126:
394:
is poorly preserved, it has been estimated to be 7.5 mm (0.30 in) long and 5.5 mm (0.22 in) high, and is known to cover the head and front part of the thorax, whereas the last four thoracic segments were left exposed. The segments of the thorax are short but tall, each measuring
411:
The hind margin of each thoracic and abdominal segment is almost vertical, and possesses ridges with five or six short, backward-pointing spines on each side. These spines are largest near the middle of the crustacean's back, decreasing in size down its sides. The hind segments also have slightly
399:
is split into six segments, which decrease in size along the animal's length: the first abdominal segment is 2.5 mm (0.098 in) long and 4 mm (0.16 in) tall, whereas the last is only 1.3 mm (0.051 in) long and 2 mm (0.079 in) tall. The
386:
which would have been used to capture prey. The raptorial appendages of the frontmost pair are each 1.5 mm (0.059 in) wide at the base, larger than those of the other three pairs (which are each 1 mm (0.039 in) wide at the base).
366:, with an arched body reaching a total length of around 30 mm (1.2 in). The animal's body was laterally compressed, being flattened from side to side and appearing narrow when viewed from above. Like other
313:
343:
honors D. Chabard, who was curator of the Muséum d'histoire naturelle d'Autun and therefore in charge of the
Montceau-les-Mines fossil collection. The
1657:
1616:
1545:
Smith, C.P.A.; Aubier, P.; Charbonnier, S.; Laville, T.; Olivier, N.; Escarguel, G.; Jenks, J.F.; Bylund, K.G.; Fara, E.; Brayard, A. (2023-03-31).
1672:
1667:
1677:
308:
period. A research program on the fossil content of this site began in 2000, during which the only known specimen was found in a
125:
379:
344:
1417:"New Malacostracan Crustacea from the Carboniferous (Stephanian) Lagerstätte of Montceau-Les-Mines, France"
1584:
340:
500:
485:
231:
1662:
1491:
1428:
1068:
1478:
Haug, Joachim T.; Haug, Carolin; Maas, Andreas; Kutschera, Verena; Waloszek, Dieter (2010-09-21).
1041:
450:, making it the second genus assigned to this family. This placement was first questioned by Haug
1547:"Closing a major gap in mantis shrimp evolution - first fossils of Stomatopoda from the Triassic"
1460:
1452:
476:. However, they acknowledge this result may have been due to the lack of data on the tail fan of
305:
301:
297:
271:
120:
34:
30:
408:
has a short, blade-like base. The rest of the tail fan is not preserved and thus poorly known.
1629:
1564:
1527:
1509:
1480:"Evolution of mantis shrimps (Stomatopoda, Malacostraca) in the light of new Mesozoic fossils"
1444:
1634:
1554:
1517:
1499:
1436:
447:
190:
432:
appendages is larger than the latter pairs, Racheboeuf, Schram and Vidal (2009) recognized
1495:
1432:
1522:
1479:
1229:
957:
930:
849:
832:
816:
464:
438:
335:
specimen, MNHN-SOT 28610, consists of the external mold of a weathered, near-complete
1651:
1546:
1464:
1014:
987:
367:
328:
267:
177:
72:
412:
larger spines than the segments further front. These spines are a feature unique to
1607:
1185:
663:
646:
630:
569:
552:
536:
167:
521:
336:
47:
395:
around 1.5 mm (0.059 in) long and 4 mm (0.16 in) tall. The
1212:
970:
469:
383:
363:
309:
92:
57:
1568:
1559:
1513:
1504:
1448:
1158:
504:
492:
in an unresolved polytomy, and further suggests the incomplete knowledge of
429:
417:
375:
157:
137:
97:
41:
1531:
1601:
701:
612:
473:
391:
351:, references the rows of spines on the margins of the animal's segments.
332:
87:
82:
67:
62:
52:
1456:
1416:
1621:
396:
102:
77:
1440:
378:
maxillipeds (feeding appendages): the second to fifth segments of the
1122:
1102:
789:
769:
728:
494:
405:
401:
275:
147:
1578:
1415:
Racheboeuf, Patrick R.; Schram, Frederick R.; Vidal, Muriel (2009).
