249:
33:
1278:
56:
1320:
351:. The worker ants, sisters of these queens, may transport the queens several meters from their natal nest and drop them off at another, alien nest to promote outbreeding with wingless stationary males in a process somewhat analogous to third party matchmaking in humans. After mating during the winter, the sexual females may depart in the spring and find their own colonies.
1334:
1281:
This article incorporates text from a scholarly publication published under a copyright license that allows anyone to reuse, revise, remix and redistribute the materials in any form for any purpose:
302:, but a few of which occur in the temperate zone. Some species are also found widely separated in North America and the Pacific Islands, as a result of human introduction.
1516:
1555:
1490:
1529:
1285:"The Spatial Distribution of mtDNA and Phylogeographic Analysis of the Ant Cardiocondyla kagutsuchi (Hymenoptera: Formicidae) in Japan"
1138:"The Spatial Distribution of mtDNA and Phylogeographic Analysis of the Ant Cardiocondyla kagutsuchi (Hymenoptera: Formicidae) in Japan"
1629:
1100:
248:
1183:
1179:, with comments on their affinities, and the first descriptions of Australian species. (Hymenoptera: Formicidae: Myrmicinae)"
294:
Approximately 70 species are currently recognized as belonging to this genus, most of which are distributed in the Old World
1534:
32:
1542:
55:
1412:
1399:
1560:
1469:
1215:
Vidal, Mathilde; Königseder, Florian; Giehr, Julia; Schrempf, Alexandra; Lucas, Christophe; Heinze, Jürgen (2021).
1482:
632:
892:
572:
1338:
1012:
902:
492:
41:
912:
862:
752:
702:
592:
562:
552:
482:
472:
432:
402:
392:
382:
982:
952:
922:
872:
852:
832:
792:
732:
612:
602:
582:
542:
422:
1002:
972:
962:
932:
842:
822:
802:
782:
772:
532:
442:
412:
362:
992:
942:
762:
712:
682:
662:
522:
502:
372:
1352:
1032:
1022:
652:
642:
622:
347:
worker ants transport young queen ants to alien nests to promote outbreeding. This allows avoidance of
311:
742:
692:
1042:
882:
672:
452:
348:
174:
1417:
512:
1596:
1277:
1570:
1451:
812:
462:
315:
253:
1624:
1619:
1390:
1324:
722:
200:
50:
1578:
1521:
1438:
1246:
1583:
1296:
1236:
1228:
1192:
1149:
158:
137:
1301:
1284:
1283:
Okita, I.; Murase, K.; Sato, T.; Kato, K.; Hosoda, A.; Terayama, M.; Masuko, K. (2013).
1154:
1137:
1136:
Okita, I.; Murase, K.; Sato, T.; Kato, K.; Hosoda, A.; Terayama, M.; Masuko, K. (2013).
1477:
1464:
1264:
Emery, C. (1869). "Enumerazione dei
Formicidi che rinvengonsi nei contorni di Napoli".
1241:
1216:
1613:
1404:
1443:
1067:
326:
186:
166:
1547:
1503:
1384:
332:
320:
154:
107:
1375:
1319:
1232:
299:
283:
127:
1217:"Worker ants promote outbreeding by transporting young queens to alien nests"
280:
269:
87:
67:
1250:
1333:
1425:
1369:
1197:
1170:
257:
1430:
1495:
295:
314:, with both winged and wingless forms. These males differ not only in
1508:
1112:
97:
77:
1346:
1071:
1591:
272:
247:
1350:
318:, but also in reproductive tactics. Closely related genera are
1456:
276:
117:
1307:
Please check the source for the exact licensing terms.
1359:
310:Several species of this genus have a striking male
1131:
1129:
1266:Annali dell'Accademia degli Aspiranti Naturalisti
8:
1210:
1208:
1347:
31:
20:
1300:
1240:
1196:
1153:
1059:
7:
1597:1FAE6D10-F408-5572-4314-09A96CE429F2
1571:9a07f72f-6925-42e0-8233-2145dd36d045
1483:75ba1812-ee71-4220-99b0-6b40121c4158
1302:10.13102/sociobiology.v60i2.129-134
1155:10.13102/sociobiology.v60i2.129-134
14:
1332:
1318:
1276:
817:Seifert & Frohschammer, 2013
54:
947:Ghosh, Sheela & Kundu, 2005
1027:Collingwood & Agosti, 1996
487:Collingwood & Agosti, 1996
1:
193:
1646:
1233:10.1038/s42003-021-02016-1
1169:Taylor, Robert W. (1991).
893:Cardiocondyla stambuloffii
573:Cardiocondyla koshewnikovi
1630:Taxa named by Carlo Emery
1171:"Notes on the ant genera
1013:Cardiocondyla wroughtonii
903:Cardiocondyla strigifrons
633:Cardiocondyla mauritanica
493:Cardiocondyla gallilaeica
206:
199:
192:
185:
172:
165:
51:Scientific classification
49:
39:
30:
23:
913:Cardiocondyla tenuifrons
863:Cardiocondyla shagrinata
753:Cardiocondyla opistopsis
703:Cardiocondyla nigrocerea
593:Cardiocondyla littoralis
563:Cardiocondyla kagutsuchi
553:Cardiocondyla jacquemini
483:Cardiocondyla gallagheri
473:Cardiocondyla fajumensis
433:Cardiocondyla carbonaria
403:Cardiocondyla breviscapa
393:Cardiocondyla brachyceps
383:Cardiocondyla bicoronata
983:Cardiocondyla venustula
953:Cardiocondyla tjibodana
923:Cardiocondyla thoracica
873:Cardiocondyla shuckardi
853:Cardiocondyla semirubra
833:Cardiocondyla sahlbergi
793:Cardiocondyla parvinoda
733:Cardiocondyla obscurior
613:Cardiocondyla longinoda
603:Cardiocondyla longiceps
583:Cardiocondyla kushanica
543:Cardiocondyla israelica
507:Kuznetsov-Ugamsky, 1927
423:Cardiocondyla bulgarica
1221:Communications Biology
1003:Cardiocondyla wheeleri
973:Cardiocondyla unicalis
963:Cardiocondyla ulianini
933:Cardiocondyla tibetana
843:Cardiocondyla sekhemka
823:Cardiocondyla rugulosa
803:Cardiocondyla persiana
783:Cardiocondyla paranuda
773:Cardiocondyla paradoxa
653:Cardiocondyla minutior
533:Cardiocondyla insutura
443:Cardiocondyla cristata
413:Cardiocondyla britteni
363:Cardiocondyla atalanta
260:
1043:Cardiocondyla zoserka
993:Cardiocondyla weserka
943:Cardiocondyla tiwarii
763:Cardiocondyla papuana
713:Cardiocondyla nivalis
683:Cardiocondyla neferka
663:Cardiocondyla monardi
523:Cardiocondyla humilis
503:Cardiocondyla gibbosa
453:Cardiocondyla elegans
373:Cardiocondyla batesii
349:inbreeding depression
345:Cardiocondyla elegans
251:
175:Cardiocondyla elegans
1478:Fauna Europaea (new)
1343:at Wikimedia Commons
1033:Cardiocondyla yoruba
1023:Cardiocondyla yemeni
813:Cardiocondyla pirata
643:Cardiocondyla melana
623:Cardiocondyla luciae
463:Cardiocondyla emeryi
254:Cardiocondyla pirata
743:Cardiocondyla opaca
693:Cardiocondyla nigra
1198:10.1155/1990/29514
883:Cardiocondyla sima
723:Cardiocondyla nuda
673:Cardiocondyla nana
261:
1607:
1606:
1579:Open Tree of Life
1353:Taxon identifiers
1337:Media related to
1048:
1038:
1028:
1018:
1008:
998:
988:
978:
968:
958:
948:
938:
928:
918:
908:
898:
888:
878:
868:
858:
848:
838:
828:
818:
808:
798:
788:
778:
768:
758:
748:
738:
728:
718:
708:
698:
688:
678:
668:
658:
648:
638:
628:
618:
608:
598:
588:
578:
568:
558:
548:
538:
528:
518:
513:Cardiocondyla goa
508:
498:
488:
478:
468:
458:
448:
438:
428:
418:
408:
398:
388:
378:
368:
246:
245:
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234:
227:
220:
213:
161:
1637:
1600:
1599:
1587:
1586:
1574:
1573:
1564:
1563:
1551:
1550:
1548:NHMSYS0000875601
1538:
1537:
1525:
1524:
1512:
1511:
1499:
1498:
1486:
1485:
1473:
1472:
1460:
1459:
1447:
1446:
1434:
1433:
1421:
1420:
1408:
1407:
1395:
1394:
1393:
1380:
1379:
1378:
1348:
1336:
1323:Data related to
1322:
1306:
1304:
1280:
1273:
1255:
1254:
1244:
1212:
1203:
1202:
1200:
1191:(3–4): 281–298.
1166:
1160:
1159:
1157:
1133:
1124:
1123:
1121:
1119:
1097:
1091:
1090:
1088:
1086:
1064:
1046:
1036:
1026:
1016:
1006:
996:
986:
976:
966:
956:
946:
936:
926:
916:
906:
896:
886:
876:
866:
856:
846:
836:
826:
816:
806:
796:
786:
776:
767:(Reiskind, 1965)
766:
756:
746:
736:
726:
716:
706:
696:
686:
676:
666:
656:
646:
636:
626:
616:
606:
596:
586:
576:
566:
556:
546:
536:
526:
516:
506:
496:
486:
476:
466:
456:
447:(Santschi, 1912)
446:
436:
426:
416:
406:
396:
386:
376:
366:
239:
232:
225:
218:
211:
153:
59:
58:
35:
21:
1645:
1644:
1640:
1639:
1638:
1636:
1635:
1634:
1610:
1609:
1608:
1603:
1595:
1590:
1582:
1577:
1569:
1567:
1559:
1554:
1546:
1541:
1533:
1528:
1520:
1515:
1507:
1502:
1494:
1489:
1481:
1476:
1468:
1463:
1455:
1450:
1442:
1437:
1429:
1424:
1416:
1411:
1403:
1398:
1389:
1388:
1383:
1374:
1373:
1368:
1355:
1315:
1310:
1282:
1263:
1259:
1258:
1214:
1213:
1206:
1168:
1167:
1163:
1135:
1134:
1127:
1117:
1115:
1099:
1098:
1094:
1084:
1082:
1066:
1065:
1061:
1056:
1051:
1007:Viehmeyer, 1914
957:Karavaiev, 1935
907:Viehmeyer, 1922
707:Karavaiev, 1935
357:
342:
308:
292:
235:
228:
221:
214:
181:
178:
152:
138:Crematogastrini
53:
17:
12:
11:
5:
1643:
1641:
1633:
1632:
1627:
1622:
1612:
1611:
1605:
1604:
1602:
1601:
1588:
1575:
1565:
1552:
1539:
1526:
1513:
1500:
1487:
1474:
1465:Fauna Europaea
1461:
1448:
1435:
1422:
1409:
1396:
1381:
1365:
1363:
1357:
1356:
1351:
1345:
1344:
1330:
1329:at Wikispecies
1314:
1313:External links
1311:
1309:
1308:
1295:(2): 129–134.
1274:
1260:
1257:
1256:
1204:
1161:
1148:(2): 129–134.
