483:, with no preference for sleeping at day or night. Large, curved claws and muscles specifically adapted for strength and stamina help them to keep a strong grip on tree branches. They are able to withstand hanging inverted for extended periods of time due to fibrinous adhesions which attach their organs (such as their liver and stomach) to their lower ribs. Given that feces and urine can account for up to a third of their body weight, this adaptation prevents these organs from pressing on the lungs when hanging upside down, making breathing easier. Adult animals are solitary, except when raising young, and males have been observed to fight one another using their fore claws.
492:), as well as flowers and fruits. They do not travel far, with home ranges of only around 0.5 to 9 ha (1.2 to 22.2 acres), depending on the local environment. Within a typical, 5-hectare (12-acre) range, a brown-throated sloth will visit around 40 trees, and may specialise on one particular species, even spending up to 20% of its time in a single specific tree. Thus, although the species are generalists, individual sloths may feed on a relatively narrow range of leaf types. Although they get most of their fluids from the leaves that they eat, brown-throated sloths have been observed drinking directly from rivers.
380:
368:
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chemical communication between individuals; 4) pick up trace nutrients in their claws, that are then ingested; and 5) favor a mutualistic relationship with populations of fur moths. More recently, a new hypothesis has emerged, which presents evidence against the previous ones and proposes that all current sloths are descendants from species that defecated on the ground, and there simply has not been enough selective pressure to abandon this behavior, since cases of predation during defecation are actually very rare.
979:
496:
103:
460:
61:
1729:
350:. The algae are generally absent in the hair of young sloths, and may also be absent in particularly old individuals, where the outer cuticle of the hair has been lost. Sloth hair also harbours a rich fungal flora. Certain strains of fungi that grow on brown-throated sloth fur have been shown to possess anti-parasitic, anti-cancer, and anti-bacterial qualities.
42:
545:
mating system. Studies of the brown-throated sloth indicate that mating is most common between
January and March in at least the northern parts of its range, but this may vary elsewhere. Gestation lasts at least seven months, and the single young is born fully furred and clawed. Young sloths cling to
478:
Brown-throated sloths sleep 15 to 18 hours every day and are active for only brief periods of time, which may be during either the day or night. Although they can walk along the ground, and even swim, they spend most of their lives in the high branches of trees, descending once every eight days or so
553:
of the females do not store significant quantities of milk as most other mammals do, since the infant sloth remains attached to the nipple at all times, and consumes the milk as soon as it is generated. The young begin to take solid food as early as four days after birth, initially licking particles
507:
Sloths descend about once every eight days to defecate on the ground. The reason and mechanism behind this behavior have long been debated among scientists. There are at least five hypotheses: 1) fertilize trees when feces are deposited at the base of the tree; 2) cover feces and avoid predation; 3)
293:
The brown-throated sloth is of similar size and build to most other species of three-toed sloths, with both males and females being 42 to 80 cm (17 to 31 in) in total body length. The tail is relatively short, only 2.5 to 9 cm (1.0 to 3.5 in) long. Adults weigh from 2.25 to
239:
294:
6.3 kg (5.0 to 13.9 lb), with no significant size difference between males and females. Each foot has three fingers, ending in long, curved claws, which are 7 to 8 cm (2.8 to 3.1 in) long on the fore feet, and 5 to 5.5 cm (2.0 to 2.2 in) on the hind feet.
532:
The female of the species is known to emit a loud, shrill scream during the mating season to attract males. Its cry sounds like "ay ay", much like that of a woman screaming. The male can be identified by a black stripe surrounded by orange fur on its back between the shoulders.
1073:
Castro-Sa, M. J., Dias-Silva, R. H. P., & Barnett, A. A. (2021). Where to go when all options are terrible: ranging behavior of brown-throated three-toed sloths (Bradypus variegatus) in central
Amazonian flooded igapó forests. Canadian Journal of Zoology, 99(9), 823–831.
450:
It is found in many different kinds of environments, including evergreen and dry forests and in highly perturbed natural areas. It is generally found from sea level to 1,200 m (3,900 ft), although some individuals have been reported from much higher elevations.
