44:
235:
1374:
987:(id)s. The mesoloph is an accessory crest on the upper molars and the mesolophid is the corresponding structure on the lower molars. Only a few other animals now considered oryzomyines lack complete mesoloph(id)s, but they are absent in various non-oryzomyines, some of which had previously been regarded as close relatives of the oryzomyines that lack them. Oryzomyines with and without complete mesoloph(id)s share various other characters, however, including presence of
1616:
1177:
96:
71:
1152:; the color changes gradually over the body. The small ears are covered with short hairs. The tail is as long as or slightly longer than the head and body, and is dark above and light below. Despite the presence of short hairs, the scales on the tail are clearly visible. The hairs on the feet are pale. The hindfeet are long and narrow and have five toes, the first and fifth of which are short.
1396:, with the cusps extending higher than the central parts of the molars. They are characterized by strong cusps and absence or reduction of accessory crests. The cusps of the upper molars are opposite, but in the lower molars the labial (outer) cusps are slightly further to the front than the lingual (inner) ones. On the upper first molar, one accessory ridge, the
1172:
are reduced, other common characteristics of semiaquatic oryzomyines. The head-body length is 94 to 140 millimeters (3.7 to 5.5 in), tail length 102 to 140 mm (4.0 to 5.5 in), hindfeet length 27 to 33 mm (1.1 to 1.3 in), ear length 13 to 19 mm (0.5 to 0.7 in) and
1450:
and some other species has another labial root. The first lower molar has large roots at the front and back of the tooth and two smaller ones in between, at the labial and lingual side. The second and third lowers molars have two roots at the front, one labial and one lingual, and another at the
1751:. Most specimens for which habitat data are known were caught on the ground in humid grassland, some in seasonally flooded areas; an Argentinean specimen was captured in dense swamp vegetation. It is terrestrial and semiaquatic, living on the ground but also spending time in the water.
1693:
gene, but nothing is known about genetic variation in other parts of the range. The species has long been rare in collections; in 1991, Voss and Myers could use less than 50 specimens for their study of the species, including Lund's fragmentary material from Lagoa Santa.
1103:; among all oryzomyines, this was the only case where relationships which received strong support from morphological and DNA sequence data conflicted. Together, the three genera form part of a large group of oryzomyines ("
1674:. Paraguayan animals are somewhat smaller than those from Bolivia and Brazil and those from Bolivia have darker fur than Paraguayan specimens, but these differences are not considered significant enough to recognize
1802:. It is a widely distributed species without substantial threats to its continued existence, but degradation of its habitat may endanger some populations. It was assessed as "potentially vulnerable" in Argentina.
1542:
with a total of 54 major arms (2n = 56, FN = 54) in specimens from both
Bolivia and Brazil; a poorly prepared Paraguayan specimen seems to have a similar karyotype. In this karyotype, all
635:
are in fact the same species. In a 1991 study, American zoologists Voss and Myers confirmed this suggestion after re-examining Winge's material, finding no appreciable differences among specimens of
1740:
346:
2464:
352:. It is a medium-sized species, weighing about 50 grams (1.8 oz), with gray–brown fur, long and narrow hindfeet, and a tail that is about as long as the head and body. The
2485:
Díaz, G. and Ojeda, R. 2000. Libro Rojo de los mamíferos amenazados de la
Argentina. Mendoza: Sociedad Argentina para el Estudio de los Mamíferos, 106 pp. (in Spanish).
1830:
rodents, Hershkovitz properly identified diagnostic characters, making the name formally available for the first time. Accordingly, the correct year of publication for
2454:
Amerigo
Vespucci and the rat of Fernando de Noronha: a new genus and species of Rodentia (Muridae, Sigmodontinae) from a volcanic island off Brazil's continental shelf
4456:
1799:
353:
4521:
1823:
1446:
As in all oryzomyines, the upper molars all have one root on the inner (lingual) side and two on the outer (labial) side; in addition, the first upper molar in
1006:, which encompasses hundreds of species found across South America and into southern North America. Sigmodontinae itself is the largest subfamily of the family
4590:
1686:
region, which includes much of
Paraguay. Two specimens from Paraguay, collected 600 kilometres (400 mi) apart, differed by 1.4% in the sequence of the
1365:, to which some of the chewing muscles are attached, are entirely separate, joining only at their front edges, which are located below the first molar.
4630:
4430:
2744:
4469:
2695:
Phylogenetic relationships of oryzomyine rodents (Muroidea: Sigmodontinae): separate and combined analyses of morphological and molecular data
2573:
2490:
1439:, forming steps in the transition from the complex, low-crowned generalized oryzomyine molar pattern to the simpler, high-crowned pattern of
671:; they considered it closer to oryzomyines than to phyllotines, but declined to formally place it in Oryzomyini in the absence of explicit
1404:, is present. Unlike in most other oryzomyines, however, which have mesolophs reaching the labial margin of the molar, the mesolophs of
4625:
2427:
Small mammals of
Chapada dos Veadeiros National Park (Cerrado of Central Brazil): Ecologic, karyologic, and taxonomic considerations
4600:
4495:
342:
521:
1567:. Apparently, a whole heterochromatic arm was added to this chromosome; cases of similar variation are known from the rodents
4513:
2565:
1601:, the Y chromosome contains two large blocks of heterochromatin on its long arm. The karyotype is closely similar to that of
412:
tribe, a very diverse group including over one hundred species, mainly in South
America. This tribe is part of the subfamily
4474:
4605:
1563:
chromosome (with two equally long arms), yielding an FN of 55. One arm of the metacentric chromosome consists entirely of
1107:"), which contains tens of other species. Several of those display some adaptations to life in the water, being partially
2613:
4620:
2737:
1408:
are short and protrude only slightly from the middle of the molar. The corresponding structure in the lower molars, the
95:
499:
4610:
4508:
2426:
2551:
4615:
4526:
3943:
1334:, which are located behind the third molars, are excavated beyond the level of the palate, but not as deeply as in
1315:
396:, including specializations towards a semiaquatic lifestyle, such as the presence of membranes between the digits (
2504:
Mammals of northern
Colombia, preliminary report no. 8: Arboreal rice rats, a systematic revision of the subgenus
1717:, Argentina, outside the current distribution of the species. It is dated from the first millennium CE. The jaw's
1361:
of the lower incisor, a raising of the mandibular bone at the back end of the incisor, is well developed. The two
991:
on the chest, absence of a gall bladder, and some characters of the skull, suggesting that they form one natural,
4382:
4274:
3950:
465:
4267:
3056:
2941:
2820:
2411:
1148:
is a nondescript, medium-sized rat with long, soft fur. The upperparts are gray–brown and the underparts are
4296:
4223:
3985:
3929:
3915:
3908:
3901:
3700:
3693:
3620:
3399:
3163:
2730:
1603:
1064:
652:
1826:, because he did not explicitly name the diagnostic characteristics of the genus. In his 1962 monograph on
4344:
3978:
3936:
3686:
3571:
3213:
3141:
2827:
1551:(with one arm so short as to be almost invisible). However, in two specimens from the Brazilian states of
1519:
1059:, described in 1999 and 2008 respectively, were also recognized as members of the group. In 2006, a broad
1331:
1133:
adaptations towards a semiaquatic lifestyle may explain the morphological support for a relation between
4560:
4303:
3971:
3957:
3887:
3648:
3627:
3578:
3341:
3303:
3134:
3120:
3070:
3035:
2371:
Hershkovitz, 1962, p. 213; Bonvicino et al., 2008, p. 54; Wetzel and Lovett, 1974, p. 211
1714:
1237:
506:
was mostly ignored in the systematic literature, but from 1952 it was used briefly, in the combination "
207:
1362:
4417:
4391:
4216:
4194:
3992:
3964:
3922:
3894:
3880:
3851:
3655:
3585:
3406:
3348:
3295:
3227:
3170:
3127:
3113:
3106:
3099:
3063:
3049:
2962:
2934:
1759:
1718:
1556:
1153:
1130:
1060:
397:
2684:(Rodentia: Muridae) and the significance of Lund's collections from the caves of Lagoa Santa, Brazil
4595:
4095:
4066:
4052:
4044:
4000:
3707:
3678:
3641:
3311:
3177:
3092:
2948:
2652:
2522:
Evolution of
Neotropical cricetine rodents (Muridae) with special reference to the phyllotine group
1639:
1268:
393:
60:
1503:
1330:
has a flat palate, lacking a ridge on the middle that extends along the length of the palate. The
3858:
3767:
3753:
3613:
3549:
3287:
3241:
3206:
3042:
2927:
1491:
1475:
1463:
664:
525:
453:
251:
184:
90:
4565:
2476:
New and noteworthy records of
Rodentia (Erethizontidae, Sciuridae, and Cricetidae) from Paraguay
2447:
1725:, but the toothrow is relatively long (5.78 mm; 4.61 to 5.60 mm in three specimens of
1710:
1156:
is present between the second, third, and fourth toes, but the membranes are not as large as in
604:, and his opinion was mostly accepted in the next few decades. Scientific knowledge of the rare
4461:
1357:, an opening at the front of the mandible, just before the first molar, opens to the side. The
4547:
4404:
4187:
3837:
3788:
3634:
3606:
3599:
3592:
3535:
3521:
3499:
3492:
3355:
3220:
3199:
2955:
2603:
A new genus of oryzomyine rodent (Cricetidae: Sigmodontinae) from the
Pleistocene of Argentina
2591:
2579:
2569:
2486:
2443:
1245:
1233:
461:
2362:
Voss and Myers, 1991, p. 426; Bonvicino et al., 2005, p. 399; D'Elia et al., 2008, p. 49
1743:. In Argentina, it is mainly a species of the eastern Chaco and in Brazil it is found in the
4081:
4073:
3844:
3809:
3802:
3781:
3528:
3478:
3471:
3464:
3457:
3450:
3377:
3367:
3319:
2969:
2647:
2559:
2442:. Rio de Janeiro: Centro Pan-Americano de Febre Aftosa – OPAS/OMS, 120 pp. (in Portuguese).
