105:
35:
127:
1478:
1400:
1452:
1413:
1514:
1206:
1614:
56:
1220:
1322:
78:
1327:
1439:
1563:
1457:
1465:
1609:
1296:
49:
43:
1221:"Molecular Systematics of Boraginaceae Tribe Boragineae Based on ITS1 and trnL Sequences, with Special Reference to
1568:
1527:
1180:
1483:
126:
60:
881:
104:
1103:
1532:
1387:
642:
806:
1374:
1239:
1078:
921:
984:
775:
1392:
1379:
994:
816:
765:
555:
1061:
785:
714:
704:
673:
586:
1004:
974:
735:
652:
500:
The first two form one subclade, the other two each a separate subclade. The subclade of the subgenus
1493:
1335:
941:
911:
901:
891:
871:
796:
755:
745:
724:
683:
565:
1418:
1173:
861:
827:
469:
254:
113:
20:
387:
are sapphire blue and retain their colour a long time. The plants show numerous flowers with five
1576:
1519:
1139:
358:
266:
121:
1501:
1309:
1202:
434:
1604:
1506:
931:
662:
575:
346:
373:
164:
834:– true alkanet, bugloss, common bugloss, corn bugloss, field bugloss (type species)
376:
with the general characteristics of the borage family. They are commonly herbaceous. The
1366:
693:
596:
369:
151:
399:, simple or branched, or are clustered at the end. The flowers are much frequented by
1598:
1277:
1091:
450:
396:
237:
220:
1581:
1121:
429:
365:
350:
246:
210:
1288:
1314:
1301:
1470:
1426:
1271:
464:
411:
200:
1555:
357:
found growing in Europe, North Africa, South Africa and
Western Asia. They are
446:
1361:
1348:
1545:
1262:
395:
forming a narrow tube facing upwards. The flowers grow in several axillary
538:
needs a narrower concept. This way taxonomy and phylogeny with respect to
1540:
1256:
190:
177:
1405:
442:
423:
354:
1431:
1095:
384:
1233:
1353:
460:
438:
392:
388:
337:
138:
1444:
377:
1237:
1340:
400:
28:
441:
colouring matter. It is insoluble in water, but soluble in
16:
534:
clearly need new independent entities, while the subgenus
700:(status vulnerable to endangered in France and Italy)
519:, which diverges markedly in its floral morphology.
1246:
380:are simple or undulate, covered with stiff hairs.
1108:(Degen & Dörfl.) Bigazzi, Hilger & Selvi
8:
1234:
103:
92:
19:For former class of Royal Navy ships, see
79:Learn how and when to remove this message
582:– small bugloss, bugloss, annual bugloss
42:This article includes a list of general
1163:
459:species are used as food plants by the
1169:
1167:
1045:(Heldr. ex Guşul.) Selvi & Bigazzi
603:– Italian bugloss, large blue alkanet
287:Gray (1821 publ. 1822), nom. superfl.
7:
1569:94f08e0e-a776-47c6-a457-6994a9ce0e5f
1494:9580be08-3ecc-4a82-9fec-9855680a6a3a
1201:. London: Timber Press. p. 46.
1192:
1190:
515:s.l., then it includes the subgenus
279:Bigazzi, E.Nardi & Selvi (1997)
669:– Cape bugloss, Cape forget-me-not
391:, united at their bases, and five
48:it lacks sufficient corresponding
14:
1197:Grey-Wilson, Christopher (2009).
542:can become completely analogous.
635:(Boiss. & Hohen.) D.F.Chamb.
125:
33:
383:The small radially symmetrical
1533:urn:lsid:ipni.org:names:4451-1
437:(or alkanet-red), a red-brown
410:is commonly used in trough or
1:
1199:The Rock Garden Plant Primer
845:(Griseb.) Selvi & Bigazz
295:Bubani (1897), nom. superfl.
1615:Taxa named by Carl Linnaeus
811:Angius, Pontec. & Selvi
729:Selvi, Bigazzi & Bacch.
311:Fabr. (1759), nom. superfl.
1631:
1181:Plants of the World Online
1135:) – evergreen alkanet
1083:(All.) Vural & Kit Tan
481:There are four subgenera:
18:
1184:. Retrieved 23 July 2024.
1126:(L.) Tausch ex L.H.Bailey
1122:Pentaglottis sempervirens
550:35 species are accepted.
508:) is largely unresolved.
272:
265:
252:
245:
122:Scientific classification
120:
111:
102:
95:
1033:Braun-Blanq. & Maire
882:Anchusa pseudoochroleuca
353:). It includes about 35
1104:Gastrocotyle macedonica
1099:) – Barrelier's bugloss
63:more precise citations.
