Oryzomys couesi - Knowledge (XXG)

1741: 1732: 1390: 1381: 3046: 3037: 2171: 622: 2162: 1932: 3624:, such as drainage of wetlands, may threaten some populations. In many areas, it is so common that it is considered a plague species. Populations even persist in the Valley of Mexico, as evidenced by a photograph published in 2006. However, the species is listed as threatened in Texas, where its distribution is very limited, because of habitat loss. In 1979, Benson and Gehlbach estimated the size of the Texas population to be about 15,000. A 2001 study predicted that 1923: 96: 228: 71: 6688: 3280:) at the outer front (anterolabial) edge of the molar, before the protoconid. The third lower molar also bears an anterolophid and an anterolabial cingulum. The first lower molar has large roots at the front and back of the tooth and two smaller ones in between, at the labial and lingual side. The second and third lowers molars have two large roots, one at the front and one at the back. 52: 2905:. Head and body length is 98 to 142 mm (3.9 to 5.6 in), tail length is 107 to 152 mm (4.2 to 6.0 in), hindfoot length is 27 to 33 mm (1.1 to 1.3 in), ear length is 13 to 18 mm (0.51 to 0.71 in), and body mass is 43 to 82 g (1.5 to 2.9 oz). Studies in Texas and El Salvador found that males are slightly larger than females. 6674: 1527:, which Thomas had described in 1893, has no more precise locality than "Mexico", but the Valley of Mexico has been suggested as its origin. It is a large, coarse-furred, bright reddish, long-tailed species with a broad skull with widely spreading zygomatic arches. Goldman wrote that it was similar to 859:

to the other specimens, which fell into two large subclades—one containing animals from the Pacific seaboard from western Mexico to El Salvador and the other containing rats from the eastern seaboard from Texas to Nicaragua. The Panamanian and Costa Rican populations differed by 6.53% to 11.93% from

2884:

is a medium-sized to large rat with coarse fur that is buff to reddish above, becoming paler towards the sides and cheeks and darker on the rump and face. The underparts are white to buff. The fur is shorter, brighter, and more intense in color than in the marsh rice rat. The snout ends bluntly and

3419:

move up to 153 m (502 ft) and females up to 126 m (413 ft). The diet includes both plant material, including seeds and green parts, and animals, including small fish, crustaceans, snails, insects like ants and beetles, and other invertebrates. It probably breeds around the year

3403:

builds globular nests of woven vegetation suspended among reeds, about 1 m (3.3 ft) above the water or the ground; in Texas, larger individuals make larger nests. It does not usually make its own runways in vegetation, but may use those of other rodents, such as cotton rats.

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among shrubs. It occurs from 2,300 m (7,500 ft) altitude down to sea level. On Cozumel, the proportion of juveniles and females is higher near roads that function as habitat edges. Cozumel rice rats rarely cross roads, which may isolate subpopulations on the island.

3115:, openings in the front part of the palate, reach backward between the molars. The palate is long, extending substantially beyond the third molars, the usual condition in oryzomyines. The back part, near the third molars, is usually perforated by prominent 3092:, the flattened front part of the zygomatic arch, is broad and develops a notch at its front end. The plate's back margin is located before the first upper molar. The jugal bone, part of the zygomatic arch, is reduced, as usual in oryzomyines. The 1531:, but more intensely colored, and differed in the form of the interorbital region; he retained it as a separate species pending further investigations. Carleton and Arroyo-Cabrales noted that archival research may yet uncover the precise origin of 3628:

would drive the Texas population to extinction, because no suitable habitats would continue to exist. The Cozumel population has declined substantially since the mid-1980s, perhaps due to habitat disturbance and predation by introduced species.

3253:, is present behind the labial cuspule. As in most oryzomyines, the upper molars all have one root on the inner (lingual) side and two on the outer (labial) side; in addition, the first upper molar usually has another small labial root. 2358:

and the synonyms currently associated with it. Eliot in 1905 and Goldman in 1918 gave separate common names for each of the species and subspecies they recognized. Many authors have used "Coues' Rice Rat" or some variation thereof for

466:

previously included many other species, which were reclassified in various studies culminating in contributions by Marcelo Weksler and coworkers in 2006 that removed more than forty species from the genus. All are placed in the tribe

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range from 5 to 30 per ha (2 to 12 per acre). On Cozumel, density is around 14.5 to 16.5 per ha (5.9 to 6.7 per acre), but shows large seasonal variation. In western Mexico, one study found densities of 3 per ha (1.2 per acre) in

1913:

Allen, 1910, from Nicaragua. According to Goldman, individual variation within the subspecies is large, which has led to the large number of published synonyms, but populations from all parts of its range are essentially similar.