312:
The brown-throated sloth has grayish-brown to beige-color fur over the body, with darker brown fur on the throat, the sides of the face, and the forehead. The face is generally paler in color, with a stripe of very dark fur running beneath the eyes.
670:, which has a very similar appearance, except for the color of the fur around the throat. The two species are estimated to have diverged just 400,000 years ago, whereas their ancestors diverged from the
297:
The head is rounded, with a blunt nose and inconspicuous ears. As with other sloths, the brown-throated sloth has no incisor or canine teeth, and the cheek teeth are simple and peg-like. They have no
554:
of food from their mother's mouths. This process apparently allows them to quickly identify edible leaves, and young sloths typically have the same preferences for leaf types as their mothers.
1673:
357:. Where this overlap occurs, the three-toed sloth tends to be smaller and more numerous than its relative, being more active in moving through the forest and maintaining more
1592:
2269:
1310:
Pauli, J. N., Mendoza, J. E., Steffan, S. A., Carey, C. C., Weimer, P. J., & Peery, M. Z. (2014). A syndrome of mutualism reinforces the lifestyle of a sloth.
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Brown-throated sloths inhabit the high canopy of the forest, where they eat young leaves from a wide range of different trees (primarily in the genus
2243:
1670:
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1343:
Voirin, B., Kays, R., Wikelski, M., & Lowman, M. (2013). Why Do Sloths Poop on the Ground? In M. Lowman, S. Devy, & T. Ganesh (eds).
854:
820:
709:
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has been observed to forage for small invertebrates in the fur of the sloths, apparently without the sloth being disturbed by the attention.
1633:(1925). "Verzeichnis der saugetier-typen des Senckenbergischen Museums" [Directory of the mammal types of the Senckenberg Museum].
379:
2439:
2469:
603:
1617:
The Animal
Kingdom Divided according to the Building of the Animals as the Basis of their Natural History and the Comparative Anatomy
992:
Higginbotham, Sarah; Wong, Weng Ruh; Linington, Roger G.; Spadafora, Carmenza; Iturrado, Liliana; Arnold, A. Elizabeth (2014-01-15).
2308:
1761:
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2326:
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2454:
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Montgomery, G. G., & Sunquist, M. E. (1975). Impact of Sloths on
Neotropical Forest Energy Flow and Nutrient Cycling.
320:
are very coarse and stiff, and overlie a much softer layer of dense under-fur. The hairs are unusual in lacking a central
285:
It is the most common of the four species of three-toed sloth, and is found in the forests of South and
Central America.
2459:
2434:
1944:
1935:
500:
354:
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1612:
Das
Thierreich Eingetheilt nach dem Bau der Thiere als Grundlage ihrer Naturgeschichte und der Vergleichenden Anatomie
846:
584:
The seven recognized subspecies of the brown-throated sloth, although these are not all readily distinguishable, are:
691:
2424:
815:. Vol. 1: Marsupials, Xenarthrans, Shrews, and Bats. Chicago: University of Chicago Press. pp. 161–164.
2339:
1430:
1091:; Butcher, Michael T. (2018-12-01). "Architectural Properties of Sloth Forelimb Muscles (Pilosa: Bradypodidae)".
546:
the mother's underside for five months or more, even though they are fully weaned after just four to five weeks.
102:
2449:
614:
994:"Sloth Hair as a Novel Source of Fungi with Potent Anti-Parasitic, Anti-Cancer and Anti-Bacterial Bioactivity"
928:"Sloth hair as a novel source of fungi with potent anti-parasitic, anti-cancer and anti-bacterial bioactivity"
346:
2159:
1733:
340:
570:
978:
511:
In addition to the algae in their fur, brown-throated sloths also live commensally with a species of moth,
2121:
1863:
1606:
658:
566:
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In the wild, the lifespan of adult brown-throated three-toed sloths is typically between 30 and 40 years.