1225:s.l.; Hilger et al.".
1079:Cynoglottis barrelieri
643:Anchusa × baumgartenii
361:in the United States.
345:belongs to the borage
807:Anchusa montelinasana
760:Rech.f. & Esfand.
1229:. 2004; 94: 201-212.
1130:Anchusa sempervirens
1072:Formerly placed here
979:Boiss. & Spruner
922:Anchusa samothracica
421:(just like those of
1610:Boraginaceae genera
1115:Degen & Dörfler
985:Anchusa × thirkeana
926:Bigazzi & Selvi
850:Anchusa officinalis
838:Anchusa officinalis
828:Anchusa officinalis
790:Selvi & Bigazzi
780:Roem. & Schult.
776:Anchusa leptophylla
623:(Guşul.) D.F.Chamb.
470:Coleophora pennella
255:Anchusa officinalis
114:Anchusa officinalis
1140:Phyllocara aucheri
1112:Anchusa macedonica
1087:Anchusa barrelieri
995:Anchusa tiberiadis
817:Anchusa ochroleuca
766:Anchusa konyaensis
556:Anchusa aegyptiaca
467:species including
1592:
1591:
1502:Open Tree of Life
1240:Taxon identifiers
1208:978-0-88192-928-7
1062:Anchusa variegata
786:Anchusa leucantha
715:Anchusa dinsmorei
705:Anchusa × digenea
674:Anchusa cespitosa
587:Anchusa atlantica
334:
333:
241:
89:
88:
81:
1622:
1585:
1584:
1572:
1571:
1559:
1558:
1549:
1548:
1536:
1535:
1523:
1522:
1510:
1509:
1497:
1496:
1487:
1486:
1474:
1473:
1471:NHMSYS0000455906
1461:
1460:
1448:
1447:
1435:
1434:
1422:
1421:
1409:
1408:
1396:
1395:
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1343:
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1305:
1304:
1292:
1291:
1282:
1281:
1280:
1267:
1266:
1265:
1235:
1227:Annals of Botany
1213:
1212:
1194:
1185:
1171:
1152:
1145:
1134:
1127:
1116:
1109:
1098:
1084:
1067:
1050:Anchusa undulata
1046:
1038:Anchusa undulata
1034:
1026:Anchusa undulata
1022:
1014:Anchusa undulata
1010:
1005:Anchusa undulata
1000:
990:
980:
975:Anchusa thessala
959:
947:
937:
936:Banks & Sol.
932:Anchusa strigosa
927:
917:
907:
897:
887:
877:
867:
846:
833:
823:– yellow alkanet
822:
812:
802:
791:
781:
771:
761:
751:
741:
736:Anchusa gmelinii
730:
720:
710:
699:
689:
679:
668:
663:Anchusa capensis
658:
653:Anchusa calcarea
648:
636:
624:
602:
592:
581:
576:Anchusa arvensis
571:
561:
506:Anchusa capensis
374:perennial plants
364:They consist of
328:
320:
312:
304:
296:
288:
280:
236:
130:
129:
107:
93:
84:
77:
73:
70:
64:
59:this article by
50:inline citations
37:
36:
29:
1630:
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1621:
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1619:
1595:
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1593:
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1580:
1575:
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1562:
1554:
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1544:
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1531:
1526:
1518:
1513:
1505:
1500:
1492:
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1313:
1308:
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1285:
1276:
1275:
1270:
1261:
1260:
1255:
1242:
1217:
1216:
1209:
1196:
1195:
1188:
1172:
1165:
1160:
1150:
1148:Anchusa aucheri
1143:
1132:
1125:
1114:
1107:
1089:
1082:
1074:
1065:
1044:
1032:
1021:(Boiss.) Valdés
1020:
1008:
998:
988:
978:
963:Anchusa stylosa
957:
951:Anchusa stylosa
945:
942:Anchusa stylosa
935:
925:
915:
912:Anchusa riparia
905:
902:Anchusa pusilla
895:
892:Anchusa puechii
885:
875:
872:Anchusa procera
865:
844:
831:
820:
810:
801:Lam. ex Spreng.
