stichopterus
1289:
1282:
1277:Peipiaosteus
1275:
1268:
1245:
1238:
1233:Chondrosteus
1231:
1208:
1197:
1164:
1157:
1150:
1143:
1136:
1129:
1122:
1115:
1091:
1080:
1069:
1059:
1048:
1039:Errolichthys
1037:
1026:
983:
911:
754:
750:
744:
736:
674:
668:
657:
616:. Retrieved
609:the original
604:
592:
580:. Retrieved
573:
563:
551:. Retrieved
547:
538:
526:. Retrieved
512:
506:
500:
453:polyphyletic
444:
426:
413:
397:
392:
386:
382:
371:Vakhsh River
352:
342:
335:
328:
316:
306:
304:
296:P. kaufmanni
295:
279:and perhaps
269:Turkmenistan
252:
251:
247:
243:
241:
229:
221:
195:
193:
177:
176:
164:
18:
2123:iNaturalist
2017:Wikispecies
1937:P. spathula
1900:S. suttkusi
1849:P. hermanni
1805:H. dauricus
1754:A. sinensis
1736:A. ruthenus
1727:A. persicus
1691:A. naccarii
1588:Chondrostei
1580:Superclass
1284:Spherosteus
1131:Condorlepis
1061:Neochallaia
1007:Chondrostei
985:Chondrostei
959:Chondrostei
916:Actinopteri
877:Chondrostei
528:17 November
433:chromosomes
393:P. hermanni
343:P. hermanni
331:piscivorous
318:P. hermanni
281:Afghanistan
73:Appendix II
2249:Categories
1961:P. gladius
1673:A. mikadoi
1576:Vertebrata
1574:Subphylum
1145:Morrolepis
1124:Coccolepis
1117:Barbalepis
1050:Gualolepis
968:see below↓
553:14 January
463:References
416:restocking
300:Kyrgyzstan
287:Appearance
277:Tajikistan
273:Uzbekistan
2295:Amu Darya
1950:Psephurus
1772:A. sturio
1734:Sterlet (
1619:A. baerii
1608:Acipenser
1586:Subclass
1407:Acipenser
1377:Psephurus
1298:Yanosteus
1270:Liaosteus
1240:Gyrosteus
1159:Sunolepis
893:Kingdom:
548:cites.org
457:polyploid
429:karyotype
367:Chardzhou
355:Amu Darya
339:hybridize
172:Species:
110:Kingdom:
104:Eukaryota
2214:Species+
2141:10154851
2097:FishBase
2076:46561193
2008:Q1507886
2002:Wikidata
1926:Polyodon
1882:S. albus
1827:Pseudo-
1812:Beluga (
1803:Kaluga (
1570:Chordata
1564:Animalia
1562:Kingdom
1357:Polyodon
1138:Iyalepis
1028:Birgeria
901:Chordata
899:Phylum:
895:Animalia
676:FishBase
575:FishBase
437:genetics
423:Taxonomy
359:Aral Sea
325:Behavior
213:Synonyms
150:Family:
124:Chordata
120:Phylum:
114:Animalia
100:Domain:
60:IUCN 3.1
2115:2402094
1814:H. huso
1568:Phylum
1553:species
1549:Recent
905:Class:
377:by the
256:) is a
204:Kessler
160:Genus:
140:Order:
130:Class:
75: (
58: (
2232:282474
2193:282474
2154:573258
2128:111095
697:10–14.
649:
618:18 May
582:18 May
313:morphs
262:family
2227:WoRMS
2206:94899
2180:55290
2167:18601
2136:IRMNG
2089:17197
2084:EUNIS
2050:6WH8D
2037:50214
912:Clade
612:(PDF)
601:(PDF)
363:Kerki
341:with
77:CITES
71:CITES
2219:4111
2188:OBIS
2175:NCBI
2162:IUCN
2149:ITIS
2110:GBIF
2102:4634
2032:BOLD
1794:Huso
1434:Huso
945:see
647:ISBN
620:2017
584:2017
555:2022
530:2021
513:2010
427:The
379:IUCN
365:and
242:The
206:1877
2071:EoL
2058:CMS
2045:CoL
759:doi
517:doi
451:is
246:or
2251::
2229::
2216::
2203::
2190::
2177::
2164::
2151::
2138::
2125::
2112::
2099::
2086::
2073::
2060::
2047::
2034::
2019::
2004::
914::
755:44
753:.
749:.
716:^
702:^
685:^
673:.
628:^
603:.
572:.
546:.
511:.
505:.
471:^
333:.
275:,
271:,
1965:†
1963:)
1939:)
1902:)
1893:)
1884:)
1860:)
1851:)
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1816:)
1807:)
1783:)
1774:)
1765:)
1756:)
1747:)
1738:)
1729:)
1720:)
1711:)
1702:)
1693:)
1684:)
1675:)
1666:)
1657:)
1648:)
1639:)
1630:)
1621:)
1542:e
1535:t
1528:v
1450:†
1440:†
1423:†
1413:†
1383:†
1373:†
1363:†
1346:†
1336:†
1326:†
1258:†
1221:†
1206:†
1202:?
1195:†
1105:†
1096:?
1089:†
1085:?
1078:†
1074:?
1067:†
1057:†
1053:?
1046:†
1042:?
1035:†
1031:?
1024:†
869:e
862:t
855:v
767:.
761::
671:"
667:"
622:.
586:.
557:.
532:.
519::
503:"
499:"
443:(
385:(
250:(
79:)
62:)
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