1771:
1499:
1495:
1467:
1358:
1283:
996:
588:. Thus, he viewed the animal as a member of the phyllotine group of rodents, which includes
234:
43:
4552:
4487:
1204:
is absent, both important characters of Oryzomyini. As is characteristic of Sigmodontinae,
4534:
4059:
3830:
3823:
3795:
3774:
3760:
3542:
3485:
3234:
2722:
2531:
Moreira, C.N., Di-Nizo, C.B., Silva, M.J.d.J., Yonenaga-Yassuda, Y. and Ventura, K. 2013.
2249:
Voss and Myers, 1991, p. 423; Bonvicino et al., 2005, p. 399; Moreira et al., 2013, p. 202
1564:
1295:
1253:
572:(then used in a much broader sense than now), but it was moved to a separate genus, named
555:
2712:
2592:
Novedosos cricetidos (Mammalia, Rodentia) en el Holoceno de la Región Pampeana, Argentina
1314:
has spinous processes on its zygomatic plates. These genera also share relatively simple
1271:, located between the eyes, converges towards the front and is flanked by low beads. The
979:
When Voss and Carleton formally characterized Oryzomyini two years later, they did place
2704:
Wetzel, R.M. and Lovett, J.W. 1974. A collection of animals from the Chaco of Paraguay.
2602:
2521:
2503:
2475:
1478:
cervical (neck) vertebra and the first thoracic vertebra, an important character of the
1412:, is completely absent. The hindmost valley between cusps on the lower first molar, the
17:
4314:
4257:
4087:
1794:
The species is not known to be threatened and its conservation status is classified as
1729:) and the first molar is relatively narrow (1.28 mm; 1.30 to 1.40 mm in five
1586:
1502:(tail) vertebrae. Between the second and third caudal vertebrae, separate bones called
1354:
1299:
1249:
1181:
1108:
1067:
phylogenetic study of Oryzomyini provided further support for the relationship between
578:
4396:
2629:
1373:
1224:
The skull, which is short at the front, shows some typical oryzomyine characters. The
4584:
4482:
4206:
2881:
2871:
2799:
2638:
2555:
2532:
1795:
1523:
1479:
1413:
1291:
1272:
1030:
1003:
672:
413:
401:
357:
246:
in South America (in red); fossil record from Cueva Tixi, eastern Argentina, in blue
167:
80:
75:
2439:
Guia dos Roedores do Brasil, com chaves para gêneros baseadas em caracteres externos
1033:
studies published during the 1990s and 2000s supported a close relationship between
624:, because Bolivian animals are slightly larger and darker than those from Paraguay.
4409:
4286:
3870:
3389:
3025:
3013:
3003:
2905:
2894:
2859:
2851:
2840:
2713:
Jordfundne og nulevende Gnavere (Rodentia) fra Lagoa Santa, Minas Geraes, Brasilien
1687:
1659:
1594:
1590:
1294:, are long and narrow, extending between the first molars. The back margins of the
1276:
1264:
1201:
1165:
1149:
1055:
545:
494:
469:
457:
224:
2670:
Hershkovitz (Mammalia, Muridae) with an analysis of its phylogenetic relationships
2463:
Chebez, J.C., Pereira, J., Massoia, E., Di Giacomo, A.G. and Fortabat, S.H. 2005.
1615:
1275:
bone, located in the roof of the skull on the braincase, is nearly as wide as the
1176:
4500:
4443:
4376:
4022:
4012:
3816:
3667:
3082:
2991:
2981:
1651:
1548:
1389:
1079:. Within this group, morphological data supported a closer relationship between
389:
2495:
Hershkovitz, P.M. 1959. Two new genera of South American rodents (Cricetinae).
4325:
4245:
4235:
4119:
4108:
3731:
3720:
3561:
3153:
2917:
2810:
2793:
2754:
2547:
1827:
1779:
1683:
1675:
1671:
1598:
1569:
1560:
1539:
1526:; in some other cricetids, it perforates the far (distal) end of the humerus.
1409:
1397:
1260:
1104:
1049:
1043:
1023:
1007:
920:
845:
790:
691:
617:
597:
541:
482:
417:
409:
378:
338:
157:
4367:
436:). This was confirmed in 1991 that both names pertained to the same species.
3331:
3276:
3189:
2583:
1581:
1552:
1535:
1302:(cheekbones), are located before the first molars. Like its close relatives
1241:
1213:
1000:
992:
584:
568:
from Paraguay. In the next decades, it was viewed as an aberrant species of
552:
425:
364:
107:
51:
2296:
Musser and Carleton, 2005, p. 1164; Bonvicino et al., 2008, p. 54
1216:
part of the baculum, the central digit is smaller than those at the sides.
392:, spending much of their time in the water. The three genera share several
2042:
Voss and Myers, 1991, tables 1, 2; Bonvicino et al., 2008, p. 54
4539:
4361:
4177:
4165:
4154:
4034:
3511:
3440:
3428:
3418:
3263:
3253:
2775:
1763:
1748:
1655:
1643:
1624:
1544:
1401:
1393:
1350:
1117:
984:
879:
773:
725:
564:
488:
420:, which include many more species, mainly from Eurasia and the Americas.
384:
127:
4435:
3743:
1744:
1682:
show a similar pattern of variation: they are smaller and paler in the
1667:
1647:
1575:
1515:
1474:, as is characteristic of oryzomyines. The first ribs contact both the
1343:
1287:
1229:
1209:
1019:
1015:
613:
512:
349:
330:
608:—only three specimens were known when Hershkovitz described the genus
4448:
2787:
2781:
2769:
1506:
are present. These display a spinous process at the back, as in both
1225:
988:
473:
334:
147:
137:
117:
4338:
582:
in appearance, other features suggest it is more closely related to
2410:
Belentani, S.C. da S., Motta-Junior, J.C. and Talamoni, S.A. 2005.
1822:
in a 1959 article, but failed to make the name available under the
1346:
skull bone contains a conspicuous opening, as in most oryzomyines.
627:
In 1980, Argentinean zoologist Elio Massoia suggested that Winge's
576:, by Hershkovitz in 1959, who noted that although it is similar to
360:, although almost nothing is known about its diet or reproduction.
2694:
2661:
2453:
1663:
1614:
1372:
1175:
368:
2680:
2561:
Mammal Species of the World: A Taxonomic and Geographic Reference
1212:(penis bone) displaying large protuberances at the sides. In the
432:); and in 1921 on the basis of a live specimen from Paraguay (as
4422:
2628:
Percequillo, A.; Weksler, M.; Pardinas, U.; D'Elia, G. (2017) .
1197:
1126:
1037:
and two other oryzomyines with reduced or absent mesoloph(id)s,
1011:
970:
4342:
2726:
2412:
Notes on the food habits and prey selection of the maned wolf (
2269:
2267:
2257:
2255:
2219:
Weksler, 2006, p. 131; Carleton and Olson, 1999, pp. 49–50
540:
skull illustrated by Winge instead suggested affinities to the
1679:
1638:
is known from northeastern Argentina, probably south to about
1471:
1169:
1115:
and its relatives. Morphological data indicate that the genus
1200:, including one on the chest and three on the belly, and the
2114:
Voss and Myers, 1991, p. 422; Weksler, 2006, p. 31
1380:
was first found as a living animal in the Chaco of Paraguay.
456:
history. It was first described in 1888 by Danish zoologist
1559:, one pair of autosomes contains both an acrocentric and a
1188:) of the eastern United States is similar in appearance to
1490:, the fourth lumbar vertebra lacks the processes known as
1318:, perforations of the palate near the third molar. Unlike
612:
in 1959—increased in the following years, and in 1975 the
2612:
Pardiñas, U.F.J., Cirignoli, S. and Galliari, C.A. 2004.
2436:
Bonvicino, C.R., Oliveira, J.A. and D'Andrea, P.S. 2008.
