33:
46:
609:
This starfish has five arms, broad at the base but tapering to acute tips, four times as long as broad. The central disc of specimens preserved by drying is rather broad. The dorsal skeleton is reticulated and rather weak, thus specimens preserved in alcohol are soft and flaccid. There are numerous
618:
and small and numerous spines on the dorsal surface. The dorsal surface is entirely covered by an areolate or reticulate pattern; the areolations are 1.5-2mm across. The arm length is up to 17cm; on average, the ratio between the length of the arm and the
647:, which differs from it by virtue of less dense spines near the mouth, but longer and larger spines dorsally and marginally, and larger and more numerous pedicellariae. Compared with
641:
species within its range due to the finely and regularly areolated dorsal surface, the reticulated and flaccid skeleton, and small spines. He found it most resembling
1246:
1365:
1233:
1259:
309:
it may have first been collected in 1881-1883 by the
International Polar Expedition to Point Barrow, Alaska, where some starfish of the genus
1272:
505:
being the most divergent among them (the only specimen
Verrill was able examine), but all within the diversity to Russian specimens of
1411:
1020:
1378:
1311:
563:
453:
1264:
1143:
342:
1040:
1010:
957:
32:
653:
it has much more numerous, though smaller dorsal and lateral spines, these being differently arranged. Compared with
45:
1175:
887:"Monograph of the shallow-water starfishes of the North Pacific coast from the Arctic Ocean to California"
1077:
1036:
882:
841:
390:
318:
306:
157:
140:
509:, and thus maintaining recognition for this taxon was unwarranted. In 1950 Djakonov described one new
1383:
1316:
1220:
1112:
798:
704:
648:
643:
456:
maintains this classification despite following
Djakonov's recognition of the variety as a form of
1303:
1081:
724:
654:
419:
378:
166:
40:
695:
in the Bering Sea. It has been found at depths of 9-170m. Djakonov complies the distribution of
1251:
1406:
1352:
1285:
1207:
1016:
708:
1290:
886:
846:"Description of new genera and species of starfishes from the North Pacific coast of America"
978:
898:
845:
712:
688:
559:
469:
350:
346:
719:, usually at depths of 20m, but in America specimens have been recovered down to 170m. The
972:
742:
334:
290:
953:
680:
593:
518:
436:
1400:
659:, it has more and shorter dorsal spines. Djakonov found it to be the most similar to
489:
97:
1357:
754:
728:
611:
366:
338:
326:
1212:
1343:
1298:
974:
Report on the
International Polar Expedition to Point Barrow, Alaska (1881-1883)
741:
was first recovered in 1910 at a depth of 53m from a gravelly sea bottom in the
716:
1198:
684:
510:
498:
473:
354:
294:
107:
77:
982:
903:
57:
1082:"Asteroidea of the North Pacific and Adjacent Waters, Part 3: Forcipulata"
1337:
1192:
768:
637:
624:
597:
502:
444:, accepting all other of Verrill's forms. Fisher synonymised the variety
278:
117:
87:
1370:
1238:
282:
692:
676:
620:
615:
497:, he stated that it is only known from five specimens recovered from
370:
314:
286:
67:
1169:
535:. Jangoux and Lawrence followed Djakonov in 2001, but recognised f.
423:. In 1950 Alexander Michailovitsch Djakonov reinstated the taxon as
614:
on both the dorsal and lateral surfaces. There are also many small
1041:"A preliminary synopsis of the Asteriidae, a family of sea-stars"
1277:
1225:
1173:
381:
in the southeast of Norton Sound. In 1914 Verrill raised var.
745:
in the Bering Sea off the northeastern
Kamchatka peninsula.
703:, now considered a synonym of the nominate type, as off the
460:, and Jangoux and Lawrence also largely following Djakonov.
1154:. Zoological Institute of the Russian Academy of Sciences
663:, and Fisher went further by considering it identical to
977:. Washington D.C.: General Post Office. p. 159.
1152:
1327:
1182:
1009:Jangoux, Michel; Lawrence, John M. (1 June 2001).
1092:(3). U.S. Government Printing Office: 6, 14, 204
484:. Fisher considered all three to be synonyms of
377:was described by Verrill in this 1909 work from
321:. The species was first described by Verrill as
943:
941:
501:in Alaska, these varied in spination, with the
939:
937:
935:
933:
931:
929:
927:
925:
923:
921:
757:off Alaska, and has not been found elsewhere.
1004:
1002:
1000:
998:
996:
994:
992:
877:
875:
873:
871:
836:
834:
832:
830:
828:
826:
824:
8:
1072:
1070:
1068:
1066:
767:is known from the Sea of Okhotsk and around
1170:
1123:. World Register of Marine Species (WoRMS)
1015:. CRC Press. pp. 243, 250, 253, 262.
539:as a synonym of the nominate, and classed
523:of the nominate subspecies: the nominate,
31:
20:
902:
807:World Register of Marine Species (WoRMS)
792:
790:
788:
786:
784:
780:
1086:United States National Museum Bulletin
1045:Annals and Magazine of Natural History
687:, and in the United States around the
7:
1144:"Sea Stars (Asteroidea) Catalogue •
468:Verrill originally recognised three
723:form was originally recovered off
635:Verrill found it a quite distinct
14:
600:. The arm length is up to 14.6cm.
564:World Register of Marine Species
454:World Register of Marine Species
44:
566:by Christopher Mah as of 2008:
317:on a pebbly seabed just off of
313:were collected at a depth of 7
263:(Verrill, 1909) Djakonov, 1950
251:(Verrill, 1909) Djakonov, 1950
1:
623:of its disc is 4.3:1. It has
343:United States National Museum
187:(Verrill, 1909) Verrill, 1914
1111:Mah, Christopher L. (2008).
797:Mah, Christopher L. (2008).
850:American Journal of Science
809:. Flanders Marine Institute
385:to an independent species,
1428:
950:Морские звезды морей СССР
401:, and in 1930 synonymised
1412:Animals described in 1881
1142:Smirnov, Igor S. (2019).
1121:World Asteroidea Database
897:: 189–191, 193–194, 196.
715:off Alaska, north to the
289:in the United States and
172:
165:
146:
139:
41:Scientific classification
39:
30:
23:
671:Distribution and habitat
983:10.5962/bhl.title.31919
948:Дьяконов, А.М. (1950).
904:10.5962/bhl.title.25926
464:Intraspecific variation
1078:Fisher, Walter Kenrick
1037:Fisher, Walter Kenrick
971:Murdoch, John (1885).
891:Harriman Alaska Series
883:Verrill, Addison Emery
842:Verrill, Addison Emery
517:, and recognised four
160:, 1909) Djakonov, 1950
679:it is known from the
391:Walter Kenrick Fisher
349:. He described three
307:Addison Emery Verrill
1329:Allasterias rathbuni
1012:Echinoderm studies 6
960:. pp. 127, 128.
958:Акаде́мии Нау́к СССР
727:in the southeast of
562:are accepted in the
337:, curator of marine
323:Allasterias rathbuni
204:Allasterias rathbuni
192:Allasterias rathbuni
176:Allasterias rathbuni
705:Kamchatka peninsula
644:Asterias rollestoni
387:Allasterias anomala
184:Allasterias anomala
16:Species of starfish
1115:Asterias amurensis
725:St. Michael Island
486:Asterias amurensis
420:Asterias amurensis
379:St. Michael Island
333:. He commemorated
329:for his new genus
216:Asterias alveolata
1394:
1393:
1286:Open Tree of Life
1184:Asterias rathbuni
1176:Taxon identifiers
801:Asterias rathbuni
761:Asterias rathbuni
753:was found in the
735:Asterias rathbuni
709:Commander Islands
697:Asterias rathbuni
591:
579:
493:. Regarding var.
365:(named after the
274:Asterias rathbuni
270:
269:
264:
256:Asterias rathbuni
252:
244:Asterias rathbuni
240:
232:Asterias rathbuni
228:
220:
212:
200:
188:
180:
150:Asterias rathbuni
25:Asterias rathbuni
1419:
1387:
1386:
1374:
1373:
1361:
1360:
1348:
1347:
1346:
1320:
1319:
1307:
1306:
1294:
1293:
1281:
1280:
1268:
1267:
1255:
1254:
1242:
1241:
1229:
1228:
1216:
1215:
1203:
1202:
1201:
1171:
1164:
1163:
1161:
1159:
1139:
1133:
1132:
1130:
1128:
1108:
1102:
1101:
1099:
1097:
1074:
1061:
1060:
1058:
1056:
1033:
1027:
1026:
1006:
987:
986:
968:
962:
961:
945:
916:
915:
913:
911:
906:
879:
866:
865:
863:
861:
838:
819:
818:
816:
814:
803:(Verrill, 1909)"
794:
713:Shumagin Islands
689:Aleutian Islands
589:
577:
547:as varieties of
347:specific epithet
293:. There are two
262:
250:
238:
226:
224:Asterias anomala
219:(Djakonov, 1950)
218:
210:
198:
186:
178:
152:
132:A. rathbuni
49:
48:
35:
21:
1427:
1426:
1422:
1421:
1420:
1418:
1417:
1416:
1397:
1396:
1395:
1390:
1382:
1377:
1369:
1364:
1356:
1351:
1342:
1341:
1336:
1323:
1315:
1310:
1302:
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1284:
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1237:
1232:
1224:
1219:
1211:
1206:
1197:
1196:
1191:
1178:
1168:
1167:
1157:
1155:
1141:
1140:
1136:
1126:
1124:
1110:
1109:
1105:
1095:
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1076:
1075:
1064:
1054:
1052:
1035:
1034:
1030:
1023:
1008:
1007:
990:
970:
969:
965:
947:
946:
919:
909:
907:
881:
880:
869:
859:
857:
840:
839:
822:
812:
810:
796:
795:
782:
777:
743:Karaginsky Gulf
707:and around the
673:
633:
631:Similar species
607:
592:- Found in the
578:(Verrill, 1909)
466:
427:, and subsumed
373:). The variety
335:Richard Rathbun
325:in 1909 as the
303:
291:Far East Russia
227:(Verrill, 1909)
161:
154:
148:
135:
43:
17:
12:
11:
5:
1425:
1423:
1415:
1414:
1409:
1399:
1398:
1392:
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1388:
1375:
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1349:
1333:
1331:
1325:
1324:
1322:
1321:
1308:
1295:
1282:
1269:
1256:
1243:
1230:
1217:
1204:
1188:
1186:
1180:
1179:
1174:
1166:
1165:
1134:
1103:
1062:
1028:
1021:
988:
963:
954:St. Petersburg
952:(in Russian).
