1545:
77:
52:
335:- usually for life - together with their common offspring. They are territorial, and express this by jointly emitting sounds when pairs from other groups enter their own territory. Their diet consists mainly of fruit. To a lesser extent they also eat other parts of plants and insects.
299:. The fur is predominantly black, and in contrast to closely related species, so are the hands. The head is small and roundish, along the throat a white or yellow, collar-like marking extends to the ears.
1305:
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331:
Their behaviour has not been studied much. They live in family groups in which they stay together as a
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with a fluffy coat and a long, bushy tail. This is longer than the body and cannot be used as a
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414:(3rd ed.). Baltimore: Johns Hopkins University Press. p. 144.
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292:
128:
1444:
1297:
744:
483:
148:
118:
98:
1215:
1357:
411:
Mammal
Species of the World: A Taxonomic and Geographic Reference
262:
1271:
1219:
443:
307:
Colombian black-handed titis are restricted to southern
1560:
1373:
1228:
1130:
1062:
814:
805:
742:
665:
525:
516:
291:Colombian black-handed titis are relatively small
311:. Their range is located in the departments of
377:10.2305/IUCN.UK.2020-3.RLTS.T39944A17973462.en
1580:
455:
8:
277:. It was originally described in the genus
1587:
1573:
1216:
811:
522:
462:
448:
440:
217:
50:
31:
375:
344:
396:
394:
7:
1541:
1539:
363:IUCN Red List of Threatened Species
253:Colombian black-handed titi monkey
226:Colombian black-handed titi range
25:
319:. The habitat of this species is
1631:Taxa named by Philip Hershkovitz
1606:IUCN Red List vulnerable species
1543:
327:Behavioral ecology and nutrition
75:
1:
1168:Colombian black-handed titi (
653:Vanzolini's bald-faced saki (
1559:. You can help Knowledge by
730:Reddish-brown bearded saki (
35:Colombian black-handed titi
1100:Coastal black-handed titi (
703:Brown-backed bearded saki (
27:Species of New World monkey
18:Colombian black-handed titi
1652:
1538:
1626:Mammals described in 1963
1621:Endemic fauna of Colombia
1202:
712:Uta Hick's bearded saki (
694:Red-backed bearded saki (
478:
470:Extant species of family
237:
230:
225:
216:
197:
190:
72:Scientific classification
70:
48:
39:
34:
897:Prince Bernhard's titi (
408:; Reeder, D. M. (eds.).
303:Distribution and habitat
978:Chestnut-bellied titi (
861:Olalla brothers' titi (
1636:New World monkey stubs
1555:-related article is a
1091:Coimbra Filho's titi (
1082:Barbara Brown's titi (
1518:Paleobiology Database
1050:Stephen Nash's titi (
879:Urubamba brown titi (
771:Black-headed uakari (
370:: e.T39944A17973462.
352:Palacios, E. (2020).
1109:Black-fronted titi (
1032:Hershkovitz's titi (
969:Alta Floresta titi (
888:Baptista Lake titi (
685:Black bearded saki (
321:tropical rainforests
1616:Mammals of Colombia
1023:White-tailed titi (
870:White-coated titi (
572:Golden-faced saki (
554:White-footed saki (
42:Conservation status
942:Red-bellied titi (
933:Hoffmanns's titi (
834:White-eared titi (
721:White-nosed saki (
644:White-faced saki (
599:Rio Tapajós saki (
261:) is a species of
1568:
1567:
1533:
1532:
1505:Open Tree of Life
1405:Cheracebus medemi
1375:Cheracebus medemi
1345:Open Tree of Life
1230:Callicebus medemi
1222:Taxon identifiers
1213:
1212:
1198:
1197:
1177:Red-headed titi (
1143:
1084:C. barbarabrownae
1075:
924:Ashy black titi (
827:
801:
800:
773:C. melanocephalus
755:
678:
545:Equatorial saki (
538:
356:Cheracebus medemi
258:Cheracebus medemi
249:
248:
244:
243:Hershkovitz, 1963
240:Callicebus medemi
201:Cheracebus medemi
65:
16:(Redirected from
1643:
1589:
1582:
1575:
1553:New World monkey
1547:
1540:
1526:
1525:
1513:
1512:
1500:
1499:
1487:
1486:
1474:
1473:
1461:
1460:
1448:
1447:
1435:
1434:
1422:
1421:
1409:
1408:
1407:
1394:
1393:
1392:
1366:
1365:
1353:
1352:
1340:
1339:
1327:
1326:
1314:
1313:
1301:
1300:
1288:
1287:
1275:
1274:
1262:
1261:
1249:
1248:
1247:
1217:
1142:
1141:
1137:
1074:
1073:
1069:
826:
825:
821:
812:
789:Neblina uakari (
754:
753:
749:
677:
676:
672:
590:Burnished saki (
547:P. aequatorialis
537:
536:
532:
523:
464:
457:
450:
441:
434:
433:
398:
389:
388:
386:
384:
379:
349:
267:New World monkey
242:
221:
203:
80:
79:
59:
54:
53:
32:
21:
1651:
1650:
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1412:
1403:
1402:
1397:
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1322:
1317:
1309:
1304:
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1270:
1265:
1257:
1252:
1243:
1242:
1237:
1224:
1214:
1209:
1194:
1186:Collared titi (
1139:
1138:
1136:
1126:
1118:Atlantic titi (
1071:
1070:
1068:
1058:
1052:P. stephennashi
1007:P. aureipalatii
996:Toppin's titi (
960:Milton's titi (
951:Vieira's titi (
881:P. urubambensis
852:Rio Mayo titi (
843:Rio Beni titi (
836:P. donacophilus
823:
822:
820:
797:
751:
750:
748:
738:
675:(Bearded sakis)
674:
673:
671:
661:
626:Miller's saki (
608:Isabel's saki (
574:P.chrysocephala
563:Cazuza's saki (
534:
533:
531:
512:
474:
468:
438:
437:
422:
400:
399:
392:
382:
380:
351:
350:
346:
341:
333:monogamous pair
329:
305:
297:prehensile tail
289:
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205:
199:
186:
74:
66:
55:
51:
44:
28:
23:
22:
15:
12:
11:
5:
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1410:
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1379:
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1371:
1370:
1368:
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1328:
1315:
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1226:
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1220:
1211:
1210:
1203:
1200:
1199:
1196:
1195:
1193:
1192:
1183:
1174:
1165:
1156:
1150:Lucifer titi (
1146:
1144:
1128:
1127:
1125:
1124:
1115:
1106:
1097:
1088:
1078:
1076:
1060:
1059:
1057:
1056:
1047:
1038:
1029:
1020:
1016:P. caquetensis
1014:Caquetá titi (
1011:
1002:
993:
987:Coppery titi (
984:
975:
966:
957:
948:
939:
930:
926:P. cinerascens
921:
915:Parecis titi (
912:
903:
894:
885:
876:
867:
858:
849:
840:
830:
828:
809:
803:
802:
799:
798:
796:
795:
786:
780:Aracá uakari (
777:
768:
758:
756:
740:
739:
737:
736:
727:
718:
709:
700:
691:
681:
679:
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650:
641:
632:
623:
614:
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578:
569:
560:
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541:
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535:(Saki monkeys)
520:
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206:
195:
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183:C. medemi
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49:
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36:
26:
24:
14:
13:
10:
9:
6:
4:
3:
2:
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1614:
1612:
1609:
1607:
1604:
1603:
1601:
1590:
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1576:
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1570:
1564:
1562:
1558:
1554:
1549:
1546:
1542:
1537:
1524:
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1515:
1511:
1506:
1502:
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1467:
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1459:
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1437:
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1411:
1406:
1400:
1396:
1391:
1385:
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1380:
1378:
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1372:
1364:
1359:
1355:
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1342:
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1333:
1329:
1325:
1320:
1316:
1312:
1307:
1303:
1299:
1294:
1290:
1286:
1281:
1277:
1273:
1268:
1264:
1260:
1255:
1251:
1246:
1240:
1236:
1235:
1233:
1231:
1227:
1223:
1218:
1208:
1207:
1201:
1191:
1189:
1184:
1182:
1180:
1175:
1173:
1171:
1166:
1164:
1162:
1157:
1155:
1153:
1148:
1147:
1145:
1135:
1134:
1129:
1123:
1121:
1120:C. personatus
1116:
1114:
1112:
1111:C. nigrifrons
1107:
1105:
1103:
1102:C. melanochir
1098:
1096:
1094:
1089:
1087:
1085:
1080:
1079:
1077:
1067:
1066:
1061:
1055:
1053:
1048:
1046:
1044:
1041:Ornate titi (
1039:
1037:
1035:
1030:
1028:
1026:
1021:
1019:
1017:
1012:
1010:
1008:
1005:Madidi titi (
1003:
1001:
999:
994:
992:
990:
985:
983:
981:
976:
974:
972:
967:
965:
963:
958:
956:
954:
949:
947:
945:
940:
938:
936:
935:P. hoffmannsi
931:
929:
927:
922:
920:
918:
913:
911:
909:
904:
902:
900:
895:
893:
891:
886:
884:
882:
877:
875:
873:
872:P. pallescens
868:
866:
864:
859:
857:
855:
850:
848:
846:
841:
839:
837:
832:
831:
829:
819:
818:
817:Plecturocebus
813:
810:
808:
804:
794:
792:
787:
785:
783:
778:
776:
774:
769:
767:
765:
762:Bald uakari (
760:
759:
757:
747:
746:
741:
735:
733:
728:
726:
724:
719:
717:
715:
710:
708:
706:
701:
699:
697:
696:C. chiropotes
692:
690:
688:
683:
682:
680:
670:
669:
664:
658:
656:
655:P. vanzolinii
651:
649:
647:
642:
640:
638:
633:
631:
629:
624:
622:
620:
615:
613:
611:
606:
604:
602:
597:
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593:
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586:
584:
579:
577:
575:
570:
568:
566:
561:
559:
557:
552:
550:
548:
543:
542:
540:
530:
529:
524:
521:
519:
515:
509:
505:
503:
499:
497:
493:
491:
487:
485:
481:
480:
477:
473:
465:
460:
458:
453:
451:
446:
445:
442:
431:
427:
423:
421:0-801-88221-4
417:
413:
412:
407:
406:Wilson, D. E.