1650:
and from there eastward through Brazil in the states of
667:. Voss and Myers also re-evaluated the relationships of
2198:
Weksler, 2006, pp. 43–44; Pardiñas, 2008, table 2
1263:
bones end bluntly close to the hindmost extent of the
2474:
D'Elia, G., Mora, I., Myers, P. and Owen, R.D. 2008.
1741:
open, usually humid tropical and subtropical lowlands
1091:, but DNA sequence data favored a clustering between
408:. Together, they form a unique assemblage within the
424:
was independently described in 1888 on the basis of
4351:
4313:
4284:
4255:
4233:
4204:
4175:
4151:
4129:
4105:
4032:
4010:
3868:
3741:
3717:
3665:
3559:
3509:
3438:
3416:
3387:
3365:
3329:
3273:
3251:
3187:
3151:
3080:
3023:
3001:
2979:
2915:
2891:
2869:
2837:
2808:
2497:
Proceedings of the Biological Society of Washington
1121:is the closest relative of the group that includes
516:species from central Brazil. In his 1960 review of
27:
Genus of rodent from South America with one species
2699:Bulletin of the American Museum of Natural History
2688:Bulletin of the American Museum of Natural History
376:, its closest living relatives are the large rats
2425:Bonvicino, C.R., Lemos, B. and Weksler, M. 2005.
1416:, is severely reduced, foreshadowing its loss in
404:crowns, both of which are at incipient stages in
356:has assessed its conservation status as being of
2515:Proceedings of the United States National Museum
2437:
1173:body mass 45 to 55 g (1.6 to 1.9 oz).
2653:10.2305/IUCN.UK.2016-3.RLTS.T18598A22367785.en
2738:
2537:2n = 56; FN = 54-55 (Rodentia, Sigmodontinae)
1824:International Code of Zoological Nomenclature
1026:, all mainly from Eurasia and North America.
8:
2416:) (Mammalia, Canidae) in southeastern Brazil
1236:, which in some sigmodontines separates two
983:in the group, even though it lacks complete
643:. Since then, the species has been known as
400:), and a reduction in the complexity of the
2706:University of Connecticut Occasional Papers
2096:Weksler, 2006, pp. 28–30, table 5
2087:Weksler, 2006, pp. 27–28, table 5
1228:is long, extending past the molars and the
4339:
2745:
2731:
2723:
1597:. In addition to heterochromatin near the
1482:. Unlike in most sigmodontines, including
233:
69:
42:
31:
2651:
2533:A remarkable autosomal heteromorphism in
2340:
2338:
1754:Nothing is known about behavior or diet.
1286:, which perforate the palate between the
1125:, but DNA sequence data from the nuclear
2478:(subscription required for full paper).
2074:
2072:
2070:
2068:
2066:
1957:
1955:
1953:
1388:and its close relatives, the molars are
1240:(openings) in the skull, is absent. The
1129:gene did not support this relationship;
995:group. Oryzomyini is now one of several
2283:
2281:
2279:
2123:Weksler, 2006, p. 32, table 5
1913:
1911:
1909:
1846:
1811:
2227:
2225:
2206:
2204:
2167:
2165:
1979:Voss and Carleton, 1993, p. 31–32
1854:
1852:
1850:
1818:Hershkovitz introduced the genus name
1298:, the flattened front portions of the
536:, noting that various features of the
54:skull as illustrated by Winge in 1888
2452:Carleton, M.D. and Olson, S.L. 1999.
2051:Weksler, 2006, pp. 17, 19, 58–59
1925:
1923:
1705:" (i.e., an unnamed species close to
1279:, but does not reach the squamosals.
7:
2660:Voss, R.S. and Carleton, M.D. 1993.
1611:Distribution, ecology, and variation
1522:is absent, as in all members of the
675:justification for such a placement.
4591:IUCN Red List least concern species
2639:IUCN Red List of Threatened Species
2618:(Rodentia: Cricetidae) in Argentina
1903:Hershkovitz, 1960, pp. 519–520
1894:Voss and Carleton, 1993, p. 31
1424:. A number of molar traits support
1353:(lower jaw) is short and deep. The
895:
871:
851:
841:
765:
741:
717:
697:
687:
616:population was named as a separate
480:, and placed it in the same genus (
2469:Temas de Naturaleza y Conservación
2389:Belentani et al., 2005, p. 96
2353:Bonvicino et al., 2008, p. 54
2344:Pardiñas et al., 2004, p. 108
2287:Pardiñas et al., 2004, p. 105
2141:Weksler, 2006, pp. 35–36, 131
1988:Musser and Carleton, 2005, table 1
1876:Musser and Carleton, 2005, p. 1164
1709:) is known from a cave deposit in
1623:have been found in pellets of the
25:
2465:Mamíferos de la Reserva El Bagual
2332:Voss and Myers, 1991, p. 425
2314:Voss and Myers, 1991, pp. 426–427
2189:Voss and Myers, 1991, p. 416
2078:Voss and Myers, 1991, p. 422
2024:Voss and Myers, 1991, p. 420
1778:). It is a preferred prey of the
1010:, other members of which include
476:. Winge described the species as
4631:Taxa named by Philip Hershkovitz
2380:Chebez et al., 2005, p. 484
2323:Pardiñas, 1995, pp. 199–201
94:
2679:Voss, R.S. and Myers, P. 1991.
1400:, is lacking, but another, the
1168:on the toes and several of the
428:cave specimens from Brazil (as
2566:Johns Hopkins University Press
2541:Genetics and Molecular Biology
1938:Hershkovitz, 1962, p. 208
1208:has a complex penis, with the
969:based on morphology (top) and
528:denied any affinities between
492:and two species now placed in
464:had collected in the caves of
1:
2132:Weksler, 2006, pp. 31–32
2033:Weksler, 2006, pp. 23–25
1758:has frequently been found in
1384:As in all oryzomyines except
1196:The female has four pairs of
460:, who reviewed the materials
2431:Brazilian Journal of Biology
2273:Moreira et al., 2013, p. 204
2261:Moreira et al., 2013, p. 202
1997:Pardiñas, 2008, p. 1275
1961:Voss and Myers, 1991, p. 423
1947:Hershkovitz, 1959, p. 9
1917:Voss and Myers, 1991, p. 424
1867:Voss and Myers, 1991, p. 418
1697:A fragmentary lower jaw of "
1666:, and far in the northeast,
1642:, northward through western
1137:and the other three genera.
486:) as the species now called
347:palm savanna and thorn scrub
2598:32(2):197–203 (in Spanish).
1770:) and also in those of the
1538:generally includes 56
1316:posterolateral palatal pits
1256:. The front part is short.
663:Pine and Wetzel, 1975, are
661:Pseudoryzomys wavrini reigi
622:Pseudoryzomys wavrini reigi
295:Pseudoryzomys wavrini reigi
4647:
2550:; Carleton, M. D. (2005).
2305:D'Elia et al., 2008, p. 49
2015:Weksler, 2006, p. 131
1970:Weksler, 2006, figs. 34–36
1929:Hershkovitz, 1959, pp. 8–9
1547:(non-sex chromosomes) are
1518:, the upper arm bone, the
1494:. There are three or four
551:In 1921, renowned British
4626:Mammals described in 1888
2764:
2674:American Museum Novitates
2622:Mastozoología Neotropical
2605:(subscription required).
2539:(subscription required).
2458:American Museum Novitates
2240:Weksler, 2006, p. 55
2231:Weksler, 2006, p. 53
2210:Weksler, 2006, p. 52
2180:Weksler, 2006, p. 42
2171:Weksler, 2006, p. 41
2159:Weksler, 2006, p. 36
2150:Weksler, 2006, p. 35
2105:Weksler, 2006, p. 31
2006:Weksler, 2006, p. 72
917:
900:
893:
876:
869:
856:
849:
787:
770:
763:
746:
739:
722:
715:
702:
695:
445:Discovery and recognition
345:. It is found in lowland
257:
250:
241:
232:
213:
206:
91:Scientific classification
89:
67:
58:
50:
41:
34:
2558:; Reeder, D. M. (eds.).
2502:Hershkovitz, P.M. 1960.
2060:Weksler, 2006, pp. 55–56
1858:Percequillo et al., 2016
1244:bone lacks a suspensory
679:Oryzomyine relationships
323:Brazilian false rice rat
18:Brazilian False Rice Rat
4601:Monotypic rodent genera
2601:Pardiñas, U.F.J. 2008.
2590:Pardiñas, U.F.J. 1995.
2471:4:467–499 (in Spanish).
1604:Holochilus brasiliensis
647:(Winge, 1888), because
2646:: e.T18598A115144867.
2552:"Superfamily Muroidea"
2520:Hershkovitz, P. 1962.
