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Anomalites

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specimen is incomplete, with the lower legs missing, and the body preserved as a quartz cast attached to the matrix by a thin ridge of matrix. Frič was of the opinion that the shrunken wing casings indicated an individual still finishing pupating, the small cavity containing the beetle being a cast
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factory owned by Gabrial Zizka. During the carving process, Spigl encountered a small pocket in the freshwater quartz with the quartz cast replacement of the beetle preserved inside. The beetle and cavity were saved and shown to Zizka, who in turn entrusted same to Czech paleontologist Antonín Frič
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asking if any other fossils like it were known from the freshwater quartzes. Gaudry in turn queried a number of other French geology workers. All questioned were in agreement that the specimen was unique, with no other examples known. As such, and in compliance with paleontologist
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in 1885. Based on the limited characteristics present and visible though the outer quartz shell, he determined that the beetle was similar to the modern genus
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for examination. Frič presented the specimen in a talk on March 7, 1884, and then in a paper printed by the
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Krell, F.T. (2000). "The fossil record of Mesozoic and tertiary Scarabaeoidea (Coleoptera: Polyphaga)".
152: 301:'s philosophy that fossils should belong to their country of origin, Frič arranged the return of the 236: 561: 306: 293: 48: 538: 212: 171: 520: 494: 424: 525: 368: 298: 232: 224: 555: 397: 425:"Index to the Known Fossil Insects of the World: Including Myriapods and Arachnids" 190: 115: 499: 485: 239:, France before 1885. The quartz deposits were initially dated only as generally 244: 267: 194: 187: 85: 65: 292:
Frič sent illustrations of the specimen to prominent French paleontologist
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Fr.) aus dem tretiären Süfswasserquarz von Nogent le Rotrou in Frankreich"
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as a reference to the fossil's discovery so far from its origin spot.
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The only known specimen was recovered from "freshwater quartz" (
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and head shield are very similar to the cockchafers of genus
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of the cocoon itself. He noted that the shapes of both the
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in reflection of that. He coined the species epithet
469: 263:fossil was discovered by V. Spigl, a worker in the 211:from Central France and was first described by 8: 457: 392: 390: 388: 386: 384: 382: 311:National Museum of Natural History, France 31: 20: 351: 305:beetle to France. It was given to the 439:"Nogent-le-Rotrou (Eocene of France)" 203:. The species was found preserved in 7: 406:Deutsche entomologische Zeitschrift 273:Royal Bohemian Society of Sciences 14: 281:, and gave it the new genus name 398:"Ueber einen fossilen Maikäfer ( 52: 16:Extinct genus of scarab beetle 1: 592:Fossil taxa described in 1885 587:Taxa named by Antonin Fritsch 567:Monotypic Scarabaeidae genera 441:. The Paleobiology Database 423:Scudder, Samuel H. (1891). 608: 255:History and classification 158: 151: 49:Scientific classification 47: 39: 30: 23: 361:Invertebrate Systematics 186:is an extinct genus of 326: 228: 534:Paleobiology Database 324: 43:preserved in quartz. 