31:
55:
197:
species of vampire moths, to piercing the skins of mammals to feed on blood. The subfamily contains some large moths with wingspans longer than 5 cm (2 in).
2039:
2242:
282:
structure of this group was essentially unresolved, and in many cases it was even doubtful whether the genera were indeed correctly placed in this subfamily.
2043:
2229:
2070:
233:, previously included in the Calpinae, were found to be distantly related and were reclassified into a separate subfamily as the
1939:
213:, but the redefinition of that family has reclassified many of that family's subfamilies, including Calpinae, into the family
2234:
2141:
2195:
2247:
2288:
2208:
54:
274:
The status of the former composition of the
Calpinae was somewhat disputed; it was sometimes merged into the
184:
141:
2094:
1940:"Blood-feeding and Fruit-feeding Moths of Calpini Tribe (Noctuidae), Their Phylogeny and Classification"
30:
1262:
2020:
209:
studies have greatly revised this subfamily. The subfamily was previously classified within the
150:
49:
1967:"Annotated check list of the Noctuoidea (Insecta, Lepidoptera) of North America north of Mexico"
2283:
2255:
2177:
2260:
2010:
1974:
1540:
864:
234:
1858:
1787:
1449:
1040:
815:
372:
302:
230:
193:
191:
adapted to piercing the skins of fruit to feed on juice, and in the case of the several
2203:
2190:
2132:
1865:
1307:
977:
850:
765:
539:
456:
187:
in 1840. This subfamily includes many species of moths that have a pointed and barbed
2277:
2015:
1998:
1943:
1822:
1717:
1681:
1611:
1576:
1491:
1105:
1033:
962:
836:
681:
486:
218:
2024:
2182:
1966:
1773:
1710:
1688:
1583:
1569:
1300:
1248:
1147:
1091:
1084:
1061:
695:
525:
406:
389:
246:
206:
2216:
2146:
2126:
1908:
1872:
1850:
1724:
1703:
1667:
1632:
1604:
1498:
1435:
1386:
1364:
1255:
1098:
1076:
1069:
1047:
991:
941:
808:
801:
702:
663:
553:
476:
446:
362:
275:
222:
106:
2117:
1801:
1731:
1660:
1653:
1597:
1547:
1533:
1526:
1456:
1371:
1350:
1343:
1279:
1234:
1212:
1198:
1161:
1133:
1112:
1054:
984:
934:
723:
709:
658:
581:
574:
546:
436:
426:
342:
263:
258:
116:
217:. The Calpinae are most closely related to a clade including the subfamilies
1843:
1780:
1759:
1745:
1738:
1590:
1561:
1512:
1505:
1477:
1379:
1357:
1328:
1227:
1154:
1140:
998:
955:
927:
920:
913:
899:
779:
688:
667:
616:
332:
292:
279:
210:
188:
172:
86:
66:
1979:
2164:
2156:
2111:
1829:
1815:
1808:
1695:
1618:
1554:
1519:
1484:
1470:
1428:
1321:
1293:
1241:
1190:
1126:
1026:
1005:
948:
885:
857:
822:
794:
772:
737:
674:
644:
602:
312:
214:
180:
126:
2169:
1915:
1901:
1879:
1794:
1766:
1646:
1625:
1463:
1442:
1336:
1314:
1286:
1219:
1175:
1119:
970:
892:
829:
758:
751:
730:
651:
637:
609:
532:
518:
416:
352:
322:
253:
226:
2221:
2040:"World Checklist of Tribe Calpini (Lepidoptera: Noctuidae: Calpinae)"
1836:
1674:
1639:
1414:
1407:
1400:
1393:
1272:
1205:
1182:
1168:
1019:
1012:
871:
786:
744:
623:
595:
567:
496:
466:
96:
76:
42:
2088:
1752:
1421:
906:
878:
843:
716:
630:
588:
560:
1894:
1887:
176:
2092:
1999:"Molecular phylogenetics of Erebidae (Lepidoptera, Noctuoidea)"
278:. Most of the calpine genera were not further classified. The
1965:
Lafontaine, Donald; Schmidt, Christian (March 19, 2010).