526:
213:
2196:
2373:
1586:
468:
197:
334:
395:
The brown-throated sloth is the most widespread and common of the three-toed sloths. It is found from
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1005:
939:
1881:
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324:, and have numerous microscopic cracks across their surfaces. These cracks are host to a number of
50:
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238:
1654:"A new species of three-toed sloth (Mammalia: Xenarthra) from Panamá, with a review of the genus
1411:
1116:
513:
440:
306:
97:
2378:
1356:
Monge Nájera, J. (2021). Why sloths defecate on the ground: rejection of the mutualistic model.
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436:
271:
1140:; Avey-Arroyo, Judy A.; Arroyo, Francisco J.; Holton, Mark D.; Wilson, Rory P. (2014-04-30).
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267:
174:
164:
2419:
2347:
2300:
2021:
1747:
1677:
1522:
Taube, E.; et al. (2001). "Reproductive biology and postnatal development in sloths,
647:
625:
275:
1290:
Liberg, O. (1980). Spacing patterns in a population of rural free roaming domestic cats.
1570:
1472:
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943:
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2003:
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1415:
1120:
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1984:
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17:
2209:
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926:
Higginbotham S, Wong WR, Linington RG, Spadafora C, Iturrado L, Arnold AE (2014).
1481:
1399:
1018:
952:
840:
2391:
2313:
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2153:
1872:
1530:: review with original data from the field (French Guiana) and from captivity".
671:
522:
1728:
1104:
472:
404:
358:
317:
2144:
1691:"Phylogenetic analysis of 16S mitochondrial DNA data in sloths and anteaters"
1490:
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1808:
864:
719:
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114:
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1972:
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134:
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Over parts of its range, the brown-throated sloth overlaps the range of
2248:
424:
420:
263:
1431:"Foraging of yellow-headed caracaras in the fur of a three-toed sloth"
1327:
Chiarello, A. G. (2008). Sloth ecology: an overview of field studies.
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are among the few natural predators of the brown-throated sloth. The
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408:
154:
144:
124:
2115:
1619:] (in German). Stuttgart and Tubingen: J. G. Cotta. p. 510.
41:
2235:
1457:"Unexpected Strong Polygyny in the Brown-Throated Three-Toed Sloth"
2386:
1824:
1273:
Bailey, T. N. (1974). Social organization in a bobcat population.
911:
894:
704:(3rd ed.). Johns Hopkins University Press. pp. 100–101.
574:
494:
458:
329:
302:
87:
81:
1142:"Mitigating the squash effect: sloths breathe easily upside down"
701:
Mammal
Species of the World: A Taxonomic and Geographic Reference
432:
321:
27:
Species of New World mammals related to anteaters and armadillos
2222:
2119:
1743:
447:
found in these regions has led to some confusion in the past.
517:, which lives in their fur, and lays its eggs in the dung.
1312:
Proceedings of the Royal
Society B: Biological Sciences
1190:"Bradypus variegatus (Brown-throated three-toed sloth)"
479:
to defecate in the soil. The brown-throated sloths are
1455:
Pauli, Jonathan N.; Peery, M. Zachariah (2012-12-19).
733:
Moraes-Barros, N.; Chiarello, A.; Plese, T. (2014).
435:. It is probably not found immediately north of the
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2053:
2029:
2020:
1992:
1983:
1916:
1907:
1844:
1835:
1662:
Proceedings of the
Biological Society of Washington
1132:
1130:
666:The closest living relative of the species is the
565:The brown-throated sloth was first described by
1652:Anderson, R.P. & Handley, C.O. Jr. (2001).
1059:Dickman, Chris (1993). Macdonald, David (ed.).
1378:: a coprophagous moth that lives on a sloth".
1755:
1591:: CS1 maint: DOI inactive as of March 2024 (
1318:(1778), 20133006. DOI: 10.1098/rspb.2013.3006
888:
886:
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874:
842:Neotropical rainforest mammals. A field guide
758:10.2305/IUCN.UK.2014-1.RLTS.T3038A47437046.en
8:
1557:Mirsky, Steve (2009). "The Bradypus Bunch".
1429:Krakauer, A.H. & Krakauer, T.H. (1999).
1374:Waage, J.K. & Montgomery, G.G. (1976). "
569:in 1825 from a South American specimen. The
2116:
2026:
1989:
1966:
1913:
1841:
1818:
1762:
1748:
1740:
373:Male showing black patch between shoulders
237:
59:
40:
31:
1709:
1498:
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1035:
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1264:, 69–98. DOI:10.1007/978-3-642-88533-4_7
1218:Frontiers in Ecology and the Environment
682:
363:
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1347:(pp. 195-199). Springer, New York, NY.
1339:
1337:
1306:
1304:
1087:Olson, Rachel A.; Glenn, Zachary D.;
1076:https://doi.org/10.1139/cjz-2020-0176
7:
2475:Taxa named by Heinrich Rudolf Schinz
2415:IUCN Red List least concern species
744:IUCN Red List of Threatened Species
1689:Barros, M.C.; et al. (2003).
1275:The Journal of Wildlife Management
839:; Feer, François (February 1997).
499:Brown-throated sloth at midday in
25:
1579:10.1038/scientificamerican0509-34
573:was later specified as "probably
443:, although its similarity to the
1727:
1544:10.1111/j.1365-2907.2001.00085.x
977:
378:
366:
101:
1711:10.1590/S1415-47572003000100002
1698:Genetics and Molecular Biology
1093:Journal of Mammalian Evolution
463:Feeding brown-throated sloth (
1:
541:Brown-throated sloths have a
229:
1482:10.1371/journal.pone.0051389
1400:10.1126/science.193.4248.157
1329:The biology of the Xenarthra
1019:10.1371/journal.pone.0084549
953:10.1371/journal.pone.0084549
501:Manuel Antonio National Park
1945:Hoffmann's two-toed sloth (
1936:Linnaeus's two-toed sloth (
1061:The Encyclopedia of Mammals
847:University of Chicago Press
811:. In Gardner, A. L. (ed.).
246:Brown-throated sloth range
2491:
2440:Mammals of Central America
1438:Journal of Raptor Research
674:over 7 million years ago.
2470:Mammals described in 1825
2102:
1969:
1965:
1821:
1817:
1779:
1214:"Hanging out for a drink"
1212:; Haupt, Ryan J. (2018).
1105:10.1007/s10914-017-9411-z
845:(2nd ed.). Chicago:
355:Hoffmann's two-toed sloth
245:
236:
226:
221:
203:
196:
98:Scientific classification
96:
79:
57:
48:
39:
34:
1864:Pygmy three-toed sloth (
813:Mammals of South America
698:; Reeder, D. M. (eds.).
391:Distribution and habitat
347:Chlorococcum choloepodis
899:(Pilosa: Bradypodidae)"
803:Gardner, A. L. (2007).
690:Gardner, A. L. (2005).
341:Dictyococcus bradypodis
1891:Brown-throated sloth (
1581:(inactive 2024-03-08).
1158:10.1098/rsbl.2014.0172
567:Heinrich Rudolf Schinz
527:yellow-headed caracara
504:
475:
399:in the north, through
2465:Fauna of the Caatinga
2374:Paleobiology Database
1882:Pale-throated sloth (
1358:UNED Research Journal
498:
469:Cahuita National Park
462:
35:Brown-throated sloth
2455:Mammals of Venezuela
1195:Animal Diversity Web
893:Hayssen, V. (2010).
780:"Appendices | CITES"
751:: e.T3038A47437046.
254:brown-throated sloth
2460:Fauna of the Amazon
2435:Mammals of Colombia
2174:Bradypus_variegatus
2160:Bradypus variegatus
2130:Bradypus variegatus
2082:Southern tamandua (
2073:Northern tamandua (
1854:(Three-toed sloths)
1774:species by suborder
1734:Bradypus variegatus
1571:2009SciAm.300e..34M
1559:Scientific American
1473:2012PLoSO...751389P
1392:1976Sci...193..157W
1230:2018FrEE...16..446C
1010:2014PLoSO...984549H
944:2014PLoSO...984549H
897:Bradypus variegatus
807:Bradypus variegatus
737:Bradypus variegatus
668:pale-throated sloth
465:Bradypus variegatus
445:pale-throated sloth
259:Bradypus variegatus
207:Bradypus variegatus
51:Conservation status
18:Bradypus variegatus
2445:Mammals of Ecuador
2430:Mammals of Bolivia
1676:2010-06-13 at the
1376:Cryptoses choloepi
1262:Ecological Studies
1210:Cliffe, Rebecca N.