572:Fossil beetle genera 400:Anomalites fugitivus 331:Anomalites fugitivus 261:Anomalites fugitivus 200:Anomalites fugitivus 165:Anomalites fugitivus 41:Anomalites fugitivus 237:Centre-Val de Loire 193:which contains the 188:shining leaf chafer 327: 307:Jardin des plantes 294:Jean Albert Gaudry 582:Fossils of France 549: 548: 521:Open Tree of Life 463:Taxon identifiers 179: 178: 144:A. fugitivus 599: 542: 541: 529: 528: 516: 515: 503: 502: 490: 489: 488: 458: 451: 450: 448: 446: 435: 429: 428: 420: 414: 413: 412:: 409–410. 1889. 394: 377: 376: 356: 299:Joachim Barrande 233:Nogent-le-Rotrou 167: 163: 127: 57: 56: 35: 21: 607: 606: 602: 601: 600: 598: 597: 596: 552: 551: 550: 545: 537: 532: 524: 519: 511: 506: 498: 493: 484: 483: 478: 465: 455: 454: 444: 442: 437: 436: 432: 422: 421: 417: 396: 395: 380: 373:10.1071/IT00031 358: 357: 353: 348: 319: 259:The only known 257: 221: 175: 169: 161: 160: 147: 125: 51: 17: 12: 11: 5: 605: 603: 595: 594: 589: 584: 579: 574: 569: 564: 554: 553: 547: 546: 544: 543: 530: 517: 504: 491: 475: 473: 467: 466: 461: 453: 452: 430: 415: 378: 367:(6): 871–905. 350: 349: 347: 344: 318: 315: 309:, home of the 256: 253: 231:) quarried in 229:Süßwasserquarz 220: 217: 177: 176: 170: 156: 155: 149: 148: 140: 138: 134: 133: 123: 119: 118: 113: 109: 108: 103: 99: 98: 93: 89: 88: 83: 79: 78: 73: 69: 68: 63: 59: 58: 45: 44: 37: 36: 28: 27: 15: 13: 10: 9: 6: 4: 3: 2: 604: 593: 590: 588: 585: 583: 580: 578: 577:Eocene France 575: 573: 570: 568: 565: 563: 560: 559: 557: 540: 535: 531: 527: 522: 518: 514: 509: 505: 501: 496: 492: 487: 481: 477: 476: 474: 472: 468: 464: 459: 440: 434: 431: 426: 419: 416: 411: 407: 403: 401: 393: 391: 389: 387: 385: 383: 379: 374: 370: 366: 362: 355: 352: 345: 343: 341: 337: 332: 323: 316: 314: 312: 308: 304: 300: 295: 290: 288: 284: 280: 279: 274: 269: 266: 262: 254: 252: 250: 246: 242: 238: 234: 230: 226: 218: 216: 214: 210: 206: 202: 201: 196: 192: 191:scarab beetle 189: 185: 184: 173: 168: 166: 157: 154: 153:Binomial name 150: 146: 145: 139: 136: 135: 132: 131: 124: 121: 120: 117: 114: 111: 110: 107: 104: 101: 100: 97: 94: 91: 90: 87: 84: 81: 80: 77: 74: 71: 70: 67: 64: 61: 60: 55: 50: 46: 42: 38: 34: 29: 26: 22: 19: 470: 445:February 20, 443:. Retrieved 433: 418: 409: 405: 399: 364: 360: 354: 339: 330: 328: 325:ventral view 303:A. fugitivus 302: 291: 286: 282: 276: 260: 258: 222: 219:Distribution 213:Antonín Frič 199: 198: 182: 181: 180: 164: 159: 143: 142: 129: 128: 116:Scarabaeidae 40: 24: 18: 317:Description 556:Categories 471:Anomalites 346:References 283:Anomalites 245:Priabonian 183:Anomalites 130:Anomalites 106:Coleoptera 86:Arthropoda 25:Anomalites 562:Rutelinae 486:Q31079892 336:scutellum 287:fugitivus 268:millstone 137:Species: 72:Kingdom: 66:Eukaryota 480:Wikidata 241:Tertiary 197:species 195:solitary 112:Family: 82:Phylum: 76:Animalia 62:Domain: 526:5333972 513:6757651 340:Anomala 278:Anomala 247:of the 162:† 141:† 122:Genus: 102:Order: 96:Insecta 92:Class: 539:168837 265:Prague 249:Eocene 225:German 209:quartz 205:Eocene 174:, 1885 508:GBIF 447:2024 329:The 207:age 172:Frič 500:WRG 495:CoL 369:doi 235:of 558:: 536:: 523:: 510:: 497:: 482:: 410:II 408:. 404:. 381:^ 365:14 363:. 342:. 251:. 227:: 215:. 449:. 427:. 375:. 371:: 126:†

Index


Scientific classification
Edit this classification
Eukaryota
Animalia
Arthropoda
Insecta
Coleoptera
Scarabaeidae
Anomalites
Binomial name
Frič
shining leaf chafer
scarab beetle
solitary
Eocene
quartz
Antonín Frič
German
Nogent-le-Rotrou
Centre-Val de Loire
Tertiary
Priabonian
Eocene
Prague
millstone
Royal Bohemian Society of Sciences
Anomala
Jean Albert Gaudry
Joachim Barrande

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