51:
441:
112:
70:
285:, it was promoted to full species status in 2013 when a study of museum specimens and photographs identified distinct facial markings, which helped to differentiate it as a separate species. It is distinguished by its dark, contrasting facial features, as well as the shape and width of the stripes of its facial markings.
495:. The stripe between the eyes often varies in width, the ears are covered in hair, and the band of hair in front of the ears is wide. The colored patch on the top of the head is usually round, but is sometimes a narrower band. The body length averages 260.1 mm (10.24 in) for the species.
590:
to activate. Their toxic bite is a deterrent to predators, and the toxin is also applied to the fur during grooming as a form of protection for their infants. When threatened, slow lorises may also lick their brachial glands and bite their aggressors, delivering the toxin into the wounds. Slow
606:
was listed as "Vulnerable" as of 2012. Because that species has been divided into four distinct species, each of the new species faces a higher risk of extinction. Accordingly, each of them are expected to be listed as "Vulnerable" at the least, with some of them likely to be assigned to a
50:
1376:
Nekaris, K.A.I.; Bearder, S.K. (2007). "Chapter 3: The
Lorisiform Primates of Asia and Mainland Africa: Diversity Shrouded in Darkness". In Campbell, C.; Fuentes, C.A.; MacKinnon, K.; Panger, M.; Stumpf, R. (eds.).
483:
primates. On the ventral side of its elbow, it has a small swelling called the brachial gland, which secretes a pungent, clear oily toxin that the animal uses defensively by wiping it on its toothcomb.
432:
emerged as a new species, which had previously been overlooked. All newly recognized or elevated species showed significant differences in their "face mask"—the coloration patterns on their face.
989:
1135:
Chen, J. -H.; Pan, D.; Groves, C. P.; Wang, Y. -X.; Narushima, E.; Fitch-Snyder, H.; Crow, P.; Thanh, V. N.; Ryder, O.; Zhang, H. -W.; Fu, Y.; Zhang, Y. (2006). "Molecular phylogeny of
594:
The face mask may help the species identify potential mates by distinguishing species, and may serve as an anti-predator strategy by making its eyes appear larger than they really are.
1276:
Hagey, L.R.; Fry, B.G.; Fitch-Snyder, H. (2007). "Talking defensively, a dual use for the brachial gland exudate of slow and pygmy lorises". In Gursky, S.L.; Nekaris, K.A.I. (eds.).
1024:
2050:
570:, eating primarily insects, tree gum, nectar, and fruit. Likewise, this species has a toxic bite, a unique feature found only in slow lorises among primates. The
297:
2045:
475:
that it uses for scratching and grooming, while the other nails are straight. It also possesses a specialized arrangement of lower front teeth, called a
985:
2025:
296:
species primarily eats insects, tree gum, nectar, and fruit and has a toxic bite, a unique feature among primates. Although not yet evaluated by the
491:
are dark and contrasting. The dark rings around its eyes are usually rounded on top, though sometimes diffuse-edged, and they never reach below the
1919:
1945:
1487:
740:
467:, which distinguishes them from other slow lorises. On its front feet, the second digit is smaller than the rest; the big toe on its hind foot
1455:
1419:
1388:
1293:
1258:
1218:
1125:
1098:
610:
Between 1987 and 2012, one-third of Borneo's forests have been lost, making habitat loss one of the greatest threats to the survival of
31:
2040:
1398:
Nekaris, K.A.I.; Munds, R. (2010). "Chapter 22: Using facial markings to unmask diversity: the slow lorises (Primates: Lorisidae:
1971:
1012:
1250:
1950:
1329:
732:
1169:
1830:
701:
2035:
1480:
341:. Museum specimens of this animal had previously been identified as the Bornean slow loris using the scientific name
1232:
111:
1051:
1884:
707:. Convention on International Trade in Endangered Species of Wild Fauna and Flora (CITES). 2010. Archived from
402:
1846:
1473:
459:(the moist, naked surface around the nostrils of the nose) and a broad, flat face with large eyes. Like
2030:
1997:
1742:
1603:
686:
343:
281:
216:
440:
30:"Bornean slow loris" redirects here. For other slow lorises of Borneo that have shared that name, see
1906:
619:
359:
233:
591:
lorises can be reluctant to release their bite, which is likely to maximize the transfer of toxins.
