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Bembidion

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191: 33: 55: 1278: 1612: 1481: 1520: 254:, some of which are elevated to full genus rank by various authors; as noted above, however, no universally accepted way of splitting the genus exists yet. 219:, meaning they are found in both the Northern and Southern Hemispheres, but not in the tropics. In warmer regions it is substituted by closely related 1455: 1494: 215:
by number of species. All species are small (less than 7.5 mm) and move very fast. Most of them live close to water. The genus has a
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in 1941 and by G.G. Perrault in 1981, but none of them have been generally accepted.
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There have been many attempts to divide it into smaller genera, most notably by
1317: 126: 1416: 251: 212: 86: 66: 1277: 1377: 1311: 1356: 1460: 1382: 1473: 221: 106: 96: 76: 1288: 1421: 1556: 208: 189: 1206:. Leiden - Copenhagen: Brill - Scandinavian Science Press, 1985. 1292: 1408: 1269: 1227:. Cambridge: Harvard University Press, 1965. P. 22, 45. 1204:
The Carabidae (Coleoptera) of Fennoscandia and Denmark
1301: 1225:Biogeography of the Southern End of the World 8: 1289: 31: 20: 1161: 1198: 1196: 1194: 1192: 7: 1613:Taxa named by Pierre André Latreille 1562:EB2C87C2-FF81-FFDD-5EBF-F8BEFEA86834 1536:72a0686c-04f9-4a5f-b44d-2ae83790eb61 1448:bdffee9d-875c-4862-8454-af8c6166e2bd 250:This genus is divided into numerus 14: 1276: 53: 1149:J.Schmidt & Michalik, 2017 1: 555:J.Schmidt & Marggi, 2014 1223:Philip Jackson Darlington. 217:biantitropical distribution 1629: 451:Moret & Toledano, 2002 232: 1184:Bembidion quadrimaculatum 1140:Ortuño & Arillo, 2010 211:of beetles in the family 172: 165: 158: 151: 50:Scientific classification 48: 42:Bembidion quadrimaculatum 39: 30: 23: 755:Bonniard de Saludo, 1970 1177:Carabus quadrimaculatus 1169:Hist. Nat. Crust. Ins. 199: 1285:at Wikimedia Commons 1027:Müller-Motzfeld, 1998 931:Müller-Motzfeld, 1998 779:Müller-Motzfeld, 1986 699:Habu & Baba, 1968 523:Müller-Motzfeld, 1988 307:Habu & Ueno, 1955 193: 160:At least 1400 species 1443:Fauna Europaea (new) 1156:References and notes 851:Tschitscherine, 1895 196:Bembidion tetracolum 360:Bembidionetolitzkya 258:subgenera include: 1202:Carl H. Lindroth. 1137:Archaeophilochthus 225:and other genera. 200: 1585: 1584: 1544:Open Tree of Life 1295:Taxon identifiers 1281:Media related to 1272:A list of species 1246:Catalogue of Life 1150: 1141: 1132: 1124: 1116: 1108: 1100: 1092: 1084: 1076: 1075:Motschulsky, 1864 1068: 1060: 1052: 1044: 1043:Motschulsky, 1864 1036: 1028: 1020: 1019:Motschulsky, 1864 1012: 1004: 996: 988: 980: 972: 964: 960:Pseudophilochthus 956: 948: 940: 939:Motschulsky, 1864 932: 924: 916: 915:Motschulsky, 1864 908: 907:Motschulsky, 1844 900: 892: 888:Philochthemphanes 884: 876: 868: 860: 852: 844: 836: 828: 820: 812: 804: 796: 788: 780: 772: 764: 756: 748: 740: 732: 724: 716: 708: 700: 692: 684: 683:Motschulsky, 1864 676: 668: 660: 659:Motschulsky, 1864 652: 644: 636: 628: 627:Motschulsky, 1850 620: 612: 604: 596: 588: 580: 572: 564: 556: 548: 540: 532: 524: 516: 508: 504:Gondwanabembidion 500: 492: 484: 476: 468: 460: 459:Motschulsky, 1850 452: 444: 436: 432:Desarmatocillenus 428: 420: 412: 411:Motschulsky, 1864 404: 396: 388: 380: 376:Blepharoplataphus 372: 364: 356: 348: 340: 332: 324: 316: 308: 300: 292: 284: 276: 268: 267:Motschulsky, 1864 188: 187: 147: 1620: 1608:Carabidae genera 1578: 1577: 1565: 1564: 1552: 1551: 1539: 1538: 1529: 1528: 1516: 1515: 1513:NHMSYS0001717141 1503: 1502: 1490: 1489: 1477: 1476: 1464: 1463: 1451: 1450: 1438: 1437: 1425: 1424: 1412: 1411: 1399: 1398: 1386: 1385: 1373: 1372: 1360: 1359: 1350: 1349: 1337: 1336: 1335: 1322: 1321: 1320: 1290: 1280: 1256: 1255: 1253: 1252: 1242:Latreille, 1802" 1234: 1228: 1221: 1215: 1200: 1187: 1166: 1148: 1139: 1130: 1122: 1114: 1106: 1098: 1091:Netolitzky, 1931 1090: 1083:Netolitzky, 1914 1082: 1074: 1067:Netolitzky, 1918 1066: 1058: 1051:Netolitzky, 1935 1050: 1048:Thaumatoperyphus 1042: 1035:Ganglbauer, 1891 1034: 1026: 1018: 1010: 1008:Taiwanobembidion 1003:Netolitzky, 1931 1002: 995:Netolitzky, 1910 994: 986: 978: 970: 962: 954: 947:Netolitzky, 1920 946: 938: 930: 922: 914: 906: 898: 891:Netolitzky, 1943 890: 882: 875:Netolitzky, 1939 874: 866: 858: 850: 842: 834: 827:Netolitzky, 1914 826: 819:Netolitzky, 1920 818: 810: 803:Netolitzky, 1914 802: 795:Netolitzky, 1931 794: 786: 778: 771:Clairville, 1806 770: 762: 754: 746: 738: 730: 722: 715:Netolitzky, 1914 714: 707:Netolitzky, 1920 706: 698: 696:Nipponobembidion 690: 682: 674: 666: 658: 651:Netolitzky, 1935 650: 642: 635:Netolitzky, 1921 634: 626: 618: 610: 602: 594: 586: 578: 570: 562: 554: 546: 538: 530: 522: 514: 506: 498: 491:Netolitzky, 1931 490: 483:Netolitzky, 1911 482: 474: 466: 458: 450: 442: 435:Netolitzky, 1939 434: 426: 418: 410: 403:Netolitzky, 1942 402: 394: 386: 379:Netolitzky, 1920 378: 370: 362: 354: 346: 344:Australoemphanes 339:Netolitzky, 1943 338: 330: 322: 314: 306: 298: 290: 283:Netolitzky, 1931 282: 275:Netolitzky, 1931 274: 266: 142: 58: 57: 35: 21: 16:Genus of beetles 1628: 1627: 1623: 1622: 1621: 1619: 1618: 1617: 1588: 1587: 1586: 1581: 1573: 1568: 1560: 1555: 1547: 1542: 1534: 1532: 1524: 1519: 1511: 1506: 1498: 1493: 1485: 1480: 1472: 1467: 1459: 1454: 1446: 1441: 1433: 1428: 1420: 1415: 1407: 1402: 1394: 1389: 1381: 1376: 1368: 1363: 1355: 1353: 1345: 1340: 1331: 1330: 1325: 1316: 1315: 1310: 1297: 1287: 1265: 1260: 1259: 1250: 1248: 1236: 1235: 1231: 1222: 1218: 1201: 1190: 1167: 1163: 1158: 1153: 1080:Trichoplataphus 963:Wollaston, 1877 944:Pseudometallina 920:Plocamoperyphus 632:Microserrullula 584:Limnaeoperyphus 467:Wollaston, 1877 419:Samouelle, 1819 355:Latreille, 1802 315:Wollaston, 1877 241: 231: 207:is the largest 141: 52: 17: 12: 11: 5: 1626: 1624: 1616: 1615: 1610: 1605: 1600: 1590: 1589: 1583: 1582: 1580: 1579: 1566: 1553: 1540: 1530: 1517: 1504: 1491: 1478: 1465: 1452: 1439: 1430:Fauna Europaea 1426: 1413: 1400: 1387: 1374: 1361: 1351: 1338: 1323: 1307: 1305: 1299: 1298: 1293: 1274: 1273: 1266: 1264: 1263:External links 1261: 1258: 1257: 1229: 1216: 1188: 1160: 1159: 1157: 1154: 1152: 1151: 1142: 1133: 1131:Lindroth, 1976 1125: 1123:Lindroth, 1976 1117: 1115:Lindroth, 1976 1109: 1107:Lindroth, 1976 1101: 1099:Lindroth, 1980 1093: 1085: 1077: 1069: 1061: 1053: 1045: 1037: 1029: 1021: 1013: 1005: 997: 989: 981: 976:Pseudotrepanes 973: 971:Toledano, 2005 965: 957: 952:Pseudoperyphus 949: 941: 933: 925: 917: 909: 901: 899:Stephens, 1828 893: 885: 877: 869: 861: 853: 845: 837: 829: 824:Paraprincidium 821: 813: 805: 797: 789: 781: 773: 765: 757: 749: 741: 733: 725: 717: 709: 704:Notaphemphanes 701: 693: 685: 677: 669: 667:Bousquet, 2006 661: 653: 648:Necpericompsus 645: 643:Toledano, 1998 637: 629: 621: 613: 611:Stephens, 1828 605: 597: 595:Maddison, 2012 589: 581: 573: 571:Toledano, 2000 565: 563:Toledano, 2008 557: 549: 541: 533: 525: 517: 515:Lindroth, 1963 512:Hirmoplataphus 509: 507:Toledano, 2005 501: 496:Gnatholymnaeum 493: 485: 477: 469: 461: 453: 445: 437: 429: 421: 413: 405: 397: 392:Chilioperyphus 389: 381: 373: 371:Toledano, 2000 365: 363:E.Strand, 1929 357: 349: 347:Toledano, 2005 341: 336:Aureoplataphus 333: 325: 320:Armatocillenus 317: 309: 301: 293: 288:Antiperyphanes 285: 277: 269: 260: 230: 227: 186: 185: 184: 183: 178: 170: 169: 163: 162: 156: 155: 149: 148: 134: 130: 129: 124: 120: 119: 114: 110: 109: 104: 100: 99: 94: 90: 89: 84: 80: 79: 74: 70: 69: 64: 60: 59: 46: 45: 37: 36: 28: 27: 15: 13: 10: 9: 6: 4: 3: 2: 1625: 1614: 1611: 1609: 1606: 1604: 1601: 1599: 1596: 1595: 1593: 1576: 1571: 1567: 1563: 1558: 1554: 1550: 1545: 1541: 1537: 1531: 1527: 1522: 1518: 1514: 1509: 1505: 1501: 1496: 1492: 1488: 1483: 1479: 1475: 1470: 1466: 1462: 1457: 1453: 1449: 1444: 1440: 1436: 1431: 1427: 1423: 1418: 1414: 1410: 1405: 1401: 1397: 1392: 1388: 1384: 1379: 1375: 1371: 1366: 1362: 1358: 1352: 1348: 1343: 1339: 1334: 1328: 1324: 1319: 1313: 1309: 1308: 1306: 1304: 1300: 1296: 1291: 1286: 1284: 1279: 1271: 1268: 1267: 1262: 1247: 1243: 1241: 1233: 1230: 1226: 1220: 1217: 1214:. P. 129-199. 