2438:
1632:
1520:entepicondylar foramen
1381:
1193:
999:recognized within the
600:group, which includes
4561:Paleobiology Database
4383:Pseudoryzomys simplex
4353:Pseudoryzomys simplex
2682:Pseudoryzomys simplex
2632:Pseudoryzomys simplex
2616:Pseudoryzomys simplex
2535:Pseudoryzomys simplex
2414:Chrysocyon brachyurus
1784:Chrysocyon brachyurus
1715:Buenos Aires Province
1707:Pseudoryzomys simplex
1636:Pseudoryzomys simplex
1618:
1589:are acrocentric, and
1376:
1179:
1146:Pseudoryzomys simplex
1047:. The extinct genera
645:Pseudoryzomys simplex
633:Pseudoryzomys wavrini
606:Pseudoryzomys wavrini
450:Pseudoryzomys simplex
422:Pseudoryzomys simplex
318:Pseudoryzomys simplex
303:Pseudoryzomys simplex
298:Pine and Wetzel, 1975
286:Pseudoryzomys wavrini
217:Pseudoryzomys simplex
36:Pseudoryzomys simplex
4606:Mammals of Argentina
4326:Ekbletomys hypenemus
2860:A. praeuniversitatis
2607:Journal of Mammalogy
2398:Díaz and Ojeda, 2000
1721:agrees with that of
1455:Postcranial skeleton
1431:s relationship with
1332:parapterygoid fossae
1099:to the exclusion of
1087:to the exclusion of
562:as a new species of
398:interdigital webbing
321:, also known as the
4621:Mammals of Paraguay
1790:Conservation status
1470:, 13 of which bear
1392:, low-crowned, and
1269:interorbital region
341:from south-central
61:Conservation status
4611:Mammals of Bolivia
2693:Weksler, M. 2006.
2664:Hesperomys molitor
1834:is 1962, not 1959.
1633:
1498:and about 29
1462:has 19 or 20
1382:
1267:bones. The narrow
1194:
1186:Oryzomys palustris
659:Thomas, 1921, and
629:Hesperomys simplex
579:Oryzomys palustris
526:Philip Hershkovitz
478:Hesperomys simplex
452:has had a complex
430:Hesperomys simplex
261:Hesperomys simplex
4616:Rodents of Brazil
4576:
4575:
4548:Open Tree of Life
4345:Taxon identifiers
4336:
4335:
2758:
2753:Species of tribe
2668:Holochilus magnus
2575:978-0-8018-8221-0
2526:Fieldiana Zoology
2491:978-987-98497-0-5
1363:masseteric ridges
1284:incisive foramina
1234:alisphenoid strut
977:
976:
965:Relationships of
959:
958:
950:
949:
941:
940:
932:
931:
858:Other Oryzomyini
838:
837:
829:
828:
820:
819:
811:
810:
802:
801:
704:Other Oryzomyini
462:Peter Wilhem Lund
314:
313:
308:
299:
291:
290:Hershkovitz, 1959
282:
274:
265:
188:
84:
16:(Redirected from
4638:
4569:
4568:
4556:
4555:
4543:
4542:
4530:
4529:
4517:
4516:
4504:
4503:
4491:
4490:
4478:
4477:
4465:
4464:
4452:
4451:
4439:
4438:
4426:
4425:
4413:
4412:
4400:
4399:
4387:
4386:
4385:
4372:
4371:
4370:
4340:
3944:O. longicaudatus
3368:Microakodontomys
2992:D. albimaculatus
2942:C. maracajuensis
2821:A. galapagoensis
2756:
2747:
2740:
2733:
2724:
2711:Winge, H. 1888.
2662:A new genus for
2657:
2655:
2614:Distribution of
2609:89(5):1270–1278.
2587:
2568:. p. 1164.
2564:(3rd ed.).
2441:
2405:Literature cited
2399:
2396:
2390:
2387:
2381:
2378:
2372:
2369:
2363:
2360:
2354:
2351:
2345:
2342:
2333:
2330:
2324:
2321:
2315:
2312:
2306:
2303:
2297:
2294:
2288:
2285:
2274:
2271:
2262:
2259:
2250:
2247:
2241:
2238:
2232:
2229:
2220:
2217:
2211:
2208:
2199:
2196:
2190:
2187:
2181:
2178:
2172:
2169:
2160:
2157:
2151:
2148:
2142:
2139:
2133:
2130:
2124:
2121:
2115:
2112:
2106:
2103:
2097:
2094:
2088:
2085:
2079:
2076:
2061:
2058:
2052:
2049:
2043:
2040:
2034:
2031:
2025:
2022:
2016:
2013:
2007:
2004:
1998:
1995:
1989:
1986:
1980:
1977:
1971:
1968:
1962:
1959:
1948:
1945:
1939:
1936:
1930:
1927:
1918:
1915:
1904:
1901:
1895:
1892:
1886:
1883:
1877:
1874:
1868:
1865:
1859:
1856:
1835:
1816:
1776:Bubo virginianus
1772:great horned owl
1468:lumbar vertebrae
1430:
1359:capsular process
1300:zygomatic arches
1296:zygomatic plates
896:
872:
852:
842:
766:
742:
718:
698:
688:
683:
682:
657:Oryzomys wavrini
655:for the animal;
560:Oryzomys wavrini
502:Winge proposed,
489:Lundomys molitor
434:Oryzomys wavrini
306:
297:
289:
280:
278:Oryzomys wavrini
273:Trouessart, 1898
272:
269:Oryzomys simplex
263:
242:Distribution of
237:
219:
183:
99:
98:
78:
73:
72:
46:
32:
21:
4646:
4645:
4641:
4640:
4639:
4637:
4636:
4635:
4581:
4580:
4577:
4572:
4564:
4559:
4551:
4546:
4538:
4535:Observation.org
4533:
4525:
4520:
4512:
4507:
4499:
4494:
4486:
4481:
4473:
4468:
4460:
4455:
4447:
4442:
4434:
4429:
4421:
4416:
4408:
4403:
4395:
4390:
4381:
4380:
4375:
4366:
4365:
4360:
4347:
4337:
4332:
4309:
4280:
4251:
4229:
4200:
4171:
4147:
4125:
4101:
4028:
4006:
3951:O. magellanicus
3864:
3852:O. sydandersoni
3737:
3713:
3661:
3621:N. maculiventer
3555:
3505:
3434:
3412:
3383:
3378:M. transitorius
3361:
3325:
3269:
3247:
3214:H. megacephalus
3183:
3164:H. brasiliensis
3147:
3076:
3019:
2997:
2975:
2911:
2887:
2865:
2833:
2804:
2760:
2751:
2627:
2576:
2546:
2407:
2402:
2397:
2393:
2388:
2384:
2379:
2375:
2370:
2366:
2361:
2357:
2352:
2348:
2343:
2336:
2331:
2327:
2322:
2318:
2313:
2309:
2304:
2300:
2295:
2291:
2286:
2277:
2272:
2265:
2260:
2253:
2248:
2244:
2239:
2235:
2230:
2223:
2218:
2214:
2209:
2202:
2197:
2193:
2188:
2184:
2179:
2175:
2170:
2163:
2158:
2154:
2149:
2145:
2140:
2136:
2131:
2127:
2122:
2118:
2113:
2109:
2104:
2100:
2095:
2091:
2086:
2082:
2077:
2064:
2059:
2055:
2050:
2046:
2041:
2037:
2032:
2028:
2023:
2019:
2014:
2010:
2005:
2001:
1996:
1992:
1987:
1983:
1978:
1974:
1969:
1965:
1960:
1951:
1946:
1942:
1937:
1933:
1928:
1921:
1916:
1907:
1902:
1898:
1893:
1889:
1884:
1880:
1875:
1871:
1866:
1862:
1857:
1848:
1844:
1839:
1838:
1817:
1813:
1808:
1792:
1756:P. simplex
1737:P. simplex
1731:P. simplex
1727:P. simplex
1723:P. simplex
1703:P. simplex
1613:
1593:is larger than
1587:sex chromosomes
1565:heterochromatin
1532:
1457:
1428:
1371:
1254:tympanic cavity
1252:, the roof the
1248:contacting the
1222:
1143:
960:
951:
942:
933:
839:
830:
821:
812:
803:
681:
665:junior synonyms
631:and the living
556:Oldfield Thomas
538:H. simplex
508:Oecomys simplex
504:H. simplex
447:
442:
228:
221:
215:
202:
199:P. simplex
182:
93:
85:
74:
70:
63:
28:
23:
22:
15:
12:
11:
5:
4644:
4642:
4634:
4633:
4628:
4623:
4618:
4613:
4608:
4603:
4598:
4593:
4583:
4582:
4574:
4573:
4571:
4570:
4557:
4544:
4531:
4518:
4505:
4492:
4479:
4466:
4453:
4440:
4427:
4414:
4401:
4388:
4373:
4357:
4355:
4349:
4348:
4343:
4334:
4333:
4331:
4330:
4319:
4317:
4315:Incertae sedis
4311:
4310:
4308:
4307:
4300:
4292:
4290:
4282:
4281:
4279:
4278:
4271:
4263:
4261:
4258:Transandinomys
4253:
4252:
4250:
4249:
4241:
4239:
4231:
4230:
4228:
4227:
4220:
4212:
4210:
4202:
4201:
4199:
4198:
4195:S. ucayalensis
4191:
4183:
4181:
4173:
4172:
4170:
4169:
4160:
4158:
4149:
4148:
4146:
4145:
4137:
4135:
4127:
4126:
4124:
4123:
4114:
4112:
4103:
4102:
4100:
4099:
4092:
4085:
4077:
4070:
4063:
4056:
4048:
4040:
4038:
4030:
4029:
4027:
4026:
4018:
4016:
4008:
4007:
4005:
4004:
3996:
3989:
3982:
3975:
3968:
3961:
3954:
3947:
3940:
3933:
3926:
3919:
3912:
3905:
3898:
3891:
3884:
3876:
3874:
3866:
3865:
3863:
3862:
3855:
3848:
3841:
3834:
3827:
3820:
3813:
3806:
3799:
3792:
3785:
3778:
3771:
3764:
3757:
3749:
3747:
3739:
3738:
3736:
3735:
3726:
3724:
3715:
3714:
3712:
3711:
3704:
3701:N. narboroughi
3697:
3694:N. fernandinae
3690:
3682:
3673:
3671:
3663:
3662:
3660:
3659:
3652:
3645:
3638:
3631:
3624:
3617:
3610:
3603:
3596:
3589:
3582:
3575:
3572:N. albigularis
3567:
3565:
3557:
3556:
3554:
3553:
3546:
3539:
3532:
3525:
3517:
3515:
3507:
3506:
3504:
3503:
3496:
3489:
3482:
3475:
3468:
3461:
3454:
3446:
3444:
3436:
3435:
3433:
3432:
3424:
3422:
3414:
3413:
3411:
3410:
3403:
3395:
3393:
3385:
3384:
3382:
3381:
3373:
3371:
3363:
3362:
3360:
3359:
3352:
3345:
3342:M. caliginosus
3337:
3335:
3327:
3326:
3324:
3323:
3315:
3307:
3304:M. desmarestii
3299:
3291:
3282:
3280:
3271:
3270:
3268:
3267:
3259:
3257:
3249:
3248:
3246:
3245:
3238:
3231:
3224:
3217:
3210:
3203:
3195:
3193:
3185:
3184:
3182:
3181:
3174:
3167:
3159:
3157:
3149:
3148:
3146:
3145:
3138:
3131:
3124:
3117:
3110:
3103:
3096:
3088:
3086:
3078:
3077:
3075:
3074:
3067:
3060:
3057:E. macconnelli
3053:
3046:
3039:
3031:
3029:
3021:
3020:
3018:
3017:
3009:
3007:
2999:
2998:
2996:
2995:
2987:
2985:
2977:
2976:
2974:
2973:
2966:
2959:
2952:
2945:
2938:
2931:
2923:
2921:
2913:
2912:
2910:
2909:
2900:
2898:
2889:
2888:
2886:
2885:
2877:
2875:
2867:
2866:
2864:
2863:
2855:
2846:
2844:
2835:
2834:
2832:
2831:
2828:A. xanthaeolus
2824:
2816:
2814:
2806:
2805:
2803:
2802:
2796:
2790:
2784:
2778:
2772:
2765:
2762:
2761:
2752:
2750:
2749:
2742:
2735:
2727:
2721:
2720:
2717:E Museo Lundii
2709:
2708:2(13):203–216.
2702:
2691:
2677:
2658:
2625:
2624:11(1):105–108.
2610:
2599:
2588:
2574:
2544:
2543:36(2):201–206.
2529:
2518:
2500:
2493:
2483:
2472:
2461:
2450:
2434:
2433:65(3):395–406.
2423:
2406:
2403:
2401:
2400:
2391:
2382:
2373:
2364:
2355:
2346:
2334:
2325:
2316:
2307:
2298:
2289:
2275:
2263:
2251:
2242:
2233:
2221:
2212:
2200:
2191:
2182:
2173:
2161:
2152:
2143:
2134:
2125:
2116:
2107:
2098:
2089:
2080:
2062:
2053:
2044:
2035:
2026:
2017:
2008:
1999:
1990:
1981:
1972:
1963:
1949:
1940:
1931:
1919:
1905:
1896:
1887:
1878:
1869:
1860:
1845:
1843:
1840:
1837:
1836:
1810:
1809:
1807:
1804:
1791:
1788:
1612:
1609:
1531:
1528:
1456:
1453:
1370:
1367:
1355:mental foramen
1250:tegmen tympani
1221:
1218:
1182:marsh rice rat
1142:
1139:
975:
974:
962:
961:
957:
956:
953:
952:
948:
947:
944:
943:
939:
938:
935:
934:
930:
929:
926:
925:
916:
913:
912:
909:
908:
899:
894:
892:
889:
888:
885:
884:
875:
870:
868:
865:
864:
861:
860:
855:
850:
848:
840:
836:
835:
832:
831:
827:
826:
823:
822:
818:
817:
814:
813:
809:
808:
805:
804:
800:
799:
796:
795:
786:
783:
782:
779:
778:
769:
764:
762:
759:
758:
755:
754:
745:
740:
738:
735:
734:
731:
730:
721:
716:
714:
711:
710:
707:
706:
701:
696:
694:
686:
680:
677:
651:is the oldest
446:
443:
441:
438:
337:in the family
327:false oryzomys
312:
311:
310:
309:
300:
292:
283:
275:
266:
255:
254:
248:
247:
239:
238:
230:
229:
222:
211:
210:
204:
203:
196:
194:
190:
189:
175:
171:
170:
165:
161:
160:
155:
151:
150:
145:
141:
140:
135:
131:
130:
125:
121:
120:
115:
111:
110:
105:
101:
100:
87:
86:
68:
65:
64:
59:
56:
55:
48:
47:
39:
38:
26:
24:
14:
13:
10:
9:
6:
4:
3:
2:
4643:
4632:
4629:
4627:
4624:
4622:
4619:
4617:
4614:
4612:
4609:
4607:
4604:
4602:
4599:
4597:
4594:
4592:
4589:
4588:
4586:
4579:
4567:
4562:
4558:
4554:
4549:
4545:
4541:
4536:
4532:
4528:
4523:
4519:
4515:
4510:
4506:
4502:
4497:
4493:
4489:
4484:
4480:
4476:
4471:
4467:
4463:
4458:
4454:
4450:
4445:
4441:
4437:
4432:
4428:
4424:
4419:
4415:
4411:
4406:
4402:
4398:
4397:Pseudoryzomys
4393:
4389:
4384:
4378:
4374:
4369:
4363:
4359:
4358:
4356:
4354:
4350:
4346:
4341:
4329:
4327:
4321:
4320:
4318:
4316:
4312:
4306:
4305:
4301:
4299:
4298:
4297:Z. brevicauda
4294:
4293:
4291:
4289:
4288:
4283:
4277:
4276:
4275:T. talamancae
4272:
4270:
4269:
4265:
4264:
4262:
4260:
4259:
4254:
4248:
4247:
4243:
4242:
4240:
4238:
4237:
4232:
4226:
4225:
4221:
4219:
4218:
4214:
4213:
4211:
4209:
4208:
4207:Sigmodontomys
4203:
4197:
4196:
4192:
4190:
4189:
4185:
4184:
4182:
4180:
4179:
4174:
4168:
4167:
4166:R. primigenus
4162:
4161:
4159:
4157:
4156:
4150:
4144:
4143:
4139:
4138:
4136:
4134:
4133:
4132:Pseudoryzomys
4128:
4122:
4121:
4116:
4115:
4113:
4111:
4110:
4104:
4098:
4097:
4096:O. peninsulae
4093:
4091:
4090:
4089:O. palustris
4086:
4084:
4083:
4078:
4076:
4075:
4071:
4069:
4068:
4067:O. dimidiatus
4064:
4062:
4061:
4057:
4055:
4054:
4053:O. antillarum
4049:
4047:
4046:
4045:O. albiventer
4042:
4041:
4039:
4037:
4036:
4031:
4025:
4024:
4020:
4019:
4017:
4015:
4014:
4009:
4003:
4002:
3997:
3995:
3994:
3990:
3988:
3987:
3986:O. stramineus
3983:
3981:
3980:
3976:
3974:
3973:
3969:
3967:
3966:
3962:
3960:
3959:
3955:
3953:
3952:
3948:
3946:
3945:
3941:
3939:
3938:
3934:
3932:
3931:
3930:O. fulvescens
3927:
3925:
3924:
3920:
3918:
3917:
3916:O. flavescens
3913:
3911:
3910:
3909:O. destructor
3906:
3904:
3903:
3902:O. chacoensis
3899:
3897:
3896:
3892:
3890:
3889:
3885:
3883:
3882:
3878:
3877:
3875:
3873:
3872:
3867:
3861:
3860:
3859:O. trinitatis
3856:
3854:
3853:
3849:
3847:
3846:
3842:
3840:
3839:
3835:
3833:
3832:
3828:
3826:
3825:
3821:
3819:
3818:
3814:
3812:
3811:
3807:
3805:
3804:
3800:
3798:
3797:
3793:
3791:
3790:
3786:
3784:
3783:
3779:
3777:
3776:
3772:
3770:
3769:
3768:O. catherinae
3765:
3763:
3762:
3758:
3756:
3755:
3754:O. auyantepui
3751:
3750:
3748:
3746:
3745:
3740:
3734:
3733:
3728:
3727:
3725:
3723:
3722:
3716:
3710:
3709:
3705:
3703:
3702:
3698:
3696:
3695:
3691:
3689:
3688:
3687:N. indefessus
3683:
3681:
3680:
3675:
3674:
3672:
3670:
3669:
3664:
3658:
3657:
3653:
3651:
3650:
3649:N. pectoralis
3646:
3644:
3643:
3639:
3637:
3636:
3632:
3630:
3629:
3628:N. meridensis
3625:
3623:
3622:
3618:
3616:
3615:
3611:
3609:
3608:
3604:
3602:
3601:
3597:
3595:
3594:
3590:
3588:
3587:
3583:
3581:
3580:
3579:N. auriventer
3576:
3574:
3573:
3569:
3568:
3566:
3564:
3563:
3558:
3552:
3551:
3547:
3545:
3544:
3540:
3538:
3537:
3533:
3531:
3530:
3526:
3524:
3523:
3519:
3518:
3516:
3514:
3513:
3508:
3502:
3501:
3497:
3495:
3494:
3490:
3488:
3487:
3483:
3481:
3480:
3476:
3474:
3473:
3469:
3467:
3466:
3462:
3460:
3459:
3455:
3453:
3452:
3448:
3447:
3445:
3443:
3442:
3437:
3431:
3430:
3426:
3425:
3423:
3421:
3420:
3415:
3409:
3408:
3404:
3402:
3401:
3400:M. altissimus
3397:
3396:
3394:
3392:
3391:
3386:
3380:
3379:
3375:
3374:
3372:
3370:
3369:
3364:
3358:
3357:
3353:
3351:
3350:
3349:M. robustulus
3346:
3344:
3343:
3339:
3338:
3336:
3334:
3333:
3328:
3322:
3321:
3316:
3314:
3313:
3308:
3306:
3305:
3300:
3298:
3297:
3296:M. curazensis
3292:
3290:
3289:
3284:
3283:
3281:
3279:
3278:
3272:
3266:
3265:
3261:
3260:
3258:
3256:
3255:
3250:
3244:
3243:
3239:
3237:
3236:
3232:
3230:
3229:
3228:H. perenensis
3225:
3223:
3222:
3218:
3216:
3215:
3211:
3209:
3208:
3204:
3202:
3201:
3197:
3196:
3194:
3192:
3191:
3186:
3180:
3179:
3175:
3173:
3172:
3168:
3166:
3165:
3161:
3160:
3158:
3156:
3155:
3150:
3144:
3143:
3142:H. saturatior
3139:
3137:
3136:
3132:
3130:
3129:
3125:
3123:
3122:
3118:
3116:
3115:
3111:
3109:
3108:
3104:
3102:
3101:
3097:
3095:
3094:
3090:
3089:
3087:
3085:
3084:
3079:
3073:
3072:
3068:
3066:
3065:
3061:
3059:
3058:
3054:
3052:
3051:
3047:
3045:
3044:
3040:
3038:
3037:
3033:
3032:
3030:
3028:
3027:
3022:
3016:
3015:
3011:
3010:
3008:
3006:
3005:
3000:
2994:
2993:
2989:
2988:
2986:
2984:
2983:
2978:
2972:
2971:
2967:
2965:
2964:
2960:
2958:
2957:
2953:
2951:
2950:
2946:
2944:
2943:
2939:
2937:
2936:
2932:
2930:
2929:
2925:
2924:
2922:
2920:
2919:
2914:
2908:
2907:
2902:
2901:
2899:
2897:
2896:
2890:
2884:
2883:
2879:
2878:
2876:
2874:
2873:
2872:Amphinectomys
2868:
2862:
2861:
2856:
2854:
2853:
2848:
2847:
2845:
2843:
2842:
2836:
2830:
2829:
2825:
2823:
2822:
2818:
2817:
2815:
2813:
2812:
2807:
2801:
2800:Sigmodontinae
2797:
2795:
2791:
2789:
2785:
2783:
2779:
2777:
2773:
2771:
2767:
2766:
2763:
2759:
2748:
2743:
2741:
2736:
2734:
2729:
2728:
2725:
2718:
2714:
2710:
2707:
2703:
2700:
2696:
2692:
2689:
2685:
2683:
2678:
2675:
2671:
2669:
2665:
2659:
2654:
2649:
2645:
2641:
2640:
2635:
2633:
2626:
2623:
2619:
2617:
2611:
2608:
2604:
2600:
2597:
2593:
2589:
2585:
2581:
2577:
2571:
2567:
2563:
2562:
2557:
2556:Wilson, D. E.
2553:
2549:
2548:Musser, G. G.
2545:
2542:
2538:
2536:
2530:
2527:
2523:
2519:
2516:
2512:
2511:
2507:
2501:
2498:
2494:
2492:
2488:
2484:
2481:
2477:
2473:
2470:
2466:
2462:
2459:
2455:
2451:
2449:
2445:
2440:
2435:
2432:
2428:
2424:
2421:
2417:
2415:
2409:
2408:
2404:
2395:
2392:
2386:
2383:
2377:
2374:
2368:
2365:
2359:
2356:
2350:
2347:
2341:
2339:
2335:
2329:
2326:
2320:
2317:
2311:
2308:
2302:
2299:
2293:
2290:
2284:
2282:
2280:
2276:
2270:
2268:
2264:
2258:
2256:
2252:
2246:
2243:
2237:
2234:
2228:
2226:
2222:
2216:
2213:
2207:
2205:
2201:
2195:
2192:
2186:
2183:
2177:
2174:
2168:
2166:
2162:
2156:
2153:
2147:
2144:
2138:
2135:
2129:
2126:
2120:
2117:
2111:
2108:
2102:
2099:
2093:
2090:
2084:
2081:
2075:
2073:
2071:
2069:
2067:
2063:
2057:
2054:
2048:
2045:
2039:
2036:
2030:
2027:
2021:
2018:
2012:
2009:
2003:
2000:
1994:
1991:
1985:
1982:
1976:
1973:
1967:
1964:
1958:
1956:
1954:
1950:
1944:
1941:
1935:
1932:
1926:
1924:
1920:
1914:
1912:
1910:
1906:
1900:
1897:
1891:
1888:
1882:
1879:
1873:
1870:
1864:
1861:
1855:
1853:
1851:
1847:
1841:
1833:
1832:Pseudoryzomys
1829:
1825:
1821:
1820:Pseudoryzomys
1815:
1812:
1805:
1803:
1801:
1797:
1796:least concern
1789:
1787:
1785:
1781:
1777:
1773:
1769:
1765:
1761:
1757:
1752:
1750:
1746:
1742:
1738:
1734:
1732:
1728:
1724:
1720:
1716:
1712:
1708:
1704:
1700:
1699:Pseudoryzomys
1695:
1692:
1691:
1685:
1681:
1677:
1673:
1669:
1665:
1662:, São Paulo,
1661:
1658:, Tocantins,
1657:
1653:
1649:
1645:
1641:
1637:
1630:
1626:
1622:
1621:Pseudoryzomys
1617:
1610:
1608:
1606:
1605:
1600:
1596:
1592:
1588:
1584:
1583:
1578:
1577:
1572:
1571:
1566:
1562:
1558:
1554:
1550:
1546:
1541:
1537:
1529:
1527:
1525:
1524:Sigmodontinae
1521:
1517:
1513:
1509:
1505:
1501:
1497:
1493:
1489:
1485:
1481:
1480:Sigmodontinae
1477:
1473:
1469:
1465:
1461:
1460:Pseudoryzomys
1454:
1452:
1449:
1448:Pseudoryzomys
1444:
1442:
1438:
1434:
1427:
1426:Pseudoryzomys
1423:
1419:
1415:
1414:posteroflexid
1411:
1407:
1406:Pseudoryzomys
1403:
1399:
1395:
1391:
1387:
1379:
1378:Pseudoryzomys
1375:
1368:
1366:
1364:
1360:
1356:
1352:
1347:
1345:
1341:
1337:
1333:
1329:
1328:Pseudoryzomys
1325:
1321:
1317:
1313:
1312:Pseudoryzomys
1309:
1305:
1301:
1297:
1293:
1289:
1285:
1280:
1278:
1274:
1273:interparietal
1270:
1266:
1262:
1257:
1255:
1251:
1247:
1243:
1239:
1235:
1231:
1227:
1219:
1217:
1215:
1214:cartilaginous
1211:
1207:
1206:Pseudoryzomys
1203:
1199:
1191:
1190:Pseudoryzomys
1187:
1183:
1178:
1174:
1171:
1167:
1166:tufts of hair
1163:
1159:
1155:
1151:
1147:
1140:
1138:
1136:
1132:
1128:
1124:
1123:Pseudoryzomys
1120:
1119:
1114:
1113:Pseudoryzomys
1110:
1106:
1102:
1098:
1097:Pseudoryzomys
1094:
1090:
1089:Pseudoryzomys
1086:
1082:
1078:
1077:Pseudoryzomys
1074:
1070:
1066:
1062:
1061:morphological
1058:
1057:
1052:
1051:
1046:
1045:
1040:
1036:
1035:Pseudoryzomys
1032:
1027:
1025:
1021:
1017:
1013:
1009:
1005:
1004:Sigmodontinae
1002:
998:
994:
990:
986:
982:
981:Pseudoryzomys
972:
968:
967:Pseudoryzomys
964:
963:
955:
954:
946:
945:
937:
936:
928:
927:
924:
923:
922:
915:
914:
911:
910:
907:
906:
905:
904:Pseudoryzomys
898:
897:
891:
890:
887:
886:
883:
882:
881:
874:
873:
867:
866:
863:
862:
859:
854:
853:
847:
844:
843:
834:
833:
825:
824:
816:
815:
807:
806:
798:
797:
794:
793:
792:
785:
784:
781:
780:
777:
776:
775:
768:
767:
761:
760:
757:
756:
753:
752:
751:
750:Pseudoryzomys
744:
743:
737:
736:
733:
732:
729:
728:
727:
720:
719:
713:
712:
709:
708:
705:
700:
699:
693:
690:
689:
685:
684:
678:
676:
674:
670:
669:Pseudoryzomys
666:
662:
658:
654:
653:specific name
650:
646:
642:
638:
634:
630:
625:
623:
619:
615:
611:
610:Pseudoryzomys
607:
603:
599:
596:, not of the
595:
591:
587:
586:
581:
580:
575:
574:Pseudoryzomys
571:
567:
566:
561:
557:
554:
549:
547:
543:
539:
535:
531:
527:
523:
519:
515:
514:
509:
505:
501:
500:other species
497:
496:
491:
490:
485:
484:
479:
475:
471:
467:
463:
459:
455:
451:
444:
439:
437:
435:
431:
427:
423:
419:
415:
414:Sigmodontinae
411:
407:
406:Pseudoryzomys
403:
399:
395:
391:
387:
386:
381:
380:
375:
374:
373:Pseudoryzomys
370:
366:
361:
359:
358:least concern
355:
351:
348:
344:
343:South America
340:
336:
332:
328:
324:
320:
319:
307:Massoia, 1980
304:
301:
296:
293:
287:
284:
279:
276:
270:
267:
262:
259:
258:
256:
253:
249:
245:
244:Pseudoryzomys
240:
236:
231:
226:
220:
218:
212:
209:
208:Binomial name
205:
201:
200:
195:
192:
191:
186:
181:
180:
179:Pseudoryzomys
176:
173:
172:
169:
168:Sigmodontinae
166:
163:
162:
159:
156:
153:
152:
149:
146:
143:
142:
139:
136:
133:
132:
129:
126:
123:
122:
119:
116:
113:
112:
109:
106:
103:
102:
97:
92:
88:
82:
77:
76:Least Concern
66:
62:
57:
53:
49:
45:
40:
37:
33:
30:
19:
4578:
4352:
4323:
4302:
4295:
4287:Zygodontomys
4285:
4273:
4268:T. bolivaris
4266:
4256:
4244:
4234:
4224:S. aphrastus
4222:
4215:
4205:
4193:
4186:
4176:
4164:
4153:
4141:
4140:
4131:
4130:
4118:
4107:
4094:
4088:
4080:
4072:
4065:
4058:
4051:
4043:
4033:
4023:O. balneator
4021:
4011:
3999:
3991:
3984:
3979:O. rupestris
3977:
3970:
3963:
3956:
3949:
3942:
3937:O. griseolus
3935:
3928:
3921:
3914:
3907:
3900:
3893:
3886:
3879:
3871:Oligoryzomys
3869:
3857:
3850:
3843:
3838:O. speciosus
3836:
3829:
3822:
3815:
3808:
3801:
3794:
3789:O. flavicans
3787:
3780:
3773:
3766:
3759:
3752:
3742:
3732:N. vespuccii
3730:
3719:
3706:
3699:
3692:
3685:
3677:
3666:
3656:N. pirrensis
3654:
3647:
3640:
3633:
3626:
3619:
3612:
3605:
3598:
3591:
3586:N. caracolus
3584:
3577:
3570:
3560:
3550:N. squamipes
3548:
3541:
3534:
3527:
3520:
3510:
3498:
3491:
3484:
3477:
3470:
3463:
3456:
3449:
3439:
3427:
3417:
3405:
3398:
3390:Microryzomys
3388:
3376:
3366:
3354:
3347:
3340:
3330:
3318:
3312:M. georginae
3310:
3302:
3294:
3286:
3275:
3262:
3252:
3240:
3233:
3226:
3219:
3212:
3205:
3198:
3188:
3176:
3171:H. chacarius
3169:
3162:
3152:
3140:
3135:H. rostratus
3133:
3126:
3121:H. melanotis
3119:
3112:
3105:
3098:
3091:
3081:
3069:
3062:
3055:
3048:
3041:
3034:
3026:Euryoryzomys
3024:
3012:
3004:Eremoryzomys
3002:
2990:
2980:
2968:
2963:C. subflavus
2961:
2954:
2947:
2940:
2935:C. langguthi
2933:
2926:
2916:
2904:
2895:Carletonomys
2893:
2880:
2870:
2858:
2850:
2841:Agathaeromys
2839:
2826:
2819:
2809:
2716:
2705:
2698:
2690:206:414–432.
2687:
2681:
2673:
2667:
2663:
2643:
2637:
2631:
2621:
2615:
2606:
2595:
2560:
2540:
2534:
2525:
2517:110:513–568.
2514:
2509:
2505:
2496:
2479:
2468:
2457:
2430:
2422:13(1):95–98.
2419:
2413:
2394:
2385:
2376:
2367:
2358:
2349:
2328:
2319:
2310:
2301:
2292:
2245:
2236:
2215:
2194:
2185:
2176:
2155:
2146:
2137:
2128:
2119:
2110:
2101:
2092:
2083:
2056:
2047:
2038:
2029:
2020:
2011:
2002:
1993:
1984:
1975:
1966:
1943:
1934:
1899:
1890:
1881:
1872:
1863:
1831:
1819:
1814:
1793:
1783:
1775:
1767:
1755:
1753:
1736:
1735:
1730:
1726:
1722:
1706:
1702:
1698:
1696:
1689:
1660:Minas Gerais
1635:
1634:
1628:
1620:
1602:
1580:
1574:
1568:
1533:
1511:
1507:
1504:hemal arches
1487:
1483:
1466:(chest) and
1459:
1458:
1447:
1445:
1440:
1436:
1432:
1425:
1421:
1417:
1405:
1385:
1383:
1377:
1348:
1339:
1335:
1327:
1323:
1319:
1311:
1307:
1303:
1281:
1265:premaxillary
1258:
1223:
1205:
1202:gall bladder
1195:
1189:
1185:
1161:
1157:
1145:
1144:
1134:
1122:
1116:
1112:
1100:
1096:
1092:
1088:
1084:
1080:
1076:
1072:
1068:
1056:Carletonomys
1054:
1048:
1042:
1038:
1034:
1031:phylogenetic
1028:
993:monophyletic
980:
978:
966:
919:
918:
903:
902:
901:
878:
877:
857:
789:
788:
772:
771:
749:
748:
747:
724:
723:
703:
673:phylogenetic
668:
660:
656:
648:
644:
640:
636:
632:
628:
626:
621:
609:
605:
601:
593:
589:
583:
577:
573:
569:
563:
559:
550:
537:
533:
529:
524:mammalogist
522:Field Museum
517:
511:
507:
503:
498:. Like most
493:
487:
481:
477:
470:Minas Gerais
458:Herluf Winge
449:
448:
433:
429:
421:
405:
388:, which are
383:
377:
372:
371:
365:only species
362:
326:
322:
317:
316:
315:
302:
294:
285:
281:Thomas, 1921
277:
268:
260:
243:
216:
214:
198:
197:
178:
177:
35:
29:
4444:iNaturalist
4377:Wikispecies
4304:Z. brunneus
4188:S. melanops
4013:Oreoryzomys
3972:O. nigripes
3958:O. microtis
3888:O. arenalis
3845:O. superans
3810:O. phaeotis
3803:O. paricola
3782:O. concolor
3668:Nesoryzomys
3642:N. nimbosus
3536:N. palmipes
3522:N. apicalis
3500:N. tenuipes
3493:N. spinosus
3429:M. hammondi
3288:M. audreyae
3242:H. yunganus
3207:H. laticeps
3178:H. sciureus
3128:H. rhabdops
3114:H. intectus
3107:H. fuscatus
3100:H. chapmani
3083:Handleyomys
3071:E. russatus
3036:E. emmonsae
2982:Drymoreomys
2949:C. marinhus
2852:A. donovani
2798:Subfamily:
2757:(rice rats)
2755:Oryzomyini
2719:1(3):1–200.
2596:Ameghiniana
2482:1784:39–57.
2420:Biocièncias
1885:Winge, 1888
1688:cytochrome
1652:Mato Grosso
1646:to eastern
1619:Remains of
1561:metacentric
1549:acrocentric
1540:chromosomes
1492:anapophyses
1390:brachyodont
1326:, however,
1232:bones. The
1141:Description
553:mammalogist
466:Lagoa Santa
416:and family
390:semiaquatic
264:Winge, 1888
185:Hershkovitz
164:Subfamily:
4596:Oryzomyini
4585:Categories
4246:S. angouya
4236:Sooretamys
4142:P. simplex
4120:P. nivalis
4109:Pennatomys
4082:O. nelsoni
4074:O. gorgasi
3993:O. vegetus
3965:O. moojeni
3923:O. fornesi
3895:O. brendae
3881:O. andinus
3831:O. rutilus
3824:O. roberti
3796:O. mamorae
3775:O. cleberi
3761:O. bicolor
3721:Noronhomys
3708:N. swarthi
3679:N. darwini
3614:N. levipes
3562:Nephelomys
3529:N. grandis
3479:N. paracou
3472:N. musseri
3465:N. minutus
3458:N. guianae
3451:N. dubosti
3407:M. minutus
3356:M. zunigae
3264:L. molitor
3221:H. oniscus
3200:H. acritus
3154:Holochilus
3093:H. alfaroi
3064:E. nitidus
3050:E. legatus
2928:C. goytaca
2918:Cerradomys
2882:A. savamis
2811:Aegialomys
2794:Cricetidae
2701:296:1–149.
2676:3085:1–39.
2666:Winge and
2460:3256:1–59.
1842:References
1828:phyllotine
1780:maned wolf
1719:morphology
1711:Cueva Tixi
1678:. Certain
1676:subspecies
1672:Pernambuco
1599:centromere
1570:Peromyscus
1508:Holochilus
1484:Holochilus
1441:Holochilus
1433:Holochilus
1422:Holochilus
1410:mesolophid
1398:anteroloph
1386:Holochilus
1336:Holochilus
1320:Holochilus
1308:Holochilus
1162:Holochilus
1131:convergent
1093:Holochilus
1081:Holochilus
1069:Holochilus
1050:Noronhomys
1044:Holochilus
1008:Cricetidae
973:(bottom).
921:Holochilus
846:Oryzomyini
791:Holochilus
692:Oryzomyini
641:P. wavrini
637:H. simplex
618:subspecies
598:oryzomyine
558:described
542:phyllotine
510:", for an
483:Hesperomys
418:Cricetidae
410:oryzomyine
394:characters
379:Holochilus
339:Cricetidae
158:Cricetidae
4217:S. alfari
4060:O. couesi
4001:O. victus
3635:N. moerex
3607:N. keaysi
3600:N. devius
3593:N. childi
3543:N. rattus
3486:N. pictus
3332:Melanomys
3320:M. luciae
3277:Megalomys
3190:Hylaeamys
3014:E. polius
2956:C. scotti
2906:C. cailoi
2768:Kingdom:
2528:46:1–524.
2448:0101-6970
1806:Footnotes
1768:Tyto alba
1739:inhabits
1629:Tyto alba
1582:Thaptomys
1557:São Paulo
1553:Tocantins
1545:autosomes
1536:karyotype
1530:Karyotype
1514:. On the
1242:squamosal
1230:maxillary
1065:molecular
1001:subfamily
594:Phyllotis
585:Phyllotis
546:sigmodont
454:taxonomic
426:subfossil
193:Species:
114:Kingdom:
108:Eukaryota
52:Lectotype
4514:13000868
4462:10914263
4362:Wikidata
4178:Scolomys
4155:Reigomys
4035:Oryzomys
3512:Nectomys
3441:Neacomys
3419:Mindomys
3254:Lundomys
3235:H. tatei
3043:E. lamia
2970:C. vivoi
2792:Family:
2788:Rodentia
2782:Mammalia
2776:Chordata
2774:Phylum:
2770:Animalia
2584:62265494
2510:Oryzomys
2508:, genus
2499:72:5–10.
1764:barn owl
1749:Caatinga
1644:Paraguay
1625:barn owl
1512:Lundomys
1488:Lundomys
1464:thoracic
1437:Lundomys
1418:Lundomys
1402:mesoloph
1394:bunodont
1351:mandible
1340:Lundomys
1324:Lundomys
1304:Lundomys
1290:and the
1288:incisors
1277:frontals
1238:foramina
1158:Lundomys
1135:Oryzomys
1118:Oryzomys
1111:, as do
1101:Lundomys
1085:Lundomys
1073:Lundomys
1039:Lundomys
1029:Several
1024:deermice
1020:hamsters
1016:lemmings
985:mesoloph
880:Lundomys
774:Lundomys
726:Oryzomys
614:Bolivian
602:Oryzomys
570:Oryzomys
565:Oryzomys
548:groups.
440:Taxonomy
385:Lundomys
350:habitats
252:Synonyms
154:Family:
148:Rodentia
138:Mammalia
128:Chordata
124:Phylum:
118:Animalia
104:Domain:
81:IUCN 3.1
4501:1002666
4436:2438678
4423:1180092
4368:Q899221
3744:Oecomys
2786:Order:
2780:Class:
2506:Oecomys
2480:Zootaxa
1798:by the
1762:of the
1760:pellets
1745:Cerrado
1668:Alagoas
1648:Bolivia
1585:. Both
1576:Clyomys
1516:humerus
1476:seventh
1344:mastoid
1246:process
1210:baculum
1154:Webbing
1109:aquatic
1105:clade D
649:simplex
590:Calomys
534:Oecomys
530:simplex
518:Oecomys
513:Oecomys
495:Calomys
367:in the
331:species
329:, is a
227:, 1888)
174:Genus:
144:Order:
134:Class:
79: (
4566:380011
4527:218834
4475:633314
3817:O. rex
2582:
2572:
2489:
2446:
1579:, and
1500:caudal
1496:sacral
1451:back.
1369:Molars
1342:. The
1292:molars
1226:palate
1164:. The
1075:, and
1022:, and
997:tribes
989:mammae
474:Brazil
335:rodent
187:, 1962
4553:38050
4540:86966
4488:18598
4457:IRMNG
4449:45387
4410:6WGTQ
2554:. In
1701:aff.
1684:Chaco
1664:Bahia
1656:Goiás
1429:'
1261:nasal
1220:Skull
1198:teats
1012:voles
402:molar
369:genus
225:Winge
4522:NCBI
4483:IUCN
4470:ITIS
4431:GBIF
2644:2016
2580:OCLC
2570:ISBN
2487:ISBN
2444:ISSN
1800:IUCN
1747:and
1680:bats
1670:and
1640:30°S
1555:and
1534:The
1510:and
1486:and
1472:ribs
1435:and
1420:and
1349:The
1338:and
1322:and
1306:and
1282:The
1259:The
1180:The
1170:pads
1150:buff
1127:IRBP
1095:and
1083:and
1063:and
1053:and
1041:and
971:IRBP
639:and
592:and
532:and
382:and
363:The
354:IUCN
4509:MSW
4496:MDD
4418:EoL
4405:CoL
4392:ADW
2648:doi
1786:).
1733:).
1160:or
544:or
333:of
325:or
4587::
4563::
4550::
4537::
4524::
4511::
4498::
4485::
4472::
4459::
4446::
4433::
4420::
4407::
4394::
4379::
4364::
2715:.
2697:.
2686:.
2672:.
2642:.
2636:.
2620:.
2594:.
2578:.
2524:.
2513:.
2467:.
2456:.
2429:.
2418:.
2337:^
2278:^
2266:^
2254:^
2224:^
2203:^
2164:^
2065:^
1952:^
1922:^
1908:^
1849:^
1713:,
1654:,
1631:).
1607:.
1573:,
1443:.
1310:,
1071:,
1018:,
1014:,
620:,
520:,
472:,
468:,
305::
288::
271::
4328:"
4324:"
4322:†
4163:†
4152:†
4117:†
4106:†
4079:†
4050:†
3998:†
3729:†
3718:†
3684:†
3676:†
3317:†
3309:†
3301:†
3293:†
3285:†
3274:†
2903:†
2892:†
2857:†
2849:†
2838:†
2746:e
2739:t
2732:v
2656:.
2650::
2634:"
2630:"
2586:.
1782:(
1774:(
1766:(
1690:b
1627:(
1595:Y
1591:X
1192:.
1184:(
223:(
83:)
20:)
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