509:(incomplete list); includes taxa sometimes separated in
2101:
2042:. University of Nebraska, Lincoln. Archived from
225:, which are also among the Erebidae. The tribes
8:
2089:
29:
20:
2014:
1978:
1938:Florida Museum of Natural History (ed.).
1992:
1990:
1930:
2075:Lepidoptera and Some Other Life Forms
2038:J.M. Zaspel and M.A. Branham (2008).
7:
2209:9b35ddf6-cb23-4515-bae1-c0806e3dd6f0
1997:Zahiri, Reza; et al. (2011).
14:
2069:Savela, Markku (August 2, 2019).
2016:10.1111/j.1365-3113.2011.00607.x
507:Genera provisionally placed here
53:
245:The Calpinae consist of three
1:
1267:Hacker & Saldaitis, 2010
367:Boisduval & Guenée, 1852
357:Boisduval & Guenée, 1852
2305:
2071:"Calpinae Boisduval, 1840"
1919:(Catocalinae: Ophiusina?)
1883:(Catocalinae: Ophiusina?)
1854:(Catocalinae: Ophiusina?)
1699:(Catocalinae: Ophiusina?)
1565:(Catocalinae: Ophiusina?)
1375:(Catocalinae: Ophiusina?)
1332:(Catocalinae: Ophiusina?)
1223:(Catocalinae: Ophiusina?)
1194:(Catocalinae: Ophiusina?)
1186:(Catocalinae: Ophiusina?)
1080:(Catocalinae: Ophiusina?)
1065:(Catocalinae: Ophiusina?)
790:(Catocalinae: Ophiusina?)
156:
149:
50:Scientific classification
48:
37:
28:
23:
185:Jean Baptiste Boisduval
1980:10.3897/zookeys.40.414
2003:Systematic Entomology
2204:Fauna Europaea (new)
2046:on December 20, 2014
39:Cyclopis caecutiens
307:Ochsenheimer, 1816
16:Subfamily of moths
2271:
2270:
2256:Open Tree of Life
2095:Taxon identifiers
1268:
502:
492:
482:
472:
462:
452:
442:
432:
422:
412:
383:Tribe Gonopterini
378:
368:
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348:
338:
328:
318:
308:
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270:Previous taxonomy
165:
164:
145:
2296:
2289:Moth subfamilies
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2263:
2251:
2250:
2238:
2237:
2225:
2224:
2212:
2211:
2199:
2198:
2186:
2185:
2173:
2172:
2160:
2159:
2150:
2149:
2137:
2136:
2135:
2122:
2121:
2120:
2090:
2085:
2083:
2081:
2056:
2055:
2053:
2051:
2035:
2029:
2028:
2018:
1994:
1985:
1984:
1982:
1962:
1956:
1955:
1953:
1951:
1946:on 11 April 2015
1942:. Archived from
1935:
1266:
500:
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450:
440:
430:
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376:
366:
356:
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235:Scoliopteryginae
140:
58:
57:
33:
21:
2304:
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2299:
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2110:
2097:
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2068:
2065:
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2049:
2047:
2037:
2036:
2032:
1996:
1995:
1988:
1964:
1963:
1959:
1949:
1947:
1937:
1936:
1932:
1927:
1922:
1859:Trigonochrostia
1788:Thoracolophotos
272:
243:
231:Scoliopterygini
203:
139:
52:
17:
12:
11:
5:
2302:
2300:
2292:
2291:
2286:
2276:
2275:
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2266:
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2252:
2239:
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2200:
2191:Fauna Europaea
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978:Eubryopterella
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966:(Catocalinae?)