1213: 1212:90-04-07727-8 1209: 1205: 1199: 1197: 1195: 1193: 1189: 1185: 1181: 1178: 1174: 1170: 1165: 1162: 1155: 1147: 1143: 1138: 1134: 1129: 1126: 1121: 1118: 1113: 1110: 1105: 1102: 1097: 1094: 1089: 1086: 1081: 1078: 1073: 1070: 1065: 1062: 1057: 1054: 1049: 1046: 1041: 1038: 1033: 1030: 1025: 1024:Terminophanes 1022: 1017: 1014: 1009: 1006: 1001: 998: 993: 990: 985: 982: 979:Jeannel, 1962 977: 974: 969: 968:Pseudosinocys 966: 961: 958: 953: 950: 945: 942: 937: 934: 929: 926: 923:Jeannel, 1962 921: 918: 913: 910: 905: 902: 897: 894: 889: 886: 881: 878: 873: 870: 865: 862: 859:Jeannel, 1941 857: 854: 849: 846: 843:Jeannel, 1941 841: 838: 833: 830: 825: 822: 817: 814: 811:Jeannel, 1962 809: 806: 801: 798: 793: 790: 787:LeConte, 1847 785: 782: 777: 774: 769: 766: 763:LeConte, 1847 761: 758: 753: 750: 747:Jeannel, 1962 745: 742: 739:Jeannel, 1962 737: 734: 731:Jeannel, 1962 729: 726: 721: 718: 713: 710: 705: 702: 697: 694: 689: 686: 681: 678: 673: 670: 665: 662: 657: 654: 649: 646: 641: 638: 633: 630: 625: 622: 617: 614: 609: 606: 601: 598: 593: 590: 585: 582: 577: 574: 569: 566: 561: 558: 553: 550: 547:Alluaud, 1917 545: 542: 539:LeConte, 1847 537: 534: 529: 526: 521: 518: 513: 510: 505: 502: 497: 494: 489: 486: 481: 478: 475:Jeannel, 1941 473: 470: 465: 462: 457: 454: 449: 446: 441: 438: 433: 430: 425: 422: 417: 414: 409: 406: 401: 400:Chinocillenus 398: 395:Jeannel, 1962 393: 390: 385: 382: 377: 374: 369: 366: 361: 358: 353: 350: 345: 342: 337: 334: 329: 326: 321: 318: 313: 310: 305: 302: 299:Jeannel, 1962 297: 294: 291:Jeannel, 1962 289: 286: 281: 278: 273: 270: 265: 262: 261: 259: 257: 253: 248: 246: 240: 238: 228: 226: 224: 223: 218: 214: 210: 206: 205: 198: 197: 192: 182: 179: 177: 174: 173: 171: 168: 164: 161: 157: 154: 150: 145: 140: 139: 135: 132: 131: 128: 125: 122: 121: 118: 115: 112: 111: 108: 105: 102: 101: 98: 95: 92: 91: 88: 85: 82: 81: 78: 75: 72: 71: 68: 65: 62: 61: 56: 51: 47: 44: 43: 38: 34: 29: 26: 22: 19: 1302: 1275: 1270:Tree of Life 1249:. Retrieved 1245: 1239: 1232: 1224: 1219: 1203: 1183: 1176: 1173:Type species 1168: 1164: 1145: 1136: 1127: 1119: 1112:Zeperyphodes 1111: 1103: 1095: 1087: 1079: 1071: 1064:Trepanedoris 1063: 1055: 1047: 1039: 1031: 1023: 1015: 1007: 999: 991: 983: 975: 967: 959: 951: 943: 935: 928:Politophanes 927: 919: 911: 903: 895: 887: 883:Dejean, 1821 879: 872:Peryphophila 871: 863: 855: 847: 839: 831: 823: 815: 807: 799: 791: 783: 775: 767: 760:Ochthedromus 759: 752:Notoperyphus 751: 743: 735: 727: 723:Dejean, 1821 719: 712:Notaphocampa 711: 703: 695: 687: 679: 671: 664:Neobembidion 663: 655: 647: 640:Microsinocys 639: 631: 623: 615: 607: 599: 592:Lindrochthus 591: 587:Nakane, 1963 583: 575: 567: 559: 551: 543: 535: 528:Hydriomicrus 527: 519: 511: 503: 495: 487: 480:Eupetedromus 479: 471: 464:Endosomatium 463: 455: 447: 439: 431: 423: 415: 407: 399: 391: 383: 375: 367: 359: 351: 343: 335: 331:Vysoky, 1986 328:Asioperyphus 327: 323:Dupuis, 1912 319: 311: 303: 296:Antiperyphus 295: 287: 279: 271: 263: 255: 249: 245:René Jeannel 242: 236: 220: 203: 202: 201: 194: 180: 175: 137: 136: 40: 24: 18: 1469:iNaturalist 