1751:
559:
418:
363:
309:
305:
289:
59:
1778:
1683:
1621:
1612:
1570:
1364:
1156:
519:
301:
276:
106:
74:
2002:
471:
the other toes, which enhances its gripping power. Its second toe on the hind foot has a curved
1963:
1984:
1893:
1724:
1674:
1579:
1451:
1434:
1415:
1384:
1356:
1304:
1289:
1264:
1254:
1214:
1121:
1104:
1094:
1989:
1810:
1793:
1769:
1733:
1715:
1407:
1348:
1316:
1281:
1148:
681:
515:
445:
428:
382:
374:
351:
335:
1244:
659:
543:
511:
348:
1081:
Alterman, L. (1995). "Toxins and toothcombs: potential allospecific chemical defenses in
1543:
615:
492:
358:, – a scientific name now assigned exclusively to the Philippine slow loris. In 1906,
313:
2019:
1958:
1657:
1533:
1465:
1378:
1240:
668:
472:
253:
173:
79:
1368:
1320:
1160:
1898:
1280:. Developments in Primatology: Progress and Prospects. Springer. pp. 253–273.
1228:
1206:
563:
527:
523:
480:
452:
293:
634:
trade. However, all slow loris species are protected from commercial trade under
17:
1411:
1976:
1932:
1878:
708:
623:
1869:
1188:
1698:
1553:
1285:
1152:
631:
370:
264:
193:
1438:
1433:. Edinburgh Univ Pubs Science & Maths, No 3. Edinburgh University Press.
1497:
1268:
1108:
583:
575:
507:
476:
468:
456:
338:
272:
183:
123:
1360:
1863:
1515:
567:
535:
143:
1047:
1937:
1924:
1527:
627:
464:
260:
256:
163:
1352:
389:) as a species, and by further recognizing four subspecies, including
1521:
1509:
587:
268:
153:
133:
1840:
986:"Three new species of venomous primate identified by MU researcher"
1639:
1594:
639:
635:
579:
571:
439:
328:
96:
90:
1911:
1246:
Mammal
Species of the World: A Taxonomic and Geographic Reference
892:
890:
847:
845:
843:
841:
677:
534:
is not found in the extreme southwest of the island. It may be
412:
A 2013 review of museum specimens and photographs attributed to
1844:
1469:
1402:
spp.) of
Indonesia". In Gursky-Doyen, S.; Supriatna, J (eds.).
1383:. New York, New York: Oxford University Press. pp. 28–33.
416:
resulted in elevating two of its former subspecies to species:
1181:
877:
875:
401:. The latter was elevated to the species level in 2006, when
862:
860:
602:
While this new species has yet to be assessed by the IUCN,
618:
is also a major factor, with loris parts commonly sold in
463:, this and all other Bornean species lack a second upper
381:). In 1971, that view was updated by distinguishing the
1431:
Primates
Comparative Anatomy and Taxonomy I—Strepsirhini
327:
is a strepsirrhine primate, and species of slow loris (
1330:"Taxonomy of the Bornean slow loris, with new species
1328:
Munds, R. A.; Nekaris, K. A. I.; Ford, S. M. (2013) .
1093:. New York, New York: Plenum Press. pp. 413–424.
1089:". In Alterman, L.; Doyle, G.A.; Izard, M.K (eds.).
920:
1853:
1791:
1696:
1655:
1638:
1592:
1551:
1542:
687:
10.2305/IUCN.UK.2020-2.RLTS.T163015906A163015915.en
1213:. Washington, DC: Smithsonian Institution Press.
1048:"Primate species: new slow loris found in Borneo"
832:
479:, which is also used for grooming, as with other
1309:Proceedings of the United States National Museum
968:
932:
896:
851:
760:
373:were lumped together into a single species, the
1091:Creatures of the Dark: The Nocturnal Prosimians
304:" or placed in a higher-risk category when its
944:
298:International Union for Conservation of Nature
1481:
8:
1187:. Zürich, Switzerland: 44–53. Archived from
1064:
908:
866:
455:tail, round head, and short ears. It has a
308:is assessed. It is primarily threatened by
1841:
1652:
1548:
1488:
1474:
1466:
784:
405:showed it to be genetically distinct from
369:from western Borneo. By 1953, all of the
68:
49:
38:
979:
977:
685:
506:is found in central south Borneo, in the
2051:Fauna of the Borneo lowland rain forests
1406:. New York: Springer. pp. 383–396.
956:
881:
741:Integrated Taxonomic Information System
650:
275:. Formerly considered a subspecies or
1054:from the original on 25 December 2012.
820:
808:
796:
300:(IUCN), it is likely to be listed as "
1027:from the original on 17 December 2012
992:from the original on 17 December 2012
7:
1141:International Journal of Primatology
1139:inferred from mitochondrial genes".