1138:Cliffe, Rebecca N.
1089:Cliffe, Rebecca N.
600:B. v. brasiliensis
514:Cryptoses choloepi
505:
476:
455:Behaviour and diet
189:B. variegatus
2425:Mammals of Brazil
2402:
2401:
2361:Open Tree of Life
2122:Taxon identifiers
2113:
2112:
2098:
2097:
2094:
2093:
2066:
2016:
2015:
1976:
1961:
1960:
1957:
1956:
1929:
1926:(Two-toed sloths)
1903:
1902:
1857:
1828:
1386:(4248): 157–158.
903:Mammalian Species
856:978-0-226-20721-6
837:Emmons, Louise H.
822:978-0-226-28240-4
711:978-0-8018-8221-0
662:
651:
644:B. v. trivittatus
640:
629:
618:
607:
596:
589:B. v. boliviensis
437:Amazon Rainforest
335:Rufusia pillicola
272:Neotropical realm
250:
249:
91:
74:
16:(Redirected from
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2041:Giant anteater (
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2004:Silky anteater (
1990:
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1914:
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1732:Data related to
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1538:(3–4): 173–188.
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1345:Treetops at Risk
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730:
724:
723:
687:
657:
655:B. v. variegatus
646:
635:
633:B. v. infuscatus
624:
613:
611:B. v. ephippiger
602:
591:
382:
370:
268:three-toed sloth
241:
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106:
105:
85:
68:
63:
62:
44:
32:
21:
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2348:Observation.org
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2084:T. tetradactyla
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2022:Myrmecophagidae
2012:
1979:
1953:
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1678:Wayback Machine
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1268:
1259:
1255:
1208:
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1203:
1188:
1187:
1183:
1152:(4): 20140172.
1146:Biology Letters
1136:
1135:
1128:
1086:
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1081:
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1058:
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1053:
991:
990:
986:
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835:
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823:
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797:
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773:
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761:
732:
731:
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712:
689:
688:
684:
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563:
539:
457:
439:or east of the
393:
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15:
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5:
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2110:
2103:
2100:
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2037:
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2024:
2018:
2017:
2014:
2013:
2011:
2010:
2000:
1998:
1987:
1981:
1980:
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1963:
1962:
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1955:
1954:
1952:
1951:
1942:
1932:
1930:
1911:
1905:
1904:
1901:
1900:
1898:
1897:
1888:
1884:B. tridactylus
1879:
1870:
1860:
1858:
1839:
1833:
1832:
1822:
1815:
1814:
1812:
1811:
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1769:
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1766:
1759:
1752:
1744:
1738:
1737:
1736:at Wikispecies
1723:
1720:
1718:
1717:
1681:
1644:
1622:
1598:
1549:
1514:
1467:(12): e51389.
1447:
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1366:
1349:
1333:
1320:
1300:
1283:
1266:
1253:
1201:
1181:
1126:
1099:(4): 573–588.
1079:
1066:
1063:. p. 777.
1051:
984:
918:
870:
855:
849:. p. 43.