346:, it eats both plant and animal food, including seeds and insects. It breeds throughout the year; females give birth to about four young after a pregnancy of 21 to 28 days. The species may be infected by several different 6361:(Alston, 1877). pp. 709–710 in Ceballos, G. and Oliva, G. (eds.). Los mamíferos silvestres de México. Mexico City: Comisión Nacional para el Conocimiento y Uso de la Biodiversidad and Fondo de Cultura Económica, 986 pp. 2889:. The small ears are black on the outside and the inside is covered with short, gray to buff or red hairs. The long tail is dark brown above and white to light brown below. The feet are long and stout. On the forefeet, the 1998:

populations; accordingly, they demoted the island form to a subspecies. Mark Engstrom and colleagues, writing in 1989, reaffirmed this conclusion. For an island form, this population is highly genetically variable. In its

3100:(opening) at the side of the skull above the molars, is small; it is much larger in the marsh rice rat. The narrowest part of the interorbital region is towards the front and the edges are lined by prominent shelves. The 341:

is active during the night and builds nests of vegetation that are suspended among reeds about 1 m (3.3 ft) above the ground. It is an excellent swimmer and dives well, but can also climb in vegetation. An

385:

and the marsh rice rat meet, were published in 1979 and underscored the distinctness of the two, they were again regarded as separate. Since then, some of the peripheral forms of the group, such as

1164:, was described by Burt in 1934 from central coastal Sonora, which extended the range of the species by 400 mi (640 km) at the time. This form is dark gray-brown, much darker than 381:), into a single widespread species; subsequently, many related, localized species retained by Goldman were also included in this taxon. After studies of the contact zone in Texas, where 317:

occurring from southernmost Texas through Mexico and Central America into northwestern Colombia. It is usually found in wet habitats, such as marshes, but also lives in drier forests and

2244:. It is a brown form, lacking the reddish tones of nearby populations, and has a broad skull with a short rostrum (front part) and ridges on the interorbital region like those of 2076:). They considered that this population probably represented a separate subspecies, but declined to propose a new name because they had only one adult specimen. In Nicaragua, 2901:

may be present in at least some specimens. Some of these traits are common adaptations to life in the water in oryzomyines. As in most other oryzomyines, females have eight

2995:, but there is a broad band of nonspinous tissue. The papilla (nipple-like projection) on the dorsal (upper) side of the penis is covered with small spines, a character 1807:

occurs further inland, while the marsh rice rat lives along the coast. In experimental conditions, the two fail to interbreed and genetic analysis yields no evidence of

2064:

are distinctive in their large skull and small external measurements, with an especially short tail, soft fur that is orange-brown above and buffish below, and lack of

3424:. In 28 pregnant females from Nicaragua, litter size varied from one to eight, averaging 4.4. The young become reproductively active when seven weeks old and the 8434: 3647: 3234:

shares with most but not all other oryzomyines. The flexi and flexids (valleys between the cusps and crests) at the labial (outer) side of the molars are closed by

2129:, but Hanson and colleagues found that two specimens from Refugio Nacional de Vida Silvestre Mixto Maquenque, northeastern Costa Rica, differed as much from other 3645:

The date of publication of Alston's name is listed alternatively in the literature as 1876 or 1877. The name was proposed on p. 756 of the 1876 issue of the

705:

contact zone in Texas published in 1979, using more specimens and characters, indicated that the two species are in fact easily distinguishable there; therefore,

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in color throughout and according to Hershkovitz almost identical to specimens from Guatemala, but distinctive in that the upper lip is white. He suggested that

1982:, and Goldman kept it as such because of its large size, dark fur, and long tail. In 1965, however, Knox Jones and Timothy Lawlor judged the differences between 1873:

Goldman united populations ranging from northern Veracruz through eastern Mexico, Guatemala, Honduras, and Nicaragua south to far northwestern Costa Rica in the

8512: 3651:, but the parts of this journal were not always published in the volume year. A 2005 review confirms that p. 756 of the 1876 volume was published in 1877. 8603: 8568: 800: 3377:, spending much time in the water, as Alston in his original description already recognized, but is also a good climber. A study in Costa Rica found that 6081: 5728:

Notes on new or little-known North American mammals, based on recent additions to the collection of mammals in the American Museum of Natural History

3119:, which are recessed into fossae (depressions). Sphenopalatine vacuities are usually absent, but have been reported in some populations. There is no 8588: 8408: 6722: 3576:

infecting the marsh rice rat, a common cause of hantavirus infections in the United States. No hantavirus infections in humans have been linked to

8447: 1500:

from the lowlands of the Baja California Peninsula they could not accept and they recommended further research to determine the relationships of

5924: 1070:, and northwestern Oaxaca. It usually lives below 1,000 m (3,300 ft) altitude, but has been found at 1,525 m (5,003 ft) in 6612:

Phylogenetic relationships of oryzomyine rodents (Muroidea: Sigmodontinae): separate and combined analyses of morphological and molecular data

6523: 6495: 6481: 6449: 6366: 3320:

is absent, as in all members of the Sigmodontinae; if present, as in some other rodents, this foramen perforates the distal (far) end of the

3420:

and after a pregnancy of 21 to 28 days, the female produces litters of two to seven young, with an average of 3.8, according to Reid's

3589: 2874:

Measurements are in millimeters and are in the form "average (minimum–maximum)", except where there are only one or two measured specimens.

3348:

cave deposits in Mexico and Honduras. It is common in watery habitats, such as marshes and small streams, but also occurs in forests and

8573: 1007:

in the strict sense, which occurs to the north and east, are separated by mountain ranges from the latter, harbor different species of

8598: 5959:

Deardorff, E.R., Forrester, N.L., Travassos da Rosa, A.P., Estrada-Franco, J.G., Navarro-Lopez, R., Tesh, R.B. and Weaver, S.C. 2010.

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Deardorff, E.R., Forrester, N.L., