31:
75:
753:
50:
320:
grow to a snout-vent length of about 36 mm (1.4 in) and females to 43 mm (1.7 in). They are dark grey or brown above, with symmetrical blackish markings and smooth skin with small scattered warts on the head and back.
327:
is a reasonably common species associated with riparian vegetation, hill streams and leaf-litter in evergreen forest habitats. These frogs breed in streams. Tadpoles are 46 mm (1.8 in) in length.
394:
596:
635:
813:
794:
437:
712:
570:
725:
609:
549:
838:
479:
787:
730:
614:
544:
290:
828:
74:
30:
843:
833:
302:
640:
780:
535:
823:
286:
186:
348:
Michael Wai Neng Lau, Geng
Baorong, Gu Huiqing, Sushil Dutta, Mohini Mohan Borah, Sabitry Bordoloi (2004).
497:
818:
388:
169:
39:
372:
195:
69:
601:
415:"On a collection of reptiles and batrachians made by Mr. J. D. La Touche in N.W. Fokien, China"
699:
557:
475:
267:
663:
367:
306:
627:
648:
764:
298:
752:
807:
622:
358:
349:
59:
54:
285:
The history of this species' discovery is highly international. It was described by
760:
704:
562:
146:
136:
690:
520:
414:
583:
529:
254:
found in southern and southeastern China (the northern border runs roughly from
474:(in Chinese). Zhengzhou: Henan Press of Science and Technology. p. 116.
116:
86:
653:
301:, a German zoologist, based on specimens collected by Irish ornithologist
684:
514:
278:). A closely related but probably as yet undescribed species in found in
263:
106:
717:
575:
255:
588:
271:
259:
96:
491:
310:
279:
275:
251:
126:
495:
343:
341:
768:
446:
Amphibian
Species of the World 6.1, an Online Reference
674:
504:
289:, a Belgian zoologist who made his career in the
419:Proceedings of the Zoological Society of London
788:
8:
393:: CS1 maint: multiple names: authors list (
373:10.2305/IUCN.UK.2004.RLTS.T57632A11667261.en
408:
406:
404:
795:
781:
492:
48:
29:
20:
371:
337:
386:
266:in the east) and north-eastern India (
465:
463:
7:
749:
747:
448:. American Museum of Natural History
814:IUCN Red List least concern species
359:IUCN Red List of Threatened Species
14:
751:
73:
291:Natural History Museum, London
1:
413:Boulenger, George A. (1899).
839:Amphibians described in 1899
767:. You can help Knowledge by
472:Atlas of Amphibians of China
248:pale-shouldered horned toad
24:Boettger's horned toad
860:
746:
436:Frost, Darrel R. (2021).
244:Boettger's spadefoot toad
201:
194:
175:
168:
70:Scientific classification
68:
46:
37:
28:
23:
676:Leptobrachium boettgeri
287:George Albert Boulenger
205:Leptobrachium boettgeri
440:Boulenophrys boettgeri
325:Boulenophrys boettgeri
318:Boulenophrys boettgeri
295:Boulenophrys boettgeri
239:Boulenophrys boettgeri
234:Boettger's horned toad
179:Boulenophrys boettgeri
366:: e.T57632A11667261.
305:in Guadun village in
16:Species of amphibian
829:Amphibians of China
550:Megophrys-boettgeri
536:Panophrys boettgeri
506:Megophrys boettgeri
352:Megophrys boettgeri
221:Megophrys boettgeri
213:Xenophrys boettgeri
40:Conservation status
844:Megophryidae stubs
834:Fauna of Hong Kong
442:(Boulenger, 1899)"
250:, is a species of
776:
775:
741:
740:
498:Taxon identifiers
268:Arunachal Pradesh
262:in the north and
242:), also known as
231:
230:
225:
224:(Boulenger, 1899)
217:
216:(Boulenger, 1899)
209:
161:B. boettgeri
63:
851:
797:
790:
783:
755:
748:
734:
733:
721:
720:
708:
707:
695:
694:
693:
667:
666:
657:
656:
644:
643:
631:
630:
618:
617:
605:
604:
592:
591:
579:
578:
566:
565:
553:
552:
540:
539:
538:
525:
524:
523:
493:
486:
485:
470:Fei, L. (1999).