800:
797:Anchusa milleri
789:
779:
769:
759:
756:Anchusa iranica
749:
746:Anchusa hybrida
739:
728:
725:Anchusa formosa
718:
708:
697:
687:
684:Anchusa cretica
677:
666:
656:
646:
634:
622:
600:
590:
579:
569:
566:Anchusa affinis
559:
548:
511:If we consider
479:
370:biennial plants
326:
318:
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261:
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235:
124:
85:
74:
68:
65:
55:Please help to
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27:
17:
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11:
5:
1628:
1626:
1618:
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1589:
1587:
1586:
1582:wfo-4000001905
1573:
1560:
1550:
1537:
1524:
1511:
1498:
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1475:
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1449:
1436:
1423:
1410:
1397:
1384:
1371:
1358:
1345:
1332:
1319:
1306:
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1283:
1268:
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1243:
1238:
1232:
1231:
1215:
1214:
1207:
1186:
1162:
1161:
1159:
1156:
1155:
1154:
1144:(A.DC.) Guşul.
1136:
1118:
1100:
1073:
1070:
1069:
1068:
1058:
1057:
1056:
1047:
1035:
1023:
1001:
991:
981:
971:
970:
969:
960:
958:(Boiss.) Nyman
938:
928:
918:
908:
898:
888:
878:
876:Besser ex Link
868:
858:
857:
856:
847:
824:
813:
803:
793:
782:
772:
762:
752:
742:
732:
721:
711:
701:
694:Anchusa crispa
690:
680:
670:
659:
649:
647:(Nyman) Guşul.
639:
638:
637:
628:Anchusa azurea
625:
616:Anchusa azurea
613:
607:Anchusa azurea
597:Anchusa azurea
593:
583:
572:
562:
547:
544:
522:The subgenera
478:
475:
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136:
132:
131:
118:
117:
109:
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100:
99:
87:
86:
41:
39:
32:
15:
13:
10:
9:
6:
4:
3:
2:
1627:
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1603:
1602:
1600:
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1578:
1574:
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1529:
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1512:
1508:
1503:
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1224:
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1210:
1204:
1200:
1193:
1191:
1187:
1183:
1182:
1178:
1176:
1170:
1168:
1164:
1157:
1149:
1142:
1141:
1137:
1131:
1124:
1123:
1119:
1113:
1106:
1105:
1101:
1097:
1093:
1088:
1081:
1080:
1076:
1075:
1071:
1064:
1063:
1059:
1055:
1051:
1048:
1043:
1039:
1036:
1031:
1027:
1024:
1019:
1015:
1012:
1011:
1007:
1006:
1002:
997:
996:
992:
987:
986:
982:
977:
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964:
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904:
903:
899:
894:
893:
889:
884:
883:
879:
874:
873:
869:
864:
863:
862:Anchusa ovata
859:
855:
851:
848:
843:
839:
836:
835:
830:
829:
825:
819:
818:
814:
809:
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804:
799:
798:
794:
792:– Greece
788:
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753:
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629:
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621:
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614:
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608:
605:
604:
599:
598:
594:
589:
588:
584:
578:
577:
573:
568:
567:
563:
558:
557:
553:
552:
551:
545:
543:
541:
537:
533:
529:
525:
520:
518:
514:
509:
507:
503:
498:
496:
492:
488:
484:
476:
474:
472:
471:
466:
462:
458:
454:
452:
448:
444:
440:
436:
432:
431:
426:
425:
420:
417:The roots of
415:
413:
409:
404:
402:
398:
394:
390:
386:
381:
379:
375:
371:
367:
366:annual plants
362:
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344:
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339:
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317:
314:
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306:
301:
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268:
264:
257:
256:
251:
248:
244:
239:
234:
233:
229:
226:
225:
222:
221:Boraginoideae
219:
216:
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209:
206:
205:
202:
199:
196:
195:
192:
189:
186:
183:
182:
179:
176:
173:
170:
169:
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163:
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157:
156:
153:
152:Tracheophytes
150:
147:
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140:
137:
134:
133:
128:
123:
119:
116:
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83:
80:
72:
62:
58:
52:
51:
45:
40:
31:
30:
25:
23:
1247:
1226:
1222:
1198:
1179:
1174:
1147:
1138:
1129:
1120:
1111:
1102:
1086:
1077:
1060:
1053:
1049:
1041:
1037:
1029:
1025:
1017:
1013:
1003:
993:
983:
973:
966:
962:
954:
950:
940:
930:
920:
910:
900:
890:
880:
870:
860:
853:
849:
841:
837:
826:
815:
805:
795:
784:
774:
764:
754:
744:
734:
723:
713:
703:
692:
682:
672:
661:
651:
641:
631:
627:
619:
615:
610:
606:
595:
585:
580:(L.) M.Bieb.
574:
570:R.Br. ex DC.