772:
1446:Smith, Andrew T.; Xie, Yan (2008).
988:. Missouri University News Bureau.
669:IUCN Red List of Threatened Species
658:Nekaris, K.A.I.; Miard, P. (2020).
32:Bornean slow loris (disambiguation)
2046:Taxa named by Marcus Ward Lyon Jr.
921:Hagey, Fry & Fitch-Snyder 2007
522:. Its range extends south of the
451:Like other slow lorises, it has a
25:
347:– first described by the English
288:As with other slow lorises, this
2026:IUCN Red List vulnerable species
1278:Primate Anti-Predator Strategies
1120:(3rd ed.). Academic Press.
267:that is native to central south
110:
1448:A Guide to the Mammals of China
1341:American Journal of Primatology
1321:10.5479/si.00963801.31-1494.527
1046:Walker, M. (13 December 2012).
1011:Bryner, J. (14 December 2012).
1450:. Princeton University Press.
1251:Johns Hopkins University Press
969:Munds, Nekaris & Ford 2013
933:Munds, Nekaris & Ford 2013
897:Munds, Nekaris & Ford 2013
852:Munds, Nekaris & Ford 2013
761:Munds, Nekaris & Ford 2013
1:
984:Wall, T. (13 December 2012).
1412:10.1007/978-1-4419-1560-3_22
1243:; Reeder, D. M (eds.).
397:was considered a synonym of
2067:
1305:"Notes on the slow lemurs"
1170:"Systematics of the genus
945:Nekaris & Bearder 2007
702:"Appendices I, II and III"
680:: e.T163015906A163015915.
444:Female, from Nanga Tayap,
391:N. coucang menagensis
29:
2041:Mammals described in 1906
1827:
1504:
1496:Extant species of family
1429:Osman Hill, W.C. (1953).
1286:10.1007/978-0-387-34810-0
1168:Groves, Colin P. (1971).
1153:10.1007/s10764-006-9032-5
1116:Ankel-Simons, F. (2007).
582:by their elbow), and the
574:is produced by licking a
554:Like other slow lorises,
393:. From then until 2005,
222:
215:
107:Scientific classification
105:
88:
66:
57:
48:
41:
1770:Kayan River slow loris (
1065:Nekaris & Munds 2010
1743:Philippine slow loris (
1604:Central African potto (
1380:Primates in Perspective
1303:Lyon, M.W. Jr. (1906).
1017:, discovered in Borneo"
487:The facial markings of
1334:(Primates, Lorisidae)"
607:higher-risk category.
448:
320:Taxonomy and phylogeny
1998:Paleobiology Database
1779:Sumatran slow loris (
1235:Nycticebus menagensis
1013:"Slow loris species,
443:
344:Nycticebus menagensis
1885:Nycticebus borneanus
1855:Nycticebus borneanus
1761:Bornean slow loris (
1684:Gray slender loris (
1622:West African potto (
1613:East African potto (
1571:Calabar angwantibo (
1253:. pp. 111–184.
882:Smith & Xie 2008
735:Nycticebus borneanus
662:Nycticebus borneanus
620:traditional medicine
436:Physical description
360:Marcus Ward Lyon Jr.
245:Nycticebus borneanus
226:Nycticebus borneanus
43:Nycticebus borneanus
1752:Bangka slow loris (
1725:Bengal slow loris (
1675:Red slender loris (
1580:Golden angwantibo (
1404:Indonesian Primates
1021:The Huffington Post
959:, pp. 421–423.
884:, pp. 159–160.
787:, pp. 156–163.
775:, pp. 535–536.
550:Habitat and ecology
542:in the province of
306:conservation status
60:Conservation status
2036:Primates of Borneo
1811:Pygmy slow loris (
1801:(Pygmy slow loris)
1734:Javan slow loris (
1716:Sunda slow loris (
1241:Wilson, D. E.
1229:Groves, C. P.
604:N. menagensis
520:Central Kalimantan
461:N. menagensis
449:
414:N. menagensis
403:molecular analysis
399:N. menagensis
379:Nycticebus coucang
282:N. menagensis
250:Bornean slow loris
27:Species of primate
18:Bornean Slow Loris
2013:
2012:
1985:Open Tree of Life
1847:Taxon identifiers
1838:
1837:
1823:
1822:
1804:
1709:
1668:
1665:(Slender lorises)
1649:
1634:
1633:
1564:
1457:978-0-691-09984-2
1421:978-1-4419-1559-7
1390:978-0-19-517133-4
1353:10.1002/ajp.22071
1315:(1494): 527–538.
