1534:
66:
41:
324:- usually for life - together with their common offspring. They are territorial, and express this by jointly emitting sounds when pairs from other groups enter their own territory. Their diet consists mainly of fruit. To a lesser extent they also eat other parts of plants and insects.
288:. The fur is predominantly black, and in contrast to closely related species, so are the hands. The head is small and roundish, along the throat a white or yellow, collar-like marking extends to the ears.
1294:
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208:
1619:
1594:
450:
1415:
1268:
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Their behaviour has not been studied much. They live in family groups in which they stay together as a
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with a fluffy coat and a long, bushy tail. This is longer than the body and cannot be used as a
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403:(3rd ed.). Baltimore: Johns Hopkins University Press. p. 144.
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137:
107:
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400:
Mammal
Species of the World: A Taxonomic and Geographic Reference
251:
1260:
1208:
432:
296:
Colombian black-handed titis are restricted to southern
1549:
1362:
1217:
1119:
1051:
803:
794:
731:
654:
514:
505:
280:Colombian black-handed titis are relatively small
300:. Their range is located in the departments of
366:10.2305/IUCN.UK.2020-3.RLTS.T39944A17973462.en
1569:
444:
8:
266:. It was originally described in the genus
1576:
1562:
1205:
800:
511:
451:
437:
429:
206:
39:
20:
364:
333:
385:
383:
7:
1530:
1528:
352:IUCN Red List of Threatened Species
1548:. You can help Knowledge (XXG) by
242:Colombian black-handed titi monkey
215:Colombian black-handed titi range
14:
308:. The habitat of this species is
1620:Taxa named by Philip Hershkovitz
1595:IUCN Red List vulnerable species
1532:
316:Behavioral ecology and nutrition
64:
1:
1157:Colombian black-handed titi (
642:Vanzolini's bald-faced saki (
719:Reddish-brown bearded saki (
24:Colombian black-handed titi
1089:Coastal black-handed titi (
692:Brown-backed bearded saki (
16:Species of New World monkey
1641:
1527:
1615:Mammals described in 1963
1610:Endemic fauna of Colombia
1191:
701:Uta Hick's bearded saki (
683:Red-backed bearded saki (
467:
459:Extant species of family
226:
219:
214:
205:
186:
179:
61:Scientific classification
59:
37:
28:
23:
886:Prince Bernhard's titi (
397:; Reeder, D. M. (eds.).
292:Distribution and habitat
967:Chestnut-bellied titi (
850:Olalla brothers' titi (
1625:New World monkey stubs
1544:-related article is a
1080:Coimbra Filho's titi (
1071:Barbara Brown's titi (
1507:Paleobiology Database
1039:Stephen Nash's titi (
868:Urubamba brown titi (
760:Black-headed uakari (
359:: e.T39944A17973462.
341:Palacios, E. (2020).
1098:Black-fronted titi (
1021:Hershkovitz's titi (
958:Alta Floresta titi (
877:Baptista Lake titi (
674:Black bearded saki (
310:tropical rainforests
1605:Mammals of Colombia
1012:White-tailed titi (
859:White-coated titi (
561:Golden-faced saki (
543:White-footed saki (
31:Conservation status
931:Red-bellied titi (
922:Hoffmanns's titi (
823:White-eared titi (
710:White-nosed saki (
633:White-faced saki (
588:Rio Tapajós saki (
250:) is a species of
1557:
1556:
1522:
1521:
1494:Open Tree of Life
1394:Cheracebus medemi
1364:Cheracebus medemi
1334:Open Tree of Life
1219:Callicebus medemi
1211:Taxon identifiers
1202:
1201:
1187:
1186:
1166:Red-headed titi (
1132:
1073:C. barbarabrownae
1064:
913:Ashy black titi (
816:
790:
789:
762:C. melanocephalus
744:
667:
534:Equatorial saki (
527:
345:Cheracebus medemi
247:Cheracebus medemi
238:
237:
233:
232:Hershkovitz, 1963
229:Callicebus medemi
190:Cheracebus medemi
54:
1632:
1578:
1571:
1564:
1542:New World monkey
1536:
1529:
1515:
1514:
1502:
1501:
1489:
1488:
1476:
1475:
1463:
1462:
1450:
1449:
1437:
1436:
1424:
1423:
1411:
1410:
1398:
1397:
1396:
1383:
1382:
1381:
1355:
1354:
1342:
1341:
1329:
1328:
1316:
1315:
1303:
1302:
1290:
1289:
1277:
1276:
1264:
1263:
1251:
1250:
1238:
1237:
1236:
1206:
1131:
1130:
1126:
1063:
1062:
1058:
815:
814:
810:
801:
778:Neblina uakari (
743:
742:
738:
666:
665:
661:
579:Burnished saki (
536:P. aequatorialis
526:
525:
521:
512:
453:
446:
439:
430:
423:
422:
387:
378:
377:
375:
373:
368:
338:
256:New World monkey
231:
210:
192:
69:
68:
48:
43:
42:
21:
1640:
1639:
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1510:
1505:
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1458:
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1440:
1432:
1427:
1419:
1414:
1406:
1401:
1392:
1391:
1386:
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1371:
1358:
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1337:
1332:
1324:
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1311:
1306:
1298:
1293:
1285:
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1267:
1259:
1254:
1246:
1241:
1232:
1231:
1226:
1213:
1203:
1198:
1183:
1175:Collared titi (
1128:
1127:
1125:
1115:
1107:Atlantic titi (
1060:
1059:
1057:
1047:
1041:P. stephennashi
996:P. aureipalatii
985:Toppin's titi (
949:Milton's titi (
940:Vieira's titi (
870:P. urubambensis
841:Rio Mayo titi (
832:Rio Beni titi (