1125:
1092:
1058:
1057:
1055:
1052:
1050:
1049:
1039:
1029:
1019:
1009:
999:
989:
979:
969:
959:
949:
939:
929:
919:
909:
899:
889:
879:
869:
859:
849:
839:
829:
819:
809:
799:
789:
779:
769:
759:
749:
739:
729:
719:
709:
699:
689:
679:
669:
667:Santschi, 1930
659:
649:
639:
629:
619:
609:
599:
589:
587:Pisarski, 1967
579:
569:
567:Terayama, 1999
559:
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226:Santschi, 1930
212:Santschi, 1930
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65:
61:
60:
47:
46:
45:ergatoid male
42:C. wroughtonii
37:
36:
28:
27:
15:
13:
10:
9:
6:
4:
3:
2:
1642:
1631:
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1432:
1427:
1423:
1419:
1414:
1410:
1406:
1405:Cardiocondyla
1401:
1397:
1392:
1391:Cardiocondyla
1386:
1382:
1377:
1371:
1367:
1366:
1364:
1362:
1361:Cardiocondyla
1358:
1354:
1349:
1342:
1341:
1340:Cardiocondyla
1335:
1331:
1328:
1327:
1326:Cardiocondyla
1321:
1317:
1316:
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1303:
1298:
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1286:
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1165:
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1151:
1147:
1143:
1139:
1132:
1130:
1126:
1114:
1110:
1106:
1104:
1103:Cardiocondyla
1096:
1093:
1081:
1077:
1075:
1074:Cardiocondyla
1069:
1063:
1060:
1053:
1045:
1044:
1040:
1035:
1034:
1030:
1025:
1024:
1020:
1017:(Forel, 1890)
1015:
1014:
1010:
1005:
1004:
1000:
995:
994:
990:
987:Wheeler, 1908
985:
984:
980:
977:Seifert, 2003
975:
974:
970:
965:
964:
960:
955:
954:
950:
945:
944:
940:
937:Seifert, 2003
935:
934:
930:
927:(Smith, 1859)
925:
924:
920:
917:Seifert, 2003
915:
914:
910:
905:
904:
900:
895:
894:
890:
887:Wheeler, 1935
885:
884:
880:
875:
874:
870:
867:Seifert, 2003
865:
864:
860:
857:Seifert, 2003
855:
854:
850:
845:
844:
840:
835:
834:
830:
827:Seifert, 2003
825:
824:
820:
815:
814:
810:
807:Seifert, 2003
805:
804:
800:
795:
794:
790:
787:Seifert, 2003
785:
784:
780:
775:
774:
770:
765:
764:
760:
757:Seifert, 2003
755:
754:
750:
747:Seifert, 2003
745:
744:
740:
737:Wheeler, 1929
735:
734:
730:
725:
724:
720:
715:
714:
710:
705:
704:
700:
695:
694:
690:
685:
684:
680:
677:Seifert, 2003
675:
674:
670:
665:
664:
660:
655:
654:
650:
647:Seifert, 2003
645:
644:
640:
635:
634:
630:
625:
624:
620:
615:
614:
610:
607:Seifert, 2003
605:
604:
600:
597:Seifert, 2003
595:
594:
590:
585:
584:
580:
575:
574:
570:
565:
564:
560:
557:Bernard, 1953
555:
554:
550:
547:Seifert, 2003
545:
544:
540:
535:
534:
530:
527:(Smith, 1858)
525:
524:
520:
517:Seifert, 2003
515:
514:
510:
505:
504:
500:
497:Seifert, 2003
495:
494:
490:
485:
484:
480:
475:
474:
470:
465:
464:
460:
455:
454:
450:
445:
444:
440:
435:
434:
430:
425:
424:
420:
417:Crawley, 1920
415:
414:
410:
407:Seifert, 2003
405:
404:
400:
397:Seifert, 2003
395:
394:
390:
387:Seifert, 2003
385:
384:
380:
375:
374:
370:
365:
364:
360:
359:
354:
352:
350:
346:
339:
337:
335:
334:
329:
328:
323:
322:
317:
313:
305:
303:
301:
297:
289:
287:
285:
282:
278:
274:
271:
267:
266:
265:Cardiocondyla
259:
256:
255:
250:
242:
238:
233:Wheeler, 1935
231:
224:
217:
210:
205:
202:
198:
195:
191:
188:
184:
177:
176:
171:
168:
164:
160:
156:
151:
150:
149:Cardiocondyla
146:
143:
142:
139:
136:
133:
132:
129:
126:
123:
122:
119:
116:
113:
112:
109:
106:
103:
102:
99:
96:
93:
92:
89:
86:
83:
82:
79:
76:
73:
72:
69:
66:
63:
62:
57:
52:
48:
44:
43:
38:
34:
29:
26:
25:Cardiocondyla
22:
19:
16:Genus of ants
1360:
1339:
1325:
1292:
1289:Sociobiology
1288:
1269:
1265:
1224:
1220:
1188:
1182:
1177:Willowsiella
1176:
1172:
1164:
1145:
1142:Sociobiology
1141:
1116:. Retrieved
1108:
1102:
1095:
1083:. Retrieved
1079:
1073:
1062:
1047:Bolton, 1982
1041:
1037:Rigato, 2002
1031:
1021:
1011:
1001:
997:Bolton, 1982
991:
981:
971:
961:
951:
941:
931:
921:
911:
901:
891:
881:
871:
861:
851:
847:Bolton, 1982
841:
831:
821:
811:
801:
791:
781:
771:
761:
751:
741:
731:
727:(Mayr, 1866)
721:
711:
701:
691:
687:Bolton, 1982
681:
671:
661:
651:
641:
631:
627:Rigato, 2002
621:
617:Rigato, 2002
611:
601:
591:
581:
577:Ruzsky, 1902
571:
561:
551:
541:
531:
521:
511:
501:
491:
481:
471:
461:
451:
441:
431:
421:
411:
401:
391:
381:
371:
361:
344:
343:
331:
327:Stereomyrmex
325:
319:
312:polymorphism
309:
293:
290:Distribution
264:
263:
262:
252:
236:
229:
222:
215:
208:
207:
173:
167:Type species
148:
147:
40:
24:
18:
1504:iNaturalist
1385:Wikispecies
1173:Romblonella
967:Emery, 1889
897:Forel, 1892
877:Forel, 1891
837:Forel, 1913
797:Forel, 1902
777:Emery, 1897
697:Forel, 1905
657:Forel, 1899
637:Forel, 1890
477:Forel, 1913
467:Forel, 1881
457:Emery, 1869
437:Forel, 1907
427:Forel, 1892
377:Forel, 1894
367:Forel, 1915
340:Outbreeding
333:Romblonella
321:Leptothorax
306:Description
240:Emery, 1917
230:Prosopidris
219:Forel, 1890
180:Emery, 1869
124:Subfamily:
108:Hymenoptera
1625:Ant genera
1620:Myrmicinae
1614:Categories
1272:(2): 1–26.
1227:(1): 515.
1118:11 October
1109:antweb.org
1068:Bolton, B.
1054:References
717:Mann, 1919
537:Zhou, 2001
316:morphology
300:subtropics
284:Myrmicinae
194:69 species
128:Myrmicinae
118:Formicidae
88:Arthropoda
281:subfamily
270:Old World
237:Xenometra
187:Diversity
74:Kingdom:
68:Eukaryota
1426:BugGuide
1376:Q1941598
1370:Wikidata
1251:33941829
1101:"Genus:
1070:(2014).
258:paratype
201:Synonyms
114:Family:
84:Phylum:
78:Animalia
64:Domain:
1584:1021159
1522:1092703
1496:1317100
1242:8093424
1085:20 July
355:Species
296:tropics
279:in the
223:Loncyda
216:Emeryia
209:Dyclona
144:Genus:
134:Tribe:
104:Order:
98:Insecta
94:Class:
1568:NZOR:
1561:235541
1535:574042
1431:515563
1249:
1239:
1184:Psyche
1113:AntWeb
1080:AntCat
268:is an
1592:Plazi
1517:IRMNG
1509:68967
1470:81086
1457:49197
1444:62KFW
273:genus
155:Emery
1556:NCBI
1530:ITIS
1491:GBIF
1418:6427
1413:BOLD
1247:PMID
1175:and
1120:2013
1087:2014
330:and
298:and
277:ants
159:1869
1543:NBN
1452:EoL
1439:CoL
1400:AFD
1297:doi
1237:PMC
1229:doi
1193:doi
1150:doi
275:of
1616::
1594::
1581::
1558::
1545::
1532::
1519::
1506::
1493::
1480::
1467::
1454::
1441::
1428::
1415::
1402::
1387::
1372::
1293:60
1291:.
1287:.
1268:.
1245:.
1235:.
1223:.
1219:.
1207:^
1189:97
1187:.
1181:.
1146:60
1144:.
1140:.
1128:^
1111:.
1107:.
1078:.
336:.
324:,
286:.
157:,
1305:.
1299::
1270:2
1253:.
1231::
1225:4
1201:.
1195::
1158:.
1152::
1122:.
1105:"
1089:.
1076:"
1072:"
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