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411:Möschler, 1890
399:incertae sedis
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317:Billberg, 1820
309:
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179:in the family
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2119:
2113:
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1718:Sphingomorpha
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1612:Pseudogyrtona
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1106:Hirsutopalpis
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682:Brachycyttara
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219:Eulepidotinae
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183:described by
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2102:
2080:November 21,
2078:. Retrieved
2074:
2048:. Retrieved
2044:the original
2033:
2006:
2002:
1970:
1960:
1948:. Retrieved
1944:the original
1933:
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1711:Spectrophysa
1709:
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1689:Schistorhynx
1687:
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1666:
1659:
1652:
1645:
1638:
1631:
1624:
1617:
1610:
1603:
1596:
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1584:Prolyncestis
1582:
1575:
1570:Polydesmiola
1568:
1560:
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1263:Lyncestoides
1261:
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1249:Litoprosopus
1247:
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1226:
1218:
1211:
1204:
1197:
1189:
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1167:
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1148:Hyposemansis
1146:
1139:
1132:
1125:
1118:
1111:
1104:
1097:
1092:Heterormista
1090:
1085:Hemeroblemma
1083:
1075:
1068:
1060:
1053:
1046:
1039:
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1025:
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531:
526:Acanthoprora
524:
517:
510:
506:
505:
501:Walker, 1862
495:
485:
475:
471:Walker, 1863
465:
461:Strand, 1911
455:
445:
441:Turner, 1908
435:
431:Turner, 1925
425:
415:
407:Cecharismena
405:
398:
396:
395:
390:Scoliopteryx
388:
382:
381:
377:Walker, 1867
371:
361:
351:
347:Walker, 1858
341:
337:Hübner, 1818
331:
327:Walker, 1858
321:
311:
301:
297:Krüger, 1939
291:
285:
284:
280:phylogenetic
273:
247:monophyletic
244:
207:phylogenetic
204:
192:
168:
166:
136:
38:
18:
2217:iNaturalist
2127:Wikispecies
2009:: 102–124.
1909:Xanthanomis
1873:Tropidtamba
1851:Toxonprucha
1725:Stenocarsia
1704:Selenoperas
1668:Rhynchodina
1605:Pseudorgyia
1499:Parachabora
1436:Orthozancla
1387:Neostichtis
1365:Metaphoenia
1256:Lophozancla
1099:Heterospila
1077:Heteranassa
1070:Hemipsectra
1048:Goniocarsia
992:Eulepidotis
942:Episparonia
809:Cultripalpa
802:Crypsiprora
703:Brevipecten
664:Catocalinae
554:Anachrostis
511:Ophiderinae
481:Grote, 1872
477:Phyprosopus
451:Perty, 1833
447:Goniapteryx
421:Moore, 1881
363:Plusiodonta
276:Catocalinae
223:Hypocalinae
159:Ophiderinae
133:Subfamily:
107:Lepidoptera
2278:Categories
1973:(40): 26.
1925:References
1802:Thyrostipa
1732:Stenoprora
1661:Rhosologia
1654:Rhesalides
1598:Prorocopis
1548:Phytometra
1534:Pilipectus
1527:Perinaenia
1457:Oxygonitis
1372:Mimophisma
