720:, is a process by which iguana eggs are hatched in an incubator and the animals are protected and fed until they are large enough to be protected from predation upon them. In the case of the Utila iguana, 50% of the animals hatched at the Center are maintained for the head-start program and the rest are released into mangrove forests after hatching. The purpose is to get the animals to a size where they are more capable of fleeing from or fighting off predators. The program has proven successful, as the iguanas behave like their wild-born counterparts. The success of the Utila program serves as a blueprint for other such programs in the Caribbean, particularly with
225:
94:
713:, the Conservation Project of the Utila Iguana (CPUI) was founded. The International Iguana Society and the CPUI have sought to purchase land to preserve habitats for the iguanas and plan to establish an outpost staffed by Iguana Research and Breeding station personnel, who will aid in monitoring the property and work with developers to select building sites that preserve as much undisturbed beach area as possible.
527:
52:
33:
1783:
568:, which typically inhabits the drier habitats on Utila. It has interbred with this very species and produced viable offspring. From evolutionary and ecological perspectives, inhabiting brackish mangrove forests entails very specific adaptations of diet, behavior, and resource utilization. It is one of only two known species of lizard, the other being a species of
405:. Although several zoos and wildlife associations have instituted programs for the iguanas on Utila, the species still finds itself threatened due to overhunting and may face more of a threat in the form of habitat loss. Extreme conservation efforts are in place to try to prevent this species from going extinct.
670:
Gunther Köhler found the species at the brink of extinction, perhaps even functionally extinct in the wild as of 1994 due to overhunting and its restricted habitat. As a result, the Iguana
Research and Breeding Station was built in April 1997 with the help and funds of various organizations such as
657:
females are forced to migrate to nearby sandy beaches in order to bury their clutches of eggs so they can incubate in the hot sun. After digging their nest burrows and laying their eggs, the females abandon the nests and return to the mangroves. Sixty to seventy-four days later the hatchlings emerge
661:
The hatchlings are 15 centimeters (5.9 in) long, the body length being a mere 3 centimeters (1.2 in) with the tail accounting for 12 centimeters (4.7 in) of its total length. The hatchlings' dark skin color enables them to blend in with the dark floor of the mangrove forests to help
586:
The Utila iguana has a grey-brown to black coloring when young, the only species of spiny-tail iguana with such a dark color when young. Other members of the genus have a green or yellow coloring when young and turn darker with age. As this animal matures it can be a blue or light gray in color,
590:
Males achieve a maximum total length (including tail) of 76 centimeters (30 in), while females are typically 30% smaller at 56 centimeters (22 in) total length. Males have a small dewlap and a dorsal crest made up of 56 large dorsal spines, making the animal
394:. Males may grow up to 76 centimeters (30 in) in length, while females are smaller, with a length of up to 56 centimeters (22 in). Eggs are laid in sandy beaches and hatch about 60–76 days later, with the hatchlings returning to live in the mangrove forests.
706:, exotic invasive plants cover the ground near the mangroves and make the area inappropriate for nesting sites. The iguana is locally hunted for meat, although efforts to educate locals have helped reduce this somewhat in recent years.
1665:
2330:
2378:
2260:
2299:
716:
The Iguana
Research and Breeding station employs a "head-starting" program for newly hatched iguanas. "Head-starting", originally used to protect hatching
684:
2408:
702:. Beach habitat is being lost as natural vegetation is removed in preparation for hotel and road construction. According to a survey conducted by the
2234:
1602:
772:
469:
2273:
397:
Brought to the brink of extinction by the 1990s due to hunting, it was brought back to international attention by German herpetologist Dr.
2388:
1697:
2398:
1374:
2403:
1639:
1582:
1548:
768:
or over-hunting. The population is currently stable, but future declines are expected as a result of the threats mentioned above.
764:
for the first time outside of Utila, an important step to ensure their survival if the species is lost from its natural habitat by
1499:
690:
This species currently has an estimated wild population of 10,000 animals in 2–3 subpopulations, but is greatly threatened by
2278:
2393:
1985:
56:
595:. This dorsal crest consists of white and black spines arranged in alternating groups of two or three of the same color.
1970:
672:
653:
for the first year of their lives. As the Utila iguana cannot successfully lay its eggs in the mangrove swamps, the
93:
2304:
2000:
1945:
256:
1908:
2200:
2095:
2070:
1918:
1690:
480:
457:
431:
742:
The Utila iguana is maintained in a number of zoos throughout Europe, as well as two in the United States (
2135:
2105:
2045:
1838:
840:
2173:
2005:
1863:
1802:
897:
633:
322:
198:
878:
2221:
2182:
1939:
1928:
1913:
1893:
752:
340:
1205:
273:
2383:
2110:
1933:
1923:
1873:
1818:
486:
41:
224:
2100:
2020:
1950:
1868:
1683:
1434:
1078:
743:
676:
650:
564:
496:
427:
232:
214:
88:
1782:
2265:
452:(Κτενός), meaning "comb" (referring to the comblike spines on the lizard's back and tail), and
2312:
2208:
1980:
1903:
1808:
1635:
1578:
592:
398:
332:
286:
2353:
260:
2325:
2317:
2010:
1843:
1450:
1378:
1200:
574:
500:. Access to Utila may have involved over-water dispersal during hurricanes, as is known for
491:
2291:
2187:
2090:
1975:
1627:
1523:
691:
679:, AFE-COHDEFOR (State Forestry Administration-Honduran Forestry Development Corporation),
551:
507:
1552:
1660:
562:
was pushed into the mangrove swamps due to competition from the larger, more aggressive
478:
is believed to have evolved from mainland-based ancestors, and may share ancestors with
2015:
1995:
1059:
747:
468:
form of
Stejneger's friend and colleague Frank Baker, who was a former director of the
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61:
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155:
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2031:
1990:
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387:
32:
2158:
645:
Adults make their homes within holes in various mangrove trees and maintain an
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165:
105:
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Maryon, D.; Ardon, D.; Martinez, A.; Clayson, S.; Pasachnik, S.A. (2018).
2152:
1736:
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543:
383:
344:
145:
125:
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Hope hatches for critically endangered lizard on the brink of extinction
2239:
1961:
1829:
1730:
1661:
Fort Worth Zoo Museum of Living Art – Species List: Utila Island iguana
994:
722:
515:
475:
424:
382:
as it is born a dark color as opposed to bright green or yellow. It is
375:
371:
355:
325:
135:
1439:"On a new species of spiny-tailed iguana from Utilla Island, Honduras"
2252:
2195:
2056:
1718:
699:
654:
612:
569:
417:
363:
115:
2129:
1275:
Gutsche, Alexander (2005). "Distribution and
Habitat Utilization of
2338:
1315:
STEJNEGER 1901 (Squamata: Iguanidae) on Isla de Utila, Honduras".
954:
616:
539:
465:
435:
336:
78:
72:
709:
In 2004, as a result of Köhler's expedition and subsequent book,
623:
that inhabit the mangroves. It has been observed eating smaller
619:, but they will opportunistically eat smaller animals, eggs, and
550:
is an inhabitant of 8 square kilometres (3.1 sq mi) of
1337:
Dirksen L, Gutsche A (2006). "Beobachtungen zur
Saurophagie bei
703:
2226:
2133:
1749:
1679:
694:, as mangrove forests are being used as garbage dump sites and
1176:
1174:
1172:
1170:
1168:
1166:
1164:
1162:
1655:
1468:
1675:
587:
depending on heat conditions or even the animal's temper.
1307:
Gutsche A, Köhler G (2004). "A fertile hybrid between
1599:"International Species Information System Abstracts"
2142:
2079:
2054:
2029:
1959:
1882:
1852:
1827:
1790:
1763:
1254:". The Reptile Database. www.reptile-database.org.
514:to the mainland lost during the close of the last
1522:: Zoological Society of San Diego. Archived from
1415:: Meet the World's fastest lizard and its kin".