467:
458:
457:
455:
453:
433:
427:
426:
410:
399:
398:
392:
384:
382:
380:
375:
345:
307:Wuyishan, Fujian
223:
215:
207:
181:
78:
77:
57:
52:
51:
33:
21:
859:
858:
854:
853:
852:
850:
849:
848:
804:
803:
802:
801:
744:
742:
737:
729:
724:
716:
711:
703:
698:
689:
688:
683:
670:
662:
660:
652:
649:Observation.org
647:
639:
634:
626:
621:
613:
608:
600:
595:
587:
582:
574:
569:
561:
556:
548:
543:
534:
533:
528:
519:
518:
513:
500:
490:
489:
482:
469:
468:
461:
451:
449:
435:
434:
430:
412:
411:
402:
385:
378:
376:
347:
346:
339:
334:
303:J. D. La Touche
258:in the west to
208:Boulenger, 1899
190:
183:
177:
164:
72:
64:
53:
49:
42:
17:
12:
11:
5:
857:
855:
847:
846:
841:
836:
831:
826:
824:Frogs of India
821:
816:
806:
805:
800:
799:
792:
785:
777:
774:
773:
756:
739:
738:
736:
735:
722:
709:
696:
680:
678:
672:
671:
669:
668:
658:
645:
632:
619:
606:
593:
580:
567:
554:
541:
526:
510:
508:
502:
501:
496:
488:
487:
480:
459:
428:
400:
336:
335:
333:
330:
299:Oskar Boettger
229:
228:
227:
226:
218:
210:
199:
198:
192:
191:
184:
173:
172:
166:
165:
158:
156:
152:
151:
144:
140:
139:
134:
130:
129:
124:
120:
119:
114:
110:
109:
104:
100:
99:
94:
90:
89:
84:
80:
79:
66:
65:
47:
44:
43:
38:
35:
34:
26:
25:
15:
13:
10:
9:
6:
4:
3:
2:
856:
845:
842:
840:
837:
835:
832:
830:
827:
825:
822:
820:
817:
815:
812:
811:
809:
798:
793:
791:
786:
784:
779:
778:
772:
770:
766:
763:article is a
762:
757:
754:
750:
745:
732:
727:
723:
719:
714:
710:
706:
701:
697:
692:
686:
682:
681:
679:
677:
673:
665:
659:
655:
650:
646:
642:
637:
633:
629:
624:
620:
616:
611:
607:
603:
598:
594:
590:
585:
581:
577:
572:
568:
564:
559:
555:
551:
546:
542:
537:
531:
527:
522:
516:
512:
511:
509:
507:
503:
499:
494:
483:
481:7-5349-1835-9
477:
473:
466:
464:
460:
447:
443:
441:
432:
429:
424:
420:
416:
409:
407:
405:
401:
396:
390:
374:
369:
365:
361:
360:
355:
353:
344:
342:
338:
331:
329:
326:
322:
319:
314:
312:
308:
304:
300:
297:in honour of
296:
292:
288:
283:
281:
277:
273:
269:
265:
261:
257:
253:
249:
245:
241:
240:
235:
222:
219:
214:
211:
206:
203:
202:
200:
197:
193:
188:
182:
180:
174:
171:
170:Binomial name
167:
163:
162:
157:
154:
153:
150:
149:
145:
142:
141:
138:
135:
132:
131:
128:
125:
122:
121:
118:
115:
112:
111:
108:
105:
102:
101:
98:
95:
92:
91:
88:
85:
82:
81:
76:
71:
67:
61:
56:
55:Least Concern
45:
41:
36:
32:
27:
22:
19:
819:Boulenophrys
769:expanding it
761:Megophryidae
758:
743:
675:
505:
471:
450:. Retrieved
445:
439:
431:
422:
418:
389:cite journal
377:. Retrieved
363:
357:
351:
324:
323:
317:
315:
294:
284:
247:
243:
238:
237:
233:
232:
220:
212:
204:
178:
176:
160:
159:
148:Boulenophrys
147:
137:Megophryidae
18:
584:iNaturalist
530:Wikispecies
379:20 November
293:. He named
808:Categories
425:: 159–172.
332:References
691:Q28026112
521:Q28026107
187:Boulenger
155:Species:
93:Kingdom:
87:Eukaryota
685:Wikidata
602:10248569
515:Wikidata
452:2 August
264:Zhejiang
196:Synonyms
133:Family:
117:Amphibia
107:Chordata
103:Phylum:
97:Animalia
83:Domain:
60:IUCN 3.1
731:1103097
718:8765146
664:4804223
641:1249233
615:1103096
576:4817022
256:Sichuan
246:or the
189:, 1899)
143:Genus:
123:Order:
113:Class:
58: (
661:uBio:
654:957182
589:476423
478:
274:, and
272:Sikkim
260:Shanxi
759:This
705:3TKBF
628:57632
597:IRMNG
563:3Z29Z
316:Male
311:China
280:Tibet
276:Assam
127:Anura
765:stub
726:ITIS
713:GBIF
636:NCBI
623:IUCN
610:ITIS
571:GBIF
476:ISBN
454:2022
423:1899
395:link
381:2021
364:2004
252:toad
700:CoL
558:CoL
545:ASW
368:doi
810::
728::
715::
702::
687::
651::
638::
625::
612::
599::
586::
573::
560::
547::
532::
517::
462:^
444:.
421:.
417:.
403:^
391:}}
387:{{
362:.
356:.
340:^
313:.
309:,
282:.
270:,
796:e
789:t
782:v
771:.
484:.
456:.
438:"
397:)
383:.
370::
354:"
350:"
236:(
185:(
62:)
Text is available under the Creative Commons Attribution-ShareAlike License. Additional terms may apply.