1295:978-0-387-34807-0
1260:978-0-8018-8221-0
1220:978-1-56098-872-4
1127:978-0-12-372576-9
1100:978-0-306-45183-6
947:, pp. 28–33.
909:Ankel-Simons 2007
867:Ankel-Simons 2007
612:N. borneanus
556:N. borneanus
532:N. borneanus
504:N. borneanus
489:N. borneanus
424:N. borneanus
395:N. borneanus
367:N. borneanus
325:N. borneanus
241:
240:
208:N. borneanus
100:
83:
16:(Redirected from
2058:
2006:
2005:
1993:
1992:
1980:
1979:
1967:
1966:
1954:
1953:
1941:
1940:
1928:
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1915:
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1902:
1901:
1889:
1888:
1887:
1874:
1873:
1872:
1842:
1803:
1802:
1798:
1794:Xanthonycticebus
1708:
1707:
1703:
1686:L. lydekkerianus
1667:
1666:
1662:
1653:
1648:
1647:
1643:
1563:
1562:
1558:
1549:
1490:
1483:
1476:
1467:
1461:
1442:
1425:
1394:
1372:
1338:
1332:Nycticebus kayan
1324:
1299:
1272:
1249:(3rd ed.).
1224:
1211:Primate Taxonomy
1207:Groves, Colin P.
1202:
1200:
1199:
1193:
1178:
1164:
1147:(4): 1187–1200.
1131:
1112:
1074:Literature cited
1068:
1062:
1056:
1055:
1043:
1037:
1036:
1034:
1032:
1015:Nycticebus kayan
1008:
1002:
1001:
999:
997:
981:
972:
966:
960:
954:
948:
942:
936:
935:, p. 52–53.
930:
924:
918:
912:
906:
900:
894:
885:
879:
870:
864:
855:
849:
836:
833:Chen et al. 2006
830:
824:
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806:
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794:
788:
782:
776:
770:
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751:
749:
747:
729:
723:
722:
720:
719:
713:
706:
698:
692:
691:
689:
655:
540:N. bancanus
526:and east to the
446:Ketapang Regency
426:. Additionally,
419:N. bancanus
383:pygmy slow loris
375:Sunda slow loris
356:Lemur menagensis
352:Richard Lydekker
312:and the illegal
228:
115:
114:
94:
77:
72:
71:
53:
39:
21:
2066:
2065:
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2059:
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2015:
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1996:
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1705:
1704:
1702:
1692:
1664:
1663:
1661:
1645:
1644:
1642:
1630:
1588:
1573:A. calabarensis
1560:
1559:
1557:
1538:
1500:
1494:
1464:
1458:
1445:
1428:
1422:
1397:
1391:
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1327:
1302:
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1275:
1261:
1227:
1221:
1205:
1197:
1195:
1191:
1176:
1167:
1134:
1128:
1118:Primate Anatomy
1115:
1101:
1080:
1076:
1071:
1063:
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1028:
1010:
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1005:
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927:
919:
915:
907:
903:
895:
888:
880:
873:
865:
858:
850:
839:
835:, p. 1198.
831:
827:
819:
815:
807:
803:
795:
791:
785:Osman Hill 1953
783:
779:
771:
767:
759:
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745:
743:
731:
730:
726:
717:
715:
711:
704:
700:
699:
695:
657:
656:
652:
648:
614:. The illegal
600:
586:mixes with its
552:
544:West Kalimantan
501:
438:
407:N. coucang
322:
237:
230:
224:
211:
109:
101:
84:
73:
69:
62:
35:
28:
23:
22:
15:
12:
11:
5:
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2054:
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2017:
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1807:
1805:
1789:
1788:
1786:
1785:
1776:
1767:
1758:
1749:
1740:
1731:
1727:N. bengalensis
1722:
1712:
1710:
1706:(Slow lorises)
1694:
1693:
1691:
1690:
1681:
1677:L. tardigradus
1671:
1669:
1650:
1636:
1635:
1632:
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1628:
1619:
1610:
1600:
1598:
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1565:
1546:
1544:Perodicticinae
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1132:
1126:
1113:
1099:
1077:
1075:
1072:
1070:
1069:
1067:, p. 390.
1057:
1038:
1003:
973:
961:
949:
937:
925:
923:, p. 253.
913:
911:, p. 246.
901:
886:
871:
856:
837:
825:
823:, p. 122.