1206:10.2305/IUCN.UK.2018-1.RLTS.T44181A125203850.en
1547:Burgess, Rachel; Fiallos, Maria (2003-03-31).
370:, forced there due to competition from larger
1691:
1634:. Offenbach, Germany: Herpeton. p. 368.
558:and rare among reptiles, it is believed that
8:
1577:. Berkeley: University of California Press.
1549:"Utila For Sale: Where will the Iguanas Go?"
1411:Malfatti, Mark (2007). "A Look at the Genus
698:for the construction of homes, resorts, and
2379:IUCN Red List critically endangered species
781:
681:BICA (Bay Islands Conservation Association)
378:of iguana found on Utila, and unique among
2130:
1760:
1746:
1698:
1684:
1676:
1406:
1404:
1402:
1400:
1398:
1396:
1394:
775:, the following zoological parks maintain
685:National Autonomous University of Honduras
223:
50:
31:
20:
1204:
756:breeding center. In September, 2007, the
649:existence whereas the young are strictly
578:, that lives solely in mangrove forests.
1603:International Species Information System
1493:
1491:
1375:"Rare Utila Iguanas Hatch at London Zoo"
773:International Species Information System
525:
1158:
1270:
1268:
1266:
1264:
1262:
1260:
390:, although it can be an opportunistic
1542:
1540:
1476:STEJNEGER 1901 auf Utila, Honduras".
7:
1656:Iguana Research and Breeding Station
1551:. Honduras This Week. Archived from
1368:
1366:
1364:
1362:
1360:
1358:
1356:
1332:
1330:
1302:
1300:
1298:
1296:
1294:
1192:IUCN Red List of Threatened Species
677:Senckenberg Nature Research Society
1508:Iguana Specialist Group Newsletter
750:), each institution serving as an
530:Utila Spiny-tailed Iguana, in the
494:closer to these two than it is to
374:. It is the smallest of the three
14:
1919:Honduran paleate spinytail iguana
1575:Iguanas: Biology and Conservation
2409:Taxa named by Leonhard Stejneger
1781:
470:National Zoo in Washington, D.C.
366:to exclusively inhabit brackish
281:Ctenosaura (Loganiosaura) bakeri
92:
456:(σαύρα), meaning "lizard". Its
430:in 1901, while working for the
362:and one of only two species of
1373:Eccleston, Paul (2007-09-09).
760:successfully managed to breed
1:
1986:Northern Bahamian rock iguana
1894:Northeastern spinytail iguana
993:Museum of Natural History of
658:and move back to the swamps.
542:, an island off the northern
354:The Utila iguana is the only
1971:Turks and Caicos rock iguana
1671:Profile on West Coast Iguana
1455:10.5479/si.00963801.1217.467
673:Frankfurt Zoological Society
1934:Guatemalan spinytail iguana
1909:Yellowback spinytail iguana
1632:Reptiles of Central America
711:Reptiles of Central America
403:Reptiles of Central America
2425:
2389:Reptiles described in 1901
1869:Galapagos pink land iguana
2399:Endemic fauna of Honduras
1839:Campeche spinytail iguana
1779:
1759:
1745:
1713:
1573:Alberts, Allison (2004).
784:
305:Utila spiny-tailed iguana
238:
231:
222:
204:
197:
89:Scientific classification
87:
70:
48:
39:
30:
23:
2404:Lizards of the Caribbean
1991:Grand Cayman blue iguana
1924:Oaxacan spinytail iguana
1844:Yucatán spinytail iguana
1341:(Squamata: Iguanidae)".
796:
793:
790:
787:
522:Distribution and habitat
309:Baker's spinytail iguana
2096:Angel Island chuckwalla
2071:Lesser Antillean iguana
1929:Roatán spinytail iguana
738:Zoological institutions
432:Smithsonian Institution
416:was first described by
2106:San Esteban chuckwalla
2046:Catalina desert iguana
1951:Black spinytail iguana
1199:: e.T44181A125203850.