917:
867:
820:
779:
778:
776:
773:
711:, east to the
681:Sea of Okhotsk
672:
669:
632:
629:
606:
603:
602:
601:
594:Sea of Okhotsk
590:Djakonov, 1950
580:
465:
462:
302:
299:
281:native to the
268:
267:
266:
265:
253:
241:
239:Djakonov, 1950
229:
221:
213:
201:
189:
181:
170:
169:
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155:
144:
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137:
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129:
127:
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100:
95:
91:
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75:
71:
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65:
61:
60:
55:
51:
50:
37:
36:
28:
27:
15:
13:
10:
9:
6:
4:
3:
2:
1424:
1413:
1410:
1408:
1405:
1404:
1402:
1385:
1380:
1376:
1372:
1367:
1363:
1359:
1354:
1350:
1345:
1339:
1335:
1334:
1332:
1330:
1326:
1318:
1313:
1309:
1305:
1300:
1296:
1292:
1287:
1283:
1279:
1274:
1270:
1266:
1261:
1257:
1253:
1248:
1244:
1240:
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1227:
1222:
1218:
1214:
1209:
1205:
1200:
1194:
1190:
1189:
1187:
1185:
1181:
1177:
1172:
1153:
1149:
1147:
1138:
1135:
1122:
1118:
1117:Lutken, 1871"
1116:
1107:
1104:
1091:
1087:
1083:
1079:
1073:
1071:
1069:
1067:
1063:
1051:(9): 248, 598
1050:
1046:
1042:
1038:
1032:
1029:
1024:
1022:9789058093011
1018:
1014:
1013:
1005:
1003:
1001:
999:
997:
995:
993:
989:
984:
980:
976:
975:
967:
964:
959:
955:
951:
944:
942:
940:
938:
936:
934:
932:
930:
928:
926:
924:
922:
918:
905:
900:
896:
892:
888:
884:
878:
876:
874:
872:
868:
855:
851:
847:
843:
837:
835:
833:
831:
829:
827:
825:
821:
808:
804:
802:
793:
791:
789:
787:
785:
781:
774:
772:
770:
766:
762:
758:
756:
752:
748:
744:
740:
736:
732:
730:
726:
722:
718:
714:
710:
706:
702:
698:
694:
690:
686:
682:
678:
670:
668:
666:
662:
658:
657:
652:
651:
650:A. versicolor
646:
645:
640:
639:
630:
628:
626:
622:
617:
613:
612:pedicellariae
604:
599:
595:
588:
584:
581:
576:
572:
569:
568:
567:
565:
561:
556:
554:
550:
546:
542:
538:
534:
530:
526:
522:
521:
516:
512:
508:
504:
500:
496:
492:
491:
490:sensu stricto
487:
483:
479:
475:
472:in 1909: the
471:
463:
461:
459:
455:
452:in 1930; the
451:
447:
443:
440:
439:
434:
430:
426:
422:
421:
416:
412:
408:
404:
400:
396:
392:
388:
384:
380:
376:
372:
368:
364:
360:
356:
352:
348:
344:
340:
339:invertebrates
336:
332:
328:
324:
320:
319:Port Clarence
316:
312:
308:
305:According to
300:
298:
296:
292:
288:
284:
280:
276:
275:
261:
257:
254:
249:
245:
242:
237:
233:
230:
225:
222:
217:
214:
211:Verrill, 1909
209:
205:
202:
199:Verrill, 1909
197:
193:
190:
185:
182:
179:Verrill, 1909
177:
174:
173:
171:
168:
164:
159:
153:
151:
145:
142:
141:Binomial name
138:
134:
133:
128:
125:
124:
121:
120:
116:
113:
112:
109:
106:
103:
102:
99:
98:Forcipulatida
96:
93:
92:
89:
86:
83:
82:
79:
78:Echinodermata
76:
73:
72:
69:
66:
63:
62:
59:
56:
53:
52:
47:
42:
38:
34:
29:
26:
22:
19:
1328:
1183:
1156:. Retrieved
1151:
1145:
1137:
1125:. Retrieved
1120:
1114:
1106:
1094:. Retrieved
1089:
1085:
1053:. Retrieved
1048:
1044:
1031:
1011:
973:
966:
949:
908:. Retrieved
894:
890:
858:. Retrieved
853:
849:
811:. Retrieved
806:
800:
765:crassispinus
764:
760:
759:
755:Norton Sound
750:
746:
738:
734:
733:
729:Norton Sound
720:
700:
696:
674:
665:A. amurensis
664:
661:A. amurensis
660:
656:A. amurensis
655:
649:
642:
636:
634:
608:
587:crassispinis
586:
582:
574:
570:
557:
552:
548:
544:
540:
536:
532:
528:
524:
519:
515:crassispinis
514:
507:A. amurensis
506:
494:
488:
485:
481:
477:
467:
457:
450:A. amurensis
449:
445:
441:
437:
432:
428:
424:
418:
414:
410:
406:
402:
398:
394:
393:synonymised
386:
382:
374:
367:Norton Sound
362:
358:
330:
327:type species
322:
310:
304:
273:
272:
271:
259:
255:
247:
243:
235:
231:
223:
215:
207:
203:
195:
191:
183:
175:
149:
147:
131:
130:
118:
24:
18:
1299:SeaLifeBase
1158:12 November
1127:10 November
1096:14 November
1055:15 November
910:11 November
860:11 November
813:11 November
751:nortonensis
747:A. rathbuni
717:Chukchi Sea
605:Description
596:and around
583:A. rathbuni
571:A. rathbuni
549:A. rathbuni
545:nortonensis
533:nortonensis
495:nortonensis
482:nortonensis
458:A. rathbuni
446:nortonensis
433:A. rathbuni
425:A. rathbuni
415:nortonensis
395:Allasterias
363:nortonensis
331:Allasterias
260:nortonensis
208:nortonensis
1401:Categories
775:References
731:, Alaska.
685:Bering Sea
625:planktonic
560:subspecies
511:subspecies
499:Norton Bay
429:A. anomala
389:. In 1923
295:subspecies
285:coasts of
108:Asteriidae
88:Asteroidea
1344:Q62011216
739:alveolata
627:larvae.
541:alveolata
525:alveolata
480:and var.
470:varieties
361:and var.
351:varieties
345:, in the
236:alveolata
126:Species:
64:Kingdom:
58:Eukaryota
1407:Asterias
1338:Wikidata
1252:10798453
1199:Q2045020
1193:Wikidata
1146:Asterias
1080:(1930).
1039:(1923).
885:(1914).
856:: 65, 66
844:(1909).
769:Sakhalin
638:Asterias
598:Sakhalin
575:rathbuni
553:rathbuni
474:nominate
411:rathbuni
407:rathbuni
399:Asterias
355:nominate
311:Asterias
301:Taxonomy
279:starfish
167:Synonyms
119:Asterias
104:Family:
74:Phylum:
68:Animalia
54:Domain:
1371:8897597
1291:3657160
1239:5187518
763:subsp.
721:anomala
701:anomala
691:and in
616:papulae
537:anomala
529:anomala
478:anomala
476:, var.
442:anomala
403:anomala
383:anomala
375:anomala
359:anomala
357:, var.
341:at the
315:fathoms
283:Pacific
248:anomala
196:anomala
158:Verrill
114:Genus:
94:Order:
84:Class:
1384:880779
1317:378804
1278:378804
1265:157218
1226:597576
1019:
693:Alaska
677:Russia
621:radius
610:small
520:formae
431:under
371:Alaska
353:: the
287:Alaska
1379:WoRMS
1358:5TVHD
1312:WoRMS
1304:48660
1247:IRMNG
585:ssp.
573:ssp.
551:ssp.
448:with
438:forma
417:with
413:var.
397:with
277:is a
206:var.
194:var.
1366:GBIF
1273:OBIS
1260:ITIS
1234:GBIF
1213:HRM3
1160:2019
1129:2019
1098:2019
1057:2019
1017:ISBN
912:2019
862:2019
815:2019
683:and
558:Two
543:and
531:and
503:type
409:and
1353:CoL
1221:EoL
1208:CoL
979:doi
899:doi
771:.
749:f.
737:f.
699:f.
675:In
667:.
435:as
369:in
258:f.
246:f.
234:f.
1403::
1381::
1368::
1355::
1340::
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