403:
402:Groves, C. P.
397:
395:
391:
378:
373:
369:
365:
364:
359:
357:
348:
345:
338:
336:
334:
326:
324:
322:
318:
314:
310:
302:
300:
298:
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286:
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281:
276:
272:
268:
264:
260:
259:
254:
245:
241:
236:
233:
229:
224:
220:
215:
210:
204:
202:
196:
193:
192:Binomial name
189:
185:
184:
179:
176:
175:
172:
171:
167:
164:
163:
160:
157:
154:
153:
150:
147:
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137:
134:
133:
130:
127:
124:
123:
120:
117:
114:
113:
110:
107:
104:
103:
100:
97:
94:
93:
90:
87:
84:
83:
78:
73:
69:
63:
58:
47:
43:
38:
33:
30:
19:
1561:expanding it
1550:
1535:
1374:
1229:
1204:
1188:C. torquatus
1187:
1178:
1169:
1167:
1160:
1159:Black titi (
1151:
1131:
1119:
1110:
1101:
1092:
1083:
1063:
1051:
1042:
1033:
1024:
1015:
1006:
997:
988:
980:P. caligatus
979:
970:
961:
952:
943:
934:
925:
916:
907:
906:Brown titi (
899:P. bernhardi
898:
889:
880:
871:
862:
853:
844:
835:
815:
807:Callicebinae
790:
781:
772:
763:
743:
732:C. sagulatus
731:
723:C. albinasus
722:
714:C. utahickae
713:
705:C. israelita
704:
695:
686:
666:
654:
645:
636:
627:
618:
609:
600:
591:
582:
581:Hairy saki (
573:
564:
555:
546:
526:
410:
381:. Retrieved
367:
361:
355:
347:
330:
306:
290:
278:
265:, a type of
257:
256:
252:
250:
239:
238:
200:
198:
182:
181:
169:
145:Infraorder:
29:
1440:iNaturalist
1399:Wikispecies
1293:iNaturalist
1093:C. coimbrai
1025:P. discolor
908:P. brunneus
890:P. baptista
854:P. oenanthe
845:P. modestus
646:P. pithecia
637:P. napensis
635:Napo saki (
619:P. monachus
617:Monk saki (
601:P. irrorata
556:P. albicans
518:Pitheciinae
472:Pitheciidae
383:19 November
287:Description
209:Hershkovitz
159:Pitheciidae
149:Simiiformes
1611:Cheracebus
1600:Categories
1179:C. regulus
1152:C. lucifer
1133:Cheracebus
1065:Callicebus
1043:P. ornatus
998:P. toppini
989:P. cupreus
971:P. grovesi
962:P. miltoni
953:P. vieirai
917:P. parecis
863:P. olallae
782:C. ayresii
687:C. satanas
668:Chiropotes
628:P. milleri
610:P. isabela
583:P. hirsuta
565:P. cazuzai
508:Haplorhini
506:Suborder:
339:References
280:Callicebus
170:Cheracebus
139:Haplorhini
135:Suborder:
57:Vulnerable
1390:Q56120647
1170:C. medemi
1161:C. lugens
1034:P. dubius
944:P. moloch
791:C. hosomi
764:C. calvus
752:(Uakaris)
592:P. inusta
482:Kingdom:
283:in 1963.
177:Species:
95:Kingdom:
89:Eukaryota
1384:Wikidata
1358:Species+
1337:12100334
1311:10601211
1245:Q1284488
1239:Wikidata
1206:Category
528:Pithecia
502:Primates
496:Mammalia
490:Chordata
488:Phylum:
484:Animalia
430:62265494
404:(2005).
317:Putumayo
309:Colombia
293:primates
275:Colombia
232:Synonyms
155:Family:
129:Primates
119:Mammalia
109:Chordata
105:Phylum:
99:Animalia
85:Domain:
62:IUCN 3.1
1510:3607588
1497:1812106
1484:1000865
1458:1025105
1432:9407885
1350:3607588
1285:2436402
1140:(Titis)
1072:(Titis)
824:(Titis)
745:Cacajao
500:Order:
494:Class:
313:Caquetá
271:endemic
211:, 1963)
165:Genus:
125:Order:
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426:OCLC
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368:2020
315:and
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