841:Cotswold Wildlife Park
797:Born in the last year
535:
444:, is derived from two
2006:Anegada ground iguana
2001:Lesser Caymans iguana
1914:Cape spinytail iguana
1864:Galapagos land iguana
1803:Brachylophus bulabula
898:Durrell Wildlife Park
634:Hemidactylus frenatus
631:) and geckos such as
615:, leaves, stems, and
529:
323:critically endangered
318:wishiwilly del suampo
57:Critically Endangered
2394:Reptiles of Honduras
2101:Monserrat chuckwalla
1940:Ctenosaura pectinata
1874:Santa Fe land iguana
1498:Binns, John (2003).
1443:Proc. U.S. Natl. Mus
1228:"Appendices | CITES"
726:species such as the
603:Like most iguanids,
380:spiny-tailed iguanas
303:, also known as the
251:Enyaliosaurus bakeri
2016:San Salvador iguana
1904:Balsas armed lizard
1819:Fiji crested iguana
1435:Stejneger, Leonhard
779:in their exhibits.
666:Conservation status
42:Conservation status
2111:Spotted chuckwalla
2021:Mona ground iguana
1502:Ctenosaura bakeri
1470:Ctenosaura similis
1309:Ctenosaura similis
1079:Fresno Chaffee Zoo
744:Fresno Chaffee Zoo
593:sexually dimorphic
536:
428:Leonhard Stejneger
2364:
2363:
2313:Open Tree of Life
2201:ctenosaura-bakeri
2188:Ctenosaura_bakeri
2174:Ctenosaura bakeri
2144:Ctenosaura bakeri
2136:Taxon identifiers
2127:
2126:
2123:
2122:
2119:
2118:
2091:Common chuckwalla
1981:Rhinoceros iguana
1809:Lau banded iguana
1474:Ctenosaura bakeri
1449:(1217): 467–468.
1417:Reptiles Magazine
1339:Ctenosaura bakeri
1277:Ctenosaura bakeri
1252:Ctenosaura bakeri
1185:Ctenosaura bakeri
1150:
1149:
1033:European Subtotal
777:Ctenosaura bakeri
771:According to the
762:Ctenosaura bakeri
662:elude predators.
605:Ctenosaura bakeri
548:Ctenosaura bakeri
414:Ctenosaura bakeri
343:off the coast of
341:Islas de la Bahía
335:to the island of
300:Ctenosaura bakeri
296:
295:
268:Ctenosaura bakeri
242:Ctenosaura bakeri
208:Ctenosaura bakeri
82:
65:
25:Ctenosaura bakeri
16:Species of lizard
2416:
2357:
2356:
2347:
2346:
2334:
2333:
2321:
2320:
2308:
2307:
2295:
2294:
2282:
2281:
2269:
2268:
2256:
2255:
2243:
2242:
2230:
2229:
2217:
2216:
2204:
2203:
2191:
2190:
2178:
2177:
2176:
2163:
2162:
2161:
2131:
1946:Club tail iguana
1785:
1761:
1747:
1700:
1693:
1686:
1677:
1645:
1614:
1613:
1611:
1610:
1595:
1589:
1588:
1570:
1564:
1563:
1561:
1560:
1555:on June 21, 2008
1544:
1535:
1534:
1532:
1531:
1495:
1486:
1485:
1472:(GRAY 1831) und
1465:
1459:
1458:
1431:
1425:
1424:
1408:
1389:
1388:
1386:
1385:
1379:London Telegraph
1370:
1351:
1350:
1334:
1325:
1324:
1311:(GRAY 1831) and
1304:
1289:
1288:
1272:
1255:
1248:
1242:
1241:
1239:
1238:
1224:
1218:
1217:
1215:
1213:
1208:
1178:
782:
575:Anolis utilensis
554:. Unique among
552:mangrove forests
492:phylogenetically
330:spinytail iguana
290:
277:
264:
247:
227:
210:
97:
96:
76:
59:
54:
53:
35:
21:
2424:
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2413:
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2238:
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2220:
2212:
2207:
2199:
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2181:
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2171:
2166:
2157:
2156:
2151:
2138:
2128:
2115:
2075:
2050:
2025:
2011:Ricord's iguana
1976:Jamaican iguana
1955:
1878:
1848:
1823:
1786:
1777:
1755:
1741:
1709:
1704:
1652:
1642:
1628:Köhler, Gunther
1626:
1623:
1621:Further reading
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1608:
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1527:
1500:"Taxon Reports
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1410:
1409:
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1371:
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1328:
1323:(3/4): 201–206.
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879:Zoo d'Amnéville
740:
692:loss of habitat
668:
643:
601:
584:
524:
508:Lesser Antilles
481:C. melanosterna
411:
368:mangrove swamps
284:
283:
272:— Wilson &
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246:Stejneger, 1901
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1650:External links
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1565:
1536:
1526:on May 9, 2008
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1060:Fort Worth Zoo
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597:
583:
580:
523:
520:
410:
407:
399:Gunther Köhler
386:and primarily
294:
293:
292:
291:
278:
265:
248:
236:
235:
229:
228:
220:
219:
213:
202:
201:
195:
194:
190:C. bakeri
187:
185:
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180:
173:
169:
168:
163:
159:
158:
153:
149:
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143:
139:
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71:
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49:
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40:
37:
36:
28:
27:
15:
13:
10:
9:
6:
4:
3:
2:
2421:
2410:
2407:
2405:
2402:
2400:
2397:
2395:
2392:
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2387:
2385:
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2380:
2377:
2376:
2374:
2367:
2355:
2349:
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2327:
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2319:
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2301:
2297:
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2249:
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2236:
2232:
2228:
2223:
2219:
2215:
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2206:
2202:
2197:
2193:
2189:
2184:
2180:
2175:
2169:
2165:
2160:
2154:
2150:
2149:
2147:
2145:
2141:
2137:
2132:
2112:
2109:
2107:
2104:
2102:
2099:
2097:
2094:
2092:
2089:
2088:
2086:
2084:
2083:
2078:
2072:
2069:
2067:
2064:
2063:
2061:
2059:
2058:
2053:
2047:
2044:
2042:
2041:Desert iguana
2039:
2038:
2036:
2034:
2033:
2028:
2022:
2019:
2017:
2014:
2012:
2009:
2007:
2004:
2002:
1999:
1997:
1994:
1992:
1989:
1987:
1984:
1982:
1979:
1977:
1974:
1972:
1969:
1968:
1966:
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1958:
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1807:
1805:
1804:
1800:
1799:
1797:
1795:
1794:
1789:
1784:
1774:
1773:Marine iguana
1771:
1770:
1768:
1766:
1765:Amblyrhynchus
1762:
1758:
1753:
1748:
1744:
1738:
1734:
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1649:
1643:
1641:3-936180-02-4
1637:
1633:
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1625:
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1620:
1604:
1600:
1594:
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1586:
1584:0-520-23854-0
1580:
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1039:
1036:
1034:
1031:
1030:
1026:
1023:
1020:
1017:
1015:
1014:Whipsnade Zoo
1012:
1011:
1007:
1004:
1001:
998:
996:
992:
991:
987:
984:
981:
978:
976:
975:Rotterdam Zoo
973:
972:
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952:
948:
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869:
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844:
842:
839:
838:
834:
831:
828:
825:
823:
822:Blackpool Zoo
820:
819:
815:
812:
809:
806:
804:
803:Barcelona Zoo
801:
800:
783:
780:
778:
774:
769:
767:
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759:
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749:
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737:
735:
733:
729:
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724:
719:
714:
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693:
688:
686:
682:
678:
674:
665:
663:
659:
656:
652:
648:
640:
638:
636:
635:
630:
629:Iguana iguana
626:
625:green iguanas
622:
618:
614:
610:
607:is primarily
606:
598:
596:
594:
588:
581:
579:
577:
576:
571:
567:
566:
561:
557:
553:
549:
545:
541:
533:
528:
521:
519:
517:
513:
509:
505:
504:
503:Iguana iguana
499:
498:
493:
489:
488:
483:
482:
477:
472:
471:
467:
463:
459:
458:specific name
455:
451:
447:
443:
442:
437:
433:
429:
426:
423:
419:
415:
408:
406:
404:
401:and his book
400:
395:
393:
389:
385:
381:
377:
373:
369:
365:
361:
357:
352:
350:
346:
342:
339:, one of the
338:
334:
331:
327:
324:
320:
319:
314:
310:
306:
302:
301:
288:
282:
279:
275:
269:
266:
262:
258:
252:
249:
243:
240:
239:
237:
234:
230:
226:
221:
216:
211:
209:
203:
200:
199:Binomial name
196:
192:
191:
186:
183:
182:
179:
178:
174:
171:
170:
167:
164:
161:
160:
157:
154:
151:
150:
147:
144:
141:
140:
137:
134:
131:
130:
127:
124:
121:
120:
117:
114:
111:
110:
107:
104:
101:
100:
95:
90:
86:
80:
74:
69:
63:
58:
47:
43:
38:
34:
29:
26:
22:
19:
2366:
2143:
2080:
2066:Green iguana
2055:
2030:
1996:Cuban iguana
1960:
1938:
1899:Utila iguana
1898:
1883:
1853:
1828:
1801:
1793:Brachylophus
1791:
1764:
1631:
1607:. Retrieved
1605:. 2008-08-21
1593:
1574:
1568:
1557:. Retrieved
1553:the original
1528:. Retrieved
1524:the original
1511:
1507:
1501:
1481:
1477:
1473:
1469:
1463:
1446:
1442:
1429:
1423:(11): 64–73.
1420:
1416:
1412:
1382:. Retrieved
1346:
1342:
1338:
1320:
1316:
1312:
1308:
1284:
1280:
1276:
1251:
1246:
1235:. Retrieved
1231:
1222:
1210:. Retrieved
1196:
1190:
1184:
1144:
1139:
1134:
1129:
1124:
1117:
1112:
1107:
1102:
1097:
1052:
1047:
1042:
1037:
1032:
936:Paignton Zoo
785:Institution
776:
770:
761:
751:
741:
728:Cuban Iguana
721:
715:
710:
708:
689:
669:
660:
644:
641:Reproduction
632:
628:
604:
602:
589:
585:
573:
563:
559:
547:
537:
501:
495:
485:
479:
473:
461:
453:
449:
439:
413:
412:
402:
396:
353:
317:
316:
312:
308:
304:
299:
298:
297:
289:et al., 2000
280:
267:
250:
241:
207:
205:
189:
188:
176:
24:
18:
2248:iNaturalist
2168:Wikispecies
2032:Dipsosaurus
1484:(1): 55–59.
1349:(3): 51–52.
1279:on Utila".
1212:13 November
1098:US Subtotal
860:Chester Zoo
732:Blue Iguana
718:sea turtles
651:terrestrial
609:herbivorous
582:Description
538:Endemic to
512:land bridge
490:, as it is
487:C. palearis
388:herbivorous
75:Appendix II
2384:Ctenosaura
2373:Categories
2082:Sauromalus
1885:Ctenosaura
1855:Conolophus
1814:Gau iguana
1609:2008-09-05
1559:2008-09-04
1530:2008-09-04
1520:California
1413:Ctenosaura
1384:2008-09-07
1317:Salamandra
1237:2022-01-14
1153:References
917:London Zoo
791:Female(s)
766:hurricanes
758:London Zoo
696:deforested
621:arthropods
565:C. similis
532:London Zoo
497:C. similis
441:Ctenosaura
177:Ctenosaura
152:Suborder:
1717:Kingdom:
1707:Iguanidae
1516:San Diego
1313:C. bakeri
1287:(3): 143.