1327:Wikispecies 1128:Zeplataphus 1104:Zemetallina 1059:Casey, 1918 1056:Trechonepha 1032:Testediolum 1000:Sloanephila 955:Hatch, 1950 896:Philochthus 867:Casey, 1918 856:Peryphiolus 848:Peryphidium 835:Csiki, 1901 792:Omoperyphus 691:Sharp, 1903 672:Neoemphanes 619:Casey, 1918 603:Casey, 1918 579:Casey, 1918 576:Leuchydrium 552:Jammuphanes 531:Casey, 1918 499:Sharp, 1903 443:Bedel, 1896 427:Casey, 1918 387:Bedel, 1879 368:Bembidromus 312:Apteromimus 272:Ananotaphus 1592:Categories 1251:2023-03-15 1182:, 1761 (= 1120:Zeperyphus 1096:Zecillenus 1088:Zeactedium 1011:Habu, 1973 987:Ueno, 1954 936:Princidium 864:Peryphodes 840:Peryphanes 808:Pacmophena 776:Ocyturanes 744:Nothonepha 736:Notholopha 688:Nesocidium 675:Habu, 1978 600:Liocosmius 560:Jedlickion 544:Hypsipezum 488:Furcacampa 472:Euperyphus 440:Diplocampa 424:Cyclolopha 408:Chlorodium 304:Aptenidium 233:See also: 181:Zecillenus 127:Bembidiina 123:Subtribe: 107:Coleoptera 87:Arthropoda 1603:Trechinae 1598:Bembidion 1347:Bembidion 1333:Bembidion 1303:Bembidion 1283:Bembidion 1240:Bembidion 1146:Eodontium 1040:Testedium 992:Semicampa 984:Sakagutia 912:Plataphus 800:Omotaphus 768:Ocydromus 624:Metallina 616:Melomalus 520:Hoquedela 352:Bembidion 280:Andrewesa 256:Bembidion 252:subgenera 237:Bembidion 213:Carabidae 204:Bembidion 176:Sakagutia 153:Diversity 144:Latreille 138:Bembidion 117:Carabidae 73:Kingdom: 67:Eukaryota 25:Bembidion 1378:BugGuide 1354:BioLib: 1318:Q1757595 1312:Wikidata 1180:Linnaeus 1072:Trepanes 880:Peryphus 832:Pekinium 816:Pamirium 784:Odontium 728:Nothocys 720:Notaphus 608:Lymnaeum 456:Emphanes 448:Ecuadion 416:Cillenus 384:Bracteon 264:Actedium 235:List of 229:Taxonomy 167:Synonyms 113:Family: 83:Phylum: 77:Animalia 63:Domain: 1487:1024128 1461:1035920 1171:3: 82. 1016:Talanes 568:Josefia 536:Hydrium 239:species 133:Genus: 103:Order: 97:Insecta 93:Class: 1575:146896 1549:479960 1533:NZOR: 1500:109786 1474:130639 1435:384689 1422:1BEMDG 1210:  222:Tachys 146:, 1802 1570:WoRMS 1557:Plazi 1526:60800 1482:IRMNG 1409:15735 1396:8KVB5 1383:22818 1370:24735 904:Phyla 680:Nepha 209:genus 1521:NCBI 1495:ITIS 1456:GBIF 1417:EPPO 1365:BOLD 1357:3910 1208:ISBN 656:Neja 1508:NBN 1404:EoL 1391:CoL 1342:AFD 1594:: 1572:: 1559:: 1546:: 1523:: 1510:: 1497:: 1484:: 1471:: 1458:: 1445:: 1432:: 1419:: 1406:: 1393:: 1380:: 1367:: 1344:: 1329:: 1314:: 1244:. 1191:^ 1186:). 1175:: 1144:† 1135:† 1254:. 1238:"

Index


Bembidion quadrimaculatum
Scientific classification
Edit this classification
Eukaryota
Animalia
Arthropoda
Insecta
Coleoptera
Carabidae
Bembidiina
Bembidion
Latreille
Diversity
At least 1400 species
Synonyms

Bembidion tetracolum
genus
Carabidae
biantitropical distribution
Tachys
List of Bembidion species
René Jeannel
subgenera
Type species
Linnaeus


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