564:
554:
549:
539:
535:
532:Buglossoides
531:
528:Buglossellum
527:
523:
521:
516:
512:
510:
505:
504:(containing
501:
499:
494:
491:Buglossellum
490:
487:Buglossoides
486:
482:
480:
468:
456:
455:
430:Lithospermum
428:
422:
418:
416:
412:rock gardens
407:
405:
382:
363:
351:Boraginaceae
341:
340:
335:
327:Lehm. (1817)
324:Stomotechium
323:
315:
307:
303:Mill. (1754)
299:
291:
283:
275:
253:
247:Type species
231:
230:
211:Boraginaceae
184:
171:
158:
145:
112:
96:
90:
75:
69:October 2013
66:
47:
24:-class sloop
21:
1427:iNaturalist
1272:Wikispecies
1030:lamprocarpa
1018:granatensis
886:Des.-Shost.
854:officinalis
465:Lepidoptera
292:Buglossites
217:Subfamily:
201:Boraginales
165:Angiosperms
61:introducing
1599:Categories
1158:References
1066:(L.) Lehm.
731:– Sardinia
632:macrocarpa
560:(L.) A.DC.
447:chloroform
433:) contain
406:The genus
359:introduced
276:Anchusella
44:references
1362:FloraBase
524:Buglossum
483:Buglossum
319:L. (1753)
308:Echioides
300:Buglossum
135:Kingdom:
1564:VicFlora
1553:VASCAN:
1546:40013633
1541:Tropicos
1323:eFloraSA
1257:Wikidata
1054:undulata
1042:sartorii
955:spruneri
477:Taxonomy
463:of some
439:resinoid
435:anchusin
316:Lycopsis
284:Buglossa
267:Synonyms
207:Family:
191:Asterids
178:Eudicots
1605:Anchusa
1406:5341194
1328:Anchusa
1278:Anchusa
1248:Anchusa
1223:Anchusa
1175:Anchusa
1052:subsp.
1040:subsp.
1028:subsp.
1016:subsp.
967:stylosa
965:subsp.
953:subsp.
946:M.Bieb.
852:subsp.
842:intacta
840:subsp.
821:M.Bieb.
719:Rech.f.
620:kurdica
546:Species
540:Anchusa
536:Anchusa
517:Limbata
513:Anchusa
502:Anchusa
495:Anchusa
457:Anchusa
443:alcohol
424:Alkanna
419:Anchusa
408:Anchusa
385:flowers
355:species
342:Anchusa
232:Anchusa
227:Genus:
197:Order:
139:Plantae
97:Anchusa
57:improve
22:Anchusa
1515:PLANTS
1507:884763
1491:NZOR:
1445:4451-1
1393:101622
1380:101622
1289:188064
1286:APDB:
1263:Q27946
1205:
1096:Vitman
989:Guşul.
906:Gușul.
896:Valdés
740:Ledeb.
709:Guşul.
667:Thunb.
657:Boiss.
611:azurea
461:larvae
393:petals
389:sepals
378:leaves
347:family
240:(1753)
46:, but
1520:ANCHU
1484:89629
1458:31714
1432:61789
1367:21968
1354:1ANCG
1341:70954
1302:88342
1151:A.DC.
916:A.DC.
866:Lehm.
770:Yıld.
688:Mill.
630:var.
618:var.
609:var.
601:Mill.
451:ether
397:cymes
338:genus
185:Clade
172:Clade
159:Clade
146:Clade
1528:POWO
1479:NCBI
1453:ITIS
1440:IPNI
1414:GRIN
1401:GBIF
1349:EPPO
1297:APNI
1203:ISBN
1146:(as
1128:(as
1110:(as
1092:All.
1085:(as
999:Post
750:Ten.
698:Viv.
678:Lam.
591:Ball
530:and
493:and
449:and
427:and
401:bees
372:and
336:The
1577:WFO
1556:818
1466:NBN
1419:619
1388:FoC
1375:FNA
1336:EoL
1315:VHH
1310:CoL
1601::
1579::
1566::
1543::
1530::
1517::
1504::
1481::
1468::
1455::
1442::
1429::
1416::
1403::
1390::
1377::
1364::
1351::
1338::
1325::
1312::
1299::
1274::
1259::
1189:^
1177:L.
1166:^
1133:L.
1094:)
1009:L.
832:L.
526:,
497:.
489:,
485:,
473:.
453:.
445:,
414:.
403:.
368:,
260:L.
238:L.
187::
174::
161::
148::
1211:.
1153:)
1117:)
1090:(
349:(
82:)
76:(
71:)
67:(
53:.
26:.
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