1232:cites.org
611:, eating
560:C. bakeri
466:Latinized
464:, is the
425:zoologist
418:Norwegian
392:carnivore
349:Caribbean
287:G. Köhler
215:Stejneger
184:Species:
166:Iguanidae
112:Kingdom:
106:Eukaryota
2339:Species+
2266:10369467
2159:Q1409488
2153:Wikidata
1737:Squamata
1731:Reptilia
1725:Chordata
1723:Phylum:
1719:Animalia
1630:(2003).
1437:(1901).
794:Unknown
788:Male(s)
746:and the
683:and the
647:arboreal
556:iguanids
544:Honduras
422:American
409:Taxonomy
384:arboreal
345:Honduras
257:J. Meyer
233:Synonyms
162:Family:
146:Squamata
136:Reptilia
126:Chordata
122:Phylum:
116:Animalia
102:Domain:
62:IUCN 3.1
2354:5716756
2279:1056068
2240:2459608
1962:Cyclura
1830:Cachryx
1754:species
1735:Order:
1729:Class:
995:Tournai
753:ex-situ
723:Cyclura
700:marinas
613:flowers
546:coast,
516:ice age
506:in the
476:species
448:words:
436:generic
376:species
372:species
356:species
347:in the
333:endemic
326:species
321:, is a
313:swamper
172:Genus:
156:Iguania
142:Order:
132:Class:
77: (
60: (
2351:uBio:
2331:bakeri
2318:603272
2305:500334
2227:963751
2196:ARKive
2057:Iguana
1752:Extant
1638:
1581:
1478:Elaphe
1343:Elaphe
1281:Iguana
1125:Totals
675:, the
655:gravid
462:bakeri
450:ctenos
438:name,
434:. The
420:-born
364:lizard
360:iguana
276:, 1973
263:, 1973
261:Wilson
259:&
217:, 1901
2292:44181
2261:IRMNG
2253:35319
2214:32762
1514:(1).
955:Plock
617:fruit
570:anole
540:Utila
510:or a
454:saura
446:Greek
337:Utila
79:CITES
73:CITES
2344:5031
2300:NCBI
2287:IUCN
2274:ITIS
2235:GBIF
1636:ISBN
1579:ISBN
1214:2021
1197:2018
730:and
704:IUCN
671:the
599:Diet
484:and
474:The
274:Hahn
2222:EoL
2209:CoL
2183:ADW
1451:doi
1201:doi
1145:20
1027:13
957:Zoo
358:of
328:of
315:or
2375::
2341::
2328::
2326:RD
2315::
2302::
2289::
2276::
2263::
2250::
2237::
2224::
2211::
2198::
2185::
2170::
2155::
1601:.
1539:^
1518:,
1510:.
1506:.
1490:^
1482:15
1480:.
1447:23
1445:.
1441:.
1421:15
1419:.
1393:^
1377:.
1355:^
1347:14
1345:.
1329:^
1321:40
1319:.
1293:^
1285:12
1283:.
1259:^
1230:.
1195:.
1189:.
1161:^
1140:28
1135:12
1130:14
1092:0
1073:0
1053:20
1048:24
1043:10
1038:10
1024:13
1008:0
988:0
969:4
949:0
930:3
911:0
892:0
873:0
854:0
835:0
816:0
734:.
687:.
637:.
572:,
518:.
460:,
351:.
311:,
307:,
285:—
255:—
1699:e
1692:t
1685:v
1644:.
1612:.
1587:.
1562:.
1533:.
1512:6
1504:"
1457:.
1453::
1387:.
1250:"
1240:.
1216:.
1203::
1187:"
1183:"
1118:0
1113:4
1108:2
1103:4
1089:2
1086:1
1083:0
1070:2
1067:1
1064:4
1021:2
1018:1
1005:0
1002:0
999:1
985:0
982:3
979:2
966:2
963:0
960:1
946:1
943:0
940:0
927:3
924:1
921:1
908:3
905:2
902:1
889:0
886:0
883:1
870:0
867:1
864:1
851:1
848:0
845:0
832:0
829:1
826:1
813:2
810:0
807:0
627:(
534:.
81:)
64:)
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