828:
821:
795:
771:
725:
710:
692:"Order Pilosa"
681:
679:
676:
664:
663:
652:
641:
630:
619:
608:
597:
579:Robert Mertens
562:
559:
551:mammary glands
538:
535:
456:
453:
392:
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377:
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365:
290:
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248:
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2071:
2070:
2068:
2058:
2057:
2052:
2046:
2044:
2043:M. tridactyla
2039:
2038:
2036:
2034:
2033:
2028:
2025:
2023:
2019:
2009:
2007:
2006:C. didactylus
2002:
2001:
1999:
1997:
1996:
1991:
1988:
1986:
1982:
1978:
1977:
1968:
1964:
1950:
1948:
1943:
1941:
1939:
1938:C. didactylus
1934:
1933:
1931:
1921:
1920:
1915:
1912:
1910:
1909:Choloepodidae
1906:
1896:
1894:
1893:B. variegatus
1889:
1887:
1885:
1880:
1878:
1876:
1873:Maned sloth (
1871:
1869:
1867:
1862:
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1648:
1645:
1640:
1637:(in German).
1636:
1632:
1626:
1623:
1618:
1614:
1613:
1608:
1607:Schinz, H. R.
1602:
1599:
1594:
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1580:
1576:
1572:
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1564:
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1532:Mammal Review
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937:
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912:10.1644/850.1
908:
905:(42): 19–32.
904:
900:
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889:
887:
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883:
881:
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877:
875:
871:
866:
862:
858:
852:
848:
844:
843:
838:
832:
829:
824:
818:
814:
810:
809:Schinz, 1825"
808:
799:
796:
785:
781:
775:
772:
759:
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750:
746:
745:
740:
738:
729:
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721:
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707:
703:
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697:
696:Wilson, D. E.
693:
686:
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669:
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616:
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609:
605:
601:
598:
594:
590:
587:
586:
585:
582:
580:
577:(Brazil)" by
576:
572:
571:type locality
568:
560:
558:
555:
552:
547:
544:
536:
534:
530:
528:
524:
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516:
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286:
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280:South America
277:
273:
270:found in the
269:
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261:
260:
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244:
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198:Binomial name
195:
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133:
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120:
119:
116:
113:
110:
109:
104:
99:
95:
89:
83:
78:
72:
67:
66:Least Concern
56:
52:
47:
43:
38:
33:
30:
19:
2129:
2104:
2083:
2074:
2054:
2042:
2032:Myrmecophaga
2030:
2005:
1993:
1985:Cyclopedidae
1973:Vermilingua
1971:
1947:C. hoffmanni
1946:
1937:
1917:
1892:
1890:
1883:
1875:B. torquatus
1874:
1865:
1845:
1837:Bradypodidae
1823:
1701:
1697:
1684:
1665:
1661:
1655:
1647:
1638:
1634:
1625:
1616:
1611:
1601:
1587:cite journal
1562:
1558:
1552:
1535:
1531:
1527:
1523:
1517:
1464:
1460:
1450:
1441:
1437:
1424:
1383:
1379:
1375:
1369:
1361:
1357:
1352:
1344:
1328:
1323:
1315:
1311:
1298:(3),336-349.
1295:
1291:
1286:
1281:(3),435-446.
1278:
1274:
1269:
1261:
1256:
1221:
1217:
1204:
1193:
1184:
1149:
1145:
1096:
1092:
1082:
1069:
1060:
1054:
1001:
997:
987:
935:
931:
921:
902:
896:
841:
831:
812:
806:
798:
787:. Retrieved
783:
774:
762:. Retrieved
748:
742:
736:
728:
700:
685:
665:
654:
643:
632:
622:B. v. gorgon
621:
610:
599:
588:
583:
564:
556:
548:
540:
537:Reproduction
531:
523:harpy eagles
512:
510:
506:
503:, Costa Rica
487:
485:
477:
464:
449:
431:and eastern
394:
352:
345:
339:
333:
332:, including
315:
311:
299:gall bladder
296:
292:
284:
258:
257:
253:
251:
227:
206:
204:
188:
187:
175:
165:Bradypodidae
29:
2257:iNaturalist
2154:Wikispecies
2075:T. mexicana
2063:(Tamanduas)
1975:(Anteaters)
1866:B. pygmaeus
1807:Superorder
1801:Infraclass
1704:(1): 5–11.
1631:Mertens, R.
764:19 November
672:maned sloth
328:species of
318:guard hairs
289:Description
222:Subspecies
84:Appendix II
2409:Categories
1668:(1): 1–33.
1331:, 269-280.
1224:(8): 446.
789:2022-01-14
678:References
604:Blainville
543:polygynous
481:cathemeral
473:Costa Rica
405:Costa Rica
361:activity.
1919:Choloepus
1825:Folivora
1809:Xenarthra
1565:(5): 34.
1528:Choloepus
1491:1932-6203
1444:(3): 270.
1364:(1), 4-4.
1248:1540-9309
1113:1573-7055
1028:1932-6203
784:cites.org
581:in 1925.
441:Rio Negro
413:Venezuela
401:Nicaragua
326:commensal
183:Species:
121:Kingdom:
115:Eukaryota
2387:Species+
2327:11800008
2275:10228963
2181:BioLib:
2139:Wikidata
2106:Category
2056:Tamandua
1995:Cyclopes
1847:Bradypus
1827:(Sloths)
1803:Eutheria
1797:Mammalia
1791:Chordata
1785:Animalia
1783:Kingdom
1674:Archived
1671:PDF copy
1656:Bradypus
1641:: 18–37.
1609:(1825).
1524:Bradypus
1509:23284687
1461:PLOS ONE
1416:32766454
1408:17759254
1176:24759371
1121:22238582
1046:24454729
998:PLOS ONE
972:24454729
932:PLOS ONE
865:44179508
720:62265494
648:Cornalia
615:Philippi
561:Taxonomy
489:Cecropia
417:Colombia
397:Honduras
307:appendix
262:) is a
176:Bradypus
161:Family:
145:Mammalia
135:Chordata
131:Phylum:
125:Animalia
111:Domain:
71:IUCN 3.1
2314:1000432
2249:2436353
2145:Q148752
1789:Phylum
1770:Extant
1567:Bibcode
1500:3526605
1469:Bibcode
1388:Bibcode
1380:Science
1226:Bibcode
1167:4013704
1037:3893167
1006:Bibcode
963:3893167
940:Bibcode
519:Jaguars
425:Bolivia
421:Ecuador
359:diurnal
322:medulla
276:Central
264:species
228:7, see
171:Genus:
151:Order:
141:Class:
86: (
69: (
2420:Sloths
2379:426010
2366:273233
2288:624888
2236:BRDPVA
2223:328518
2197:642350
1795:Class
1772:Pilosa
1507:
1497:
1489:
1414:
1406:
1246:
1174:
1164:
1119:
1111:
1044:
1034:
1026:
970:
960:
863:
853:
819:
718:
708:
661:, 1825
659:Schinz
650:, 1849
639:, 1831
637:Wagler
628:, 1926
626:Thomas
617:, 1870
606:, 1840
595:, 1871
429:Brazil
409:Panama
385:female
344:, and
216:, 1825
214:Schinz
155:Pilosa
2353:82102
2270:IRMNG
2262:47067
2184:31890
1694:(PDF)
1615:[
1434:(PDF)
1412:S2CID
1292:Oikos
1117:S2CID
694:. In
575:Bahia
411:into
330:algae
305:, or
303:cecum
88:CITES
82:CITES
2392:5257
2340:9355
2335:NCBI
2301:3038
2296:IUCN
2283:ITIS
2244:GBIF
2231:EPPO
2210:N5F8
2192:BOLD
1593:link
1526:and
1505:PMID
1487:ISSN
1404:PMID
1244:ISSN
1172:PMID
1109:ISSN
1042:PMID
1024:ISSN
968:PMID
861:OCLC
851:ISBN
817:ISBN
766:2021
749:2014
716:OCLC
706:ISBN
593:Gray
549:The
521:and
433:Peru
407:and
316:The
278:and
252:The
230:text
2322:MSW
2309:MDD
2218:EoL
2205:CoL
2169:ADW
1706:doi
1666:114
1575:doi
1563:300
1540:doi
1495:PMC
1477:doi
1396:doi
1384:193
1316:281
1234:doi
1162:PMC
1154:doi
1101:doi
1032:PMC
1014:doi
958:PMC
948:doi
907:doi
753:doi
467:),
274:of
266:of
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