825:P. donacophilus
812:
811:
809:
786:
740:
739:
737:
727:
664:(Bearded sakis)
663:
662:
660:
650:
615:Miller's saki (
597:Isabel's saki (
563:P.chrysocephala
552:Cazuza's saki (
523:
522:
520:
501:
463:
457:
427:
426:
411:
389:
388:
381:
371:
369:
340:
339:
335:
330:
322:monogamous pair
318:
294:
286:prehensile tail
278:
201:
194:
188:
175:
63:
55:
44:
40:
33:
17:
12:
11:
5:
1638:
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1628:
1627:
1622:
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1612:
1607:
1602:
1597:
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1573:
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1537:
1520:
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1516:
1503:
1490:
1477:
1464:
1451:
1438:
1425:
1412:
1399:
1384:
1368:
1366:
1360:
1359:
1357:
1356:
1343:
1330:
1317:
1304:
1291:
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1265:
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1239:
1223:
1221:
1215:
1214:
1209:
1200:
1199:
1192:
1189:
1188:
1185:
1184:
1182:
1181:
1172:
1163:
1154:
1145:
1139:Lucifer titi (
1135:
1133:
1117:
1116:
1114:
1113:
1104:
1095:
1086:
1077:
1067:
1065:
1049:
1048:
1046:
1045:
1036:
1027:
1018:
1009:
1005:P. caquetensis
1003:Caquetá titi (
1000:
991:
982:
976:Coppery titi (
973:
964:
955:
946:
937:
928:
919:
915:P. cinerascens
910:
904:Parecis titi (
901:
892:
883:
874:
865:
856:
847:
838:
829:
819:
817:
798:
792:
791:
788:
787:
785:
784:
775:
769:Aracá uakari (
766:
757:
747:
745:
729:
728:
726:
725:
716:
707:
698:
689:
680:
670:
668:
652:
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639:
630:
621:
612:
603:
594:
585:
576:
567:
558:
549:
540:
530:
528:
524:(Saki monkeys)
509:
503:
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379:
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236:
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224:
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202:
195:
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183:
177:
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172:C. medemi
169:
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38:
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34:
29:
26:
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15:
13:
10:
9:
6:
4:
3:
2:
1637:
1626:
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1616:
1613:
1611:
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1606:
1603:
1601:
1598:
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1590:
1579:
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1565:
1560:
1559:
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1538:
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1531:
1526:
1513:
1508:
1504:
1500:
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1482:
1478:
1474:
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1461:
1456:
1452:
1448:
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1439:
1435:
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1426:
1422:
1417:
1413:
1409:
1404:
1400:
1395:
1389:
1385:
1380:
1374:
1370:
1369:
1367:
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1361:
1353:
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1335:
1331:
1327:
1322:
1318:
1314:
1309:
1305:
1301:
1296:
1292:
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1270:
1266:
1262:
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1240:
1235:
1229:
1225:
1224:
1222:
1220:
1216:
1212:
1207:
1197:
1196:
1190:
1180:
1178:
1173:
1171:
1169:
1164:
1162:
1160:
1155:
1153:
1151:
1146:
1144:
1142:
1137:
1136:
1134:
1124:
1123:
1118:
1112:
1110:
1109:C. personatus
1105:
1103:
1101:
1100:C. nigrifrons
1096:
1094:
1092:
1091:C. melanochir
1087:
1085:
1083:
1078:
1076:
1074:
1069:
1068:
1066:
1056:
1055:
1050:
1044:
1042:
1037:
1035:
1033:
1030:Ornate titi (
1028:
1026:
1024:
1019:
1017:
1015:
1010:
1008:
1006:
1001:
999:
997:
994:Madidi titi (
992:
990:
988:
983:
981:
979:
974:
972:
970:
965:
963:
961:
956:
954:
952:
947:
945:
943:
938:
936:
934:
929:
927:
925:
924:P. hoffmannsi
920:
918:
916:
911:
909:
907:
902:
900:
898:
893:
891:
889:
884:
882:
880:
875:
873:
871:
866:
864:
862:
861:P. pallescens
857:
855:
853:
848:
846:
844:
839:
837:
835:
830:
828:
826:
821:
820:
818:
808:
807:
806:Plecturocebus
802:
799:
797:
793:
783:
781:
776:
774:
772:
767:
765:
763:
758:
756:
754:
751:Bald uakari (
749:
748:
746:
736:
735:
730:
724:
722:
717:
715:
713:
708:
706:
704:
699:
697:
695:
690:
688:
686:
685:C. chiropotes
681:
679:
677:
672:
671:
669:
659:
658:
653:
647:
645:
644:P. vanzolinii
640:
638:
636:
631:
629:
627:
622:
620:
618:
613:
611:
609:
604:
602:
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591:
586:
584:
582:
577:
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568:
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564:
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557:
555:
550:
548:
546:
541:
539:
537:
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531:
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518:
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494:
492:
488:
486:
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469:
466:
462:
454:
449:
447:
442:
440:
435:
434:
431:
420:
416:
412:
410:0-801-88221-4
406:
402:
401:
396:
395:Wilson, D. E.