1351:Metalectra
1344:Mesosciera
1280:Malatrogia
1235:Lineopalpa
1213:Leptotroga
1199:Latebraria
1162:Hypsoropha
1134:Hyperlopha
1113:Hopetounia
1055:Gracillina
985:Eudesmeola
935:Episparina
724:Calymniops
710:Brontypena
659:Bendisodes
582:Anticarsia
575:Antiblemma
547:Amphigonia
491:Dyar, 1921
437:Epicyrtica
427:Euryschema
343:Graphigona
264:Phyllodini
259:Ophiderini
117:Noctuoidea
87:Arthropoda
1844:Tolpiodes
1781:Thiacidas
1760:Sypnoides
1746:Syllectra
1739:Sundwarda
1591:Propatria
1562:Polydesma
1541:Phyllodes
1513:Parolulis
1506:Parapadna
1478:Pangrapta
1380:Neogabara
1358:Metallata
1329:Melipotis
1228:Lignicida
1155:Hypospila
1141:Hypoprora
999:Eurythmus
956:Eporectis
928:Episparis
921:Epidromia
914:Ephyrodes
900:Egnasides
865:Diphthera
780:Corcobara
689:Brachyona
668:Ophiusina
617:Ascalapha
333:Gonodonta
293:Africalpe
211:Noctuidae
189:proboscis
173:subfamily
142:Boisduval
73:Kingdom:
67:Eukaryota
24:Calpinae
2284:Calpinae
2165:BugGuide
2154:BioLib:
2147:Calpinae
2133:Calpinae
2118:Q2934400
2112:Wikidata
2103:Calpinae
2025:84249695
1950:21 April
1830:Tiruvaca
1816:Tinolius
1809:Thysania
1696:Selenisa
1619:Ramadasa
1555:Pleurona
1520:Perciana
1485:Panopoda
1471:Palpidia
1450:Ossonoba
1429:Orodesma
1322:Mecodina
1294:Marapana
1242:Litocala
1191:Kakopoda
1127:Hypocala
1041:Gloriana
1027:Forsebia
1006:Facidina
949:Epitausa
886:Eclipsea
858:Diatenes
823:Cyclopis
816:Cyclodes
795:Crithote
773:Conosema
738:Chilkasa
675:Boryzops
645:Belciana
603:Arthisma
373:Tetrisia
313:Eudocima
303:Calyptra
249:tribes.
215:Erebidae
201:Taxonomy
194:Calyptra
181:Erebidae
169:Calpinae
151:Synonyms
137:Calpinae
127:Erebidae
123:Family:
83:Phylum:
77:Animalia
63:Domain:
2050:July 8,
1971:ZooKeys
1916:Zaleops
1902:Vestura
1880:Tyrissa
1866:Trisula
1795:Throana
1767:Talmala
1647:Rhesala
1626:Raparna
1464:Oxyodes
1443:Ortopla
1337:Meranda
1315:Matella
1308:Massala
1287:Manbuta
1220:Lesmone
1176:Isogona
1120:Hulodes
971:Ericeia
893:Egnasia
851:Diascia
830:Daddala
766:Condate
759:Concana
752:Cissusa
731:Cephena
652:Bematha
638:Baniana
610:Arugisa
540:Alabama
533:Agamana
519:Abriesa
417:Culasta
397:Genera
353:Oraesia
323:Ferenta
254:Calpini
227:Anomini
205:Recent
103:Order:
97:Insecta
93:Class:
2261:455581
2248:415138
2235:694005
2222:122490
2196:446803
2023:
1837:Tolpia
1823:Tipasa
1682:Savara
1675:Saroba
1640:Rhapsa
1577:Praxis
1492:Panula
1415:Olyssa
1408:Olulis
1401:Oglasa
1394:Obrima
1273:Maguda
1206:Ledaea
1183:Itomia
1169:Ianius
1034:Gabara
1020:Fodina
1013:Falana
963:Erebus
872:Dunira
837:Dahlia
787:Coxina
745:Chodda
624:Avitta
596:Armana
568:Anomis
497:Radara
467:Pharga
241:Tribes
171:are a
144:, 1840
43:Brazil
2183:93MSD
2170:12331
2157:53309
2021:S2CID
1774:Tamba
1753:Sypna
1422:Ophyx
1301:Masca
1062:Helia
907:Egone
879:Dyops
844:Daona
717:Bulia
696:Brana
631:Azeta
589:Arcte
561:Anoba
177:moths
41:from
2243:NCBI
2230:ITIS
2082:2019
2052:2010
1952:2015
1895:Veia
1888:Ugia
1633:Rema
229:and
221:and
167:The
2178:CoL
2142:AFD
2011:doi
1975:doi
175:of
2280::
2258::
2245::
2232::
2219::
2206::
2193::
2180::
2167::
2144::
2129::
2114::
2073:.
2019:.
2007:37
2005:.
2001:.
1989:^
1969:.
670:?)
666::
237:.
2084:.
2054:.
2027:.
2013::
1983:.
1977::
1954:.
662:(
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