813:
801:
789:
777:
765:
753:
724:
693:
649:
647:
644:
630:promoting the
616:wildlife trade
599:
596:
551:
548:
500:
497:
493:zygomatic arch
437:
434:
321:
318:
314:wildlife trade
239:
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45:
26:
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14:
13:
10:
9:
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4:
3:
2:
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1806:
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1745:N. menagensis
1741:
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1561:(Angwantibos)
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1534:Strepsirrhini
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853:
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473:grooming claw
470:
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429:N. kayan
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217:Binomial name
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2031:Slow lorises
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1812:
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1763:N. borneanus
1762:
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1736:N. javanicus
1735:
1726:
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1595:Perodicticus
1593:
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1430:
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1379:
1347:(1): 46–56.
1344:
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1210:
1196:. Retrieved
1189:the original
1184:
1180:
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1140:
1136:
1117:
1090:
1087:Perodicticus
1086:
1082:
1060:
1050:. BBC News.
1041:
1029:. Retrieved
1020:
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994:. Retrieved
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916:
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756:
744:. Retrieved
734:
727:
716:. Retrieved
709:the original
696:
673:
667:
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624:viral videos
611:
609:
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598:Conservation
593:
555:
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539:
531:
530:. However,
528:Barito River
524:Kapuas River
503:
502:
499:Distribution
488:
486:
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371:slow lorises
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310:habitat loss
287:
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249:
244:
243:
242:
225:
223:
207:
206:
194:
42:
36:
1933:iNaturalist
1879:Wikispecies
1813:X. pygmaeus
1754:N. bancanus
1606:P. edwardsi
1031:15 December
996:19 December
821:Groves 2005
809:Groves 2001
797:Groves 1971
387:N. pygmaeus
354:in 1893 as
2020:Categories
1781:N. hilleri
1718:N. coucang
1699:Nycticebus
1615:P. ibeanus
1554:Arctocebus
1532:Suborder:
1400:Nycticebus
1198:2013-01-04
1172:Nycticebus
1137:Nycticebus
1083:Nycticebus
746:28 January
718:2012-12-14
646:References
636:Appendix I
632:exotic pet
568:omnivorous
508:Indonesian
481:lemuriform
349:naturalist
332:Nycticebus
302:Vulnerable
265:slow loris
195:Nycticebus
170:Suborder:
93:Appendix I
75:Vulnerable
1964:163015906
1646:(Lorises)
1640:Lorisinae
1582:A. aureus
1508:Kingdom:
1498:Lorisidae
1439:500576914
773:Lyon 1906
584:secretion
578:gland (a
564:nocturnal
536:sympatric
477:toothcomb
457:rhinarium
453:vestigial
364:described
339:Lorisidae
294:nocturnal
273:Indonesia
202:Species:
184:Lorisidae
130:Kingdom:
124:Eukaryota
1912:42540721
1870:Q7071010
1864:Wikidata
1831:Category
1772:N. kayan
1624:P. potto
1528:Primates
1522:Mammalia
1516:Chordata
1514:Phylum:
1510:Animalia
1369:17077282
1361:23255350
1269:62265494
1231:(2005).
1209:(2001).
1161:24319996
1109:33441731
1052:Archived
1025:Archived
990:Archived
576:brachial
560:arboreal
290:arboreal
180:Family:
164:Primates
154:Mammalia
144:Chordata
140:Phylum:
134:Animalia
120:Domain:
80:IUCN 3.1
1990:6145320
1977:1001063
1938:1369277
1925:5786052
1526:Order:
1520:Class:
628:YouTube
469:opposes
465:incisor
277:synonym
261:species
257:primate
252:, is a
236:, 1906)
190:Genus:
160:Order:
150:Class:
95: (
78: (
2003:390434
1951:944111
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588:saliva
566:, and
518:, and
362:first
336:family
269:Borneo
259:and a
248:, the
1899:485JJ
1658:Loris
1365:S2CID
1337:(PDF)
1239:. In
1192:(PDF)
1177:(PDF)
1157:S2CID
712:(PDF)
705:(PDF)
640:CITES
580:gland
572:toxin
538:with
516:South
329:genus
97:CITES
91:CITES
1959:IUCN
1946:ITIS
1920:GBIF
1452:ISBN
1435:OCLC
1416:ISBN
1385:ISBN
1357:PMID
1290:ISBN
1265:OCLC
1255:ISBN
1215:ISBN
1122:ISBN
1105:OCLC
1095:ISBN
1085:and
1033:2012
998:2012
748:2016
678:IUCN
674:2020
622:and
512:West
422:and
292:and
234:Lyon
1972:MDD
1907:EoL
1894:CoL
1408:doi
1349:doi
1317:doi
1282:doi
1149:doi
682:doi
638:of
626:on
558:is
279:of
271:in
263:of
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