392:
391:Groves, C. P.
386:
384:
380:
367:
362:
358:
354:
353:
348:
346:
337:
334:
327:
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323:
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307:
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291:
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243:
234:
230:
225:
222:
218:
213:
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191:
185:
182:
181:Binomial name
178:
174:
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161:
160:
156:
153:
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149:
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99:
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67:
62:
58:
52:
47:
36:
32:
27:
22:
19:
1550:expanding it
1539:
1524:
1363:
1218:
1193:
1177:C. torquatus
1176:
1167:
1158:
1156:
1149:
1148:Black titi (
1140:
1120:
1108:
1099:
1090:
1081:
1072:
1052:
1040:
1031:
1022:
1013:
1004:
995:
986:
977:
969:P. caligatus
968:
959:
950:
941:
932:
923:
914:
905:
896:
895:Brown titi (
888:P. bernhardi
887:
878:
869:
860:
851:
842:
833:
824:
804:
796:Callicebinae
779:
770:
761:
752:
732:
721:C. sagulatus
720:
712:C. albinasus
711:
703:C. utahickae
702:
694:C. israelita
693:
684:
675:
655:
643:
634:
625:
616:
607:
598:
589:
580:
571:
570:Hairy saki (
562:
553:
544:
535:
515:
399:
370:. Retrieved
356:
350:
344:
336:
319:
295:
279:
267:
254:, a type of
246:
245:
241:
239:
228:
227:
189:
187:
171:
170:
158:
134:Infraorder:
18:
1429:iNaturalist
1388:Wikispecies
1282:iNaturalist
1082:C. coimbrai
1014:P. discolor
897:P. brunneus
879:P. baptista
843:P. oenanthe
834:P. modestus
635:P. pithecia
626:P. napensis
624:Napo saki (
608:P. monachus
606:Monk saki (
590:P. irrorata
545:P. albicans
507:Pitheciinae
461:Pitheciidae
372:19 November
276:Description
198:Hershkovitz
148:Pitheciidae
138:Simiiformes
1600:Cheracebus
1589:Categories
1168:C. regulus
1141:C. lucifer
1122:Cheracebus
1054:Callicebus
1032:P. ornatus
987:P. toppini
978:P. cupreus
960:P. grovesi
951:P. miltoni
942:P. vieirai
906:P. parecis
852:P. olallae
771:C. ayresii
676:C. satanas
657:Chiropotes
617:P. milleri
599:P. isabela
572:P. hirsuta
554:P. cazuzai
497:Haplorhini
495:Suborder:
328:References
269:Callicebus
159:Cheracebus
128:Haplorhini
124:Suborder:
46:Vulnerable
1379:Q56120647
1159:C. medemi
1150:C. lugens
1023:P. dubius
933:P. moloch
780:C. hosomi
753:C. calvus
741:(Uakaris)
581:P. inusta
471:Kingdom:
272:in 1963.
166:Species:
84:Kingdom:
78:Eukaryota
1373:Wikidata
1347:Species+
1326:12100334
1300:10601211
1234:Q1284488
1228:Wikidata
1195:Category
517:Pithecia
491:Primates
485:Mammalia
479:Chordata
477:Phylum:
473:Animalia
419:62265494
393:(2005).
306:Putumayo
298:Colombia
282:primates
264:Colombia
221:Synonyms
144:Family:
118:Primates
108:Mammalia
98:Chordata
94:Phylum:
88:Animalia
74:Domain:
51:IUCN 3.1
1499:3607588
1486:1812106
1473:1000865
1447:1025105
1421:9407885
1339:3607588
1274:2436402
1129:(Titis)
1061:(Titis)
813:(Titis)
734:Cacajao
489:Order:
483:Class:
302:Caquetá
260:endemic
200:, 1963)
154:Genus:
114:Order:
104:Class:
49: (
1512:390840
1434:738181
1313:612148
1261:323587
417:
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