317:
263:
1582:
499:
may have also led to this result. The results of their
442:, which also has this feature. The authors thus placed
1208:
1181:
1154:
1118:
1098:
1064:
1037:
1010:
983:
953:
926:
890:
812:
785:
765:
724:
697:
626:
608:
532:
1591:
296:was discovered in the Saint-Louis opencast pit in
1410:
1408:
1406:
1404:
1402:
1400:
404:is elongated and subtriangular in shape, and the
323:In 2009, Racheboeuf, Schram and Vidal described
316:and was deposited in an off-site facility in
8:
1579:
314:National Museum of Natural History, France
20:
1558:
1521:
1503:
1396:
7:
1201:
1174:
1147:
1091:
1084:
1057:
1030:
1003:
976:
946:
919:
883:
824:
758:
751:
744:
717:
638:
601:
594:
544:
527:
517:
480:, which is poorly preserved. Smith
250:Racheboeuf, Schram and Vidal, 2009
210:Racheboeuf, Schram and Vidal, 2009
14:
339:, preserving its left side. The
124:
1658:Carboniferous animals of Europe
428:Because the frontmost pair of
327:as a new genus and species of
284:as the type and only species.
16:Extinct genus of mantis shrimp
1:
1673:Prehistoric crustacean genera
1668:Fossil taxa described in 2009
382:each bear a pair of enlarged
312:. This fossil belongs to the
300:, France, which dates to the
278:. It was named in 2009, with
472:clade to form an unresolved
512:
416:, and are unknown in other
1694:
1678:Prehistoric mantis shrimps
331:based on this fossil. The
1226:
1206:
1199:
1179:
1172:
1152:
1145:
1116:
1096:
1089:
1082:
1062:
1055:
1035:
1028:
1008:
1001:
981:
974:
951:
944:
924:
917:
888:
881:
846:
829:
822:
783:
763:
756:
749:
742:
722:
715:
695:
660:
643:
636:
606:
599:
592:
566:
549:
542:
525:
304:B and C substages of the
237:
230:
121:Scientific classification
119:
23:
1560:10.3140/bull.geosci.1864
1505:10.1186/1471-2148-10-290
1484:BMC Evolutionary Biology
1186:Lysiosquilla nkporoensis
958:Tyrannophontes gigantion
931:Tyrannophontes theridion
833:Gorgonophontes fraiponti
553:Archaeocaris vermiformis
1551:Bulletin of Geosciences
1421:Journal of Paleontology
1069:Archaeosculda phoenicia
702:Bairdops beargulchensis
436:as a close relative of
292:The fossil material of
270:which lived during the
850:Gorgonophontes peleron
570:Archaeocaris graffhami
1213:Nodosculda fisherorum
988:Triassosculda ahyongi
503:are displayed in the
501:phylogenetic analyses
486:phylogenetic analysis
1015:Tyrannosculda laurae
790:Daidal acanthocercus
454:(2010), which found
288:Discovery and naming
1496:2010BMCEE..10..290H
1433:2009JPal...83..624R
1042:Pseudosculda laevis
895:Chabardella spinosa
770:Daidal schoellmanni
484:(2023) conducted a
325:Chabardella spinosa
244:Chabardella spinosa
1373:Archaeostomatopods
1159:Ursquilla yehoachi
664:Perimecturus rapax
647:Perimecturus parki
488:which also placed
306:Late Carboniferous
298:Montceau-les-Mines
272:Late Carboniferous
31:Late Carboniferous
1645:
1644:
1630:Open Tree of Life
1585:Taxon identifiers
1441:10.1666/08-171R.1
1385:
1384:
1376:Palaeostomatopods
1367:
1366:
1358:
1357:
1349:
1348:
1340:
1339:
1331:
1330:
1322:
1321:
1313:
1312:
1304:
1303:
1295:
1294:
1286:
1285:
1277:
1276:
1268:
1267:
1259:
1258:
1250:
1249:
1241:
1240:
1134:
1133:
906:
905:
870:
869:
861:
860:
801:
800:
684:
683:
675:
674:
581:
580:
255:
254:
211:
40:301.5–300.5
1685:
1638:
1637:
1625:
1624:
1612:
1611:
1610:
1580:
1573:
1572:
1562:
1542:
1536:
1535:
1525:
1507:
1475:
1469:
1468:
1412:
1210:
1202:
1183:
1175:
1156:
1148:
1120:
1100:
1092:
1085:
1066:
1058:
1039:
1031:
1012:
1004:
985:
977:
955:
947:
928:
920:
892:
884:
825:
814:
787:
767:
759:
752:
745:
726:
718:
699:
639:
628:
613:Bairdops elegans
610:
602:
595:
545:
534:
528:
518:
513:
448:Gorgonophontidae
420:mantis shrimps.
246:
242:
209:
202:
191:Gorgonophontidae
189:
129:
128:
114:
44:
29:Temporal range:
21:
1693:
1692:
1688:
1687:
1686:
1684:
1683:
1682:
1648:
1647:
1646:
1641:
1633:
1628:
1620:
1615:
1606:
1605:
1600:
1587:
1577:
1576:
1544:
1543:
1539:
1477:
1476:
1472:
1414:
1413:
1398:
1393:
1387:
1381:
1380:
1377:
1374:
1368:
1359:
1350:
1341:
1332:
1323:
1314:
1305:
1296:
1287:
1278:
1269:
1260:
1251:
1242:
1135:
907:
871:
862:
820:
802:
685:
676:
634:
582:
540:
510:
426:
357:
290:
251:
248:
240:
239:
226:
223:C. spinosa
208:
200:
187:
123:
115:
113:
112:
111:
110:
105:
100:
95:
90:
85:
80:
75:
70:
65:
60:
55:
50:
39:
38:
27:
17:
12:
11:
5:
1691:
1689:
1681:
1680:
1675:
1670:
1665:
1660:
1650:
1649:
1643:
1642:
1640:
1639:
1626:
1613:
1597:
1595:
1589:
1588:
1583:
1575:
1574:
1537:
1470:
1427:(4): 624–629.
1395:
1394:
1392:
1389:
1383:
1382:
1379:Pseudosculdids
1378:
1375:
1372:
1371:
1369:
1365:
1364:
1361:
1360:
1356:
1355:
1352:
1351:
1347:
1346:
1343:
1342:
1338:
1337:
1334:
1333:
1329:
1328:
1325:
1324:
1320:
1319:
1316:
1315:
1311:
1310:
1307:
1306:
1302:
1301:
1298:
1297:
1293:
1292:
1289:
1288:
1284:
1283:
1280:
1279:
1275:
1274:
1271:
1270:
1266:
1265:
1262:
1261:
1257:
1256:
1253:
1252:
1248:
1247:
1244:
1243:
1239:
1238:
1235:
1234:
1230:Squilla mantis
1225:
1222:
1221:
1218:
1217:
1205:
1200:
1198:
1195:
1194:
1191:
1190:
1178:
1173:
1171:
1168:
1167:
1164:
1163:
1151:
1146:
1144:
1141:
1140:
1137:
1136:
1132:
1131:
1128:
1127:
1123:Sculda syriaca
1115:
1112:
1111:
1108:
1107:
1103:Sculda pennata
1095:
1090:
1088:
1083:
1081:
1078:
1077:
1074:
1073:
1061:
1056:
1054:
1051:
1050:
1047:
1046:
1034:
1029:
1027:
1024:
1023:
1020:
1019:
1007:
1002:
1000:
997:
996:
993:
992:
980:
975:
973:
967:
966:
963:
962:
950:
945:
943:
940:
939:
936:
935:
923:
918:
916:
913:
912:
909:
908:
904:
903:
900:
899:
887:
882:
880:
877:
876:
873:
872:
868:
867:
864:
863:
859:
858:
855:
854:
845:
842:
841:
838:
837:
828:
823:
821:
817:Gorgonophontes
811:
808:
807:
804:
803:
799:
798:
795:
794:
782:
779:
778:
775:
774:
762:
757:
755:
750:
748:
743:
741:
738:
737:
734:
733:
729:Daidal pattoni
721:
716:
714:
711:
710:
707:
706:
694:
691:
690:
687:
686:
682:
681:
678:
677:
673:
672:
669:
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659:
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651:
642:
637:
635:
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622:
621:
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617:
605:
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588:
587:
584:
583:
579:
578:
575:
574:
565:
562:
561:
558:
557:
548:
543:
541:
531:
526:
524:
516:
465:Tyrannophontes
460:Gorgonophontes
439:Gorgonophontes
425:
424:Classification
422:
368:mantis shrimps
356:
353:
289:
286:
262:is an extinct
253:
252:
249:
235:
234:
228:
227:
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217:
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185:
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117:
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96:
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66:
61:
56:
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28:
15:
13:
10:
9:
6:
4:
3:
2:
1690:
1679:
1676:
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1669:
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1556:
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1548:
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1538:
1533:
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1524:
1519:
1515:
1511:
1506:
1501:
1497:
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1471:
1466:
1462:
1458:
1454:
1450:
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1442:
1438:
1434:
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1426:
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1411:
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1405:
1403:
1401:
1397:
1390:
1388:
1370:
1363:
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1327:
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1318:
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1309:
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1245:
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1233:
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1126:
1125:
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1114:
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1093:
1087:
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1080:
1079:
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1033:
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1026:
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1022:
1021:
1018:
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1016:
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1005:
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991:
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989:
979:
978:
972:
969:
968:
965:
964:
961:
960:
959:
949:
948:
942:
941:
938:
937:
934:
933:
932:
922:
921:
915:
914:
911:
910:
902:
901:
898:
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896:
886:
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879:
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866:
865:
857:
856:
853:
852:
851:
844:
843:
840:
839:
836:
835:
834:
827:
826:
819:
818:
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805:
797:
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781:
780:
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771:
761:
760:
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753:
747:
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740:
739:
736:
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732:
731:
730:
720:
719:
713:
712:
709:
708:
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703:
693:
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689:
688:
680:
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671:
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658:
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648:
641:
640:
633:
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624:
623:
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619:
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603:
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596:
590:
589:
586:
585:
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571:
564:
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556:
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547:
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511:
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453:
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441:
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431:
423:
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409:
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403:
398:
393:
388:
385:
381:
377:
373:
369:
365:
361:
354:
352:
350:
346:
345:specific name
342:
338:
334:
330:
329:mantis shrimp
326:
321:
319:
315:
311:
307:
303:
299:
295:
287:
285:
283:
282:
277:
273:
269:
268:mantis shrimp
265:
261:
260:
247:
245:
236:
233:
232:Binomial name
229:
225:
224:
218:
215:
214:
207:
206:
199:
196:
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192:
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127:
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118:
104:
99:
94:
89:
84:
79:
74:
69:
64:
59:
54:
49:
43:
36:
32:
26:
22:
19:
1592:
1550:
1540:
1487:
1483:
1473:
1424:
1420:
1386:
1228:
1227:
1211:
1207:
1184:
1180:
1157:
1153:
1121:
1117:
1101:
1097:
1067:
1063:
1040:
1036:
1013:
1009:
986:
982:
956:
952:
929:
925:
894:
893:
889:
848:
847:
831:
830:
815:
788:
784:
768:
764:
727:
723:
700:
696:
662:
661:
645:
644:
631:Perimecturus
629:
611:
607:
568:
567:
551:
550:
537:Archaeocaris
535:
509:
493:
489:
481:
477:
463:
459:
455:
451:
443:
437:
433:
427:
413:
410:
389:
371:
362:was a small
359:
358:
348:
341:generic name
324:
322:
293:
291:
280:
279:
258:
257:
256:
243:
238:
222:
221:
204:
203:
168:Malacostraca
24:
18:
1663:Stomatopoda
1593:Chabardella
522:Stomatopoda
490:Chabardella
478:Chabardella
456:Chabardella
444:Chabardella
434:Chabardella
414:Chabardella
390:Though the
384:thoracopods
372:Chabardella
360:Chabardella
355:Description
337:exoskeleton
294:Chabardella
259:Chabardella
205:Chabardella
178:Stomatopoda
25:Chabardella
1652:Categories
1608:Q124387571
1553:: 95–110.
1490:(1): 290.
1391:References
971:Unipeltata
470:Unipeltata
364:crustacean
310:concretion
302:Stephanian
281:C. spinosa
158:Arthropoda
35:Stephanian
1569:1802-8225
1514:1471-2148
1465:130784496
1449:0022-3360
505:cladogram
430:raptorial
418:Paleozoic
376:raptorial
216:Species:
144:Kingdom:
138:Eukaryota
1602:Wikidata
1532:20858249
1457:29739132
474:polytomy
462:and the
392:carapace
333:holotype
184:Family:
154:Phylum:
148:Animalia
134:Domain:
1635:7698948
1622:9288267
1523:2955030
1492:Bibcode
1429:Bibcode
507:below:
497:pattoni
397:abdomen
349:spinosa
241:†
220:†
197:Genus:
174:Order:
164:Class:
109:↓
1567:
1530:
1520:
1512:
1463:
1455:
1447:
495:Daidal
482:et al.
452:et al.
406:uropod
402:telson
380:thorax
276:France
1461:S2CID
1453:JSTOR
446:into
318:Autun
264:genus
1617:GBIF
1565:ISSN
1528:PMID
1510:ISSN
1445:ISSN
374:has
48:PreꞒ
1555:doi
1518:PMC
1500:doi
1437:doi
274:in
266:of
37:),
1654::
1632::
1619::
1604::
1563:.
1549:.
1526:.
1516:.
1508:.
1498:.
1488:10
1486:.
1482:.
1459:.
1451:.
1443:.
1435:.
1425:83
1423:.
1419:.
1399:^
458:,
370:,
347:,
320:.
98:Pg
42:Ma
1571:.
1557::
1534:.
1502::
1494::
1467:.
1439::
1431::
1209:†
1182:†
1155:†
1119:†
1099:†
1065:†
1038:†
1011:†
984:†
954:†
927:†
891:†
813:†
786:†
766:†
725:†
698:†
627:†
609:†
533:†
468:+
201:†
188:†
103:N
93:K
88:J
83:T
78:P
73:C
68:D
63:S
58:O
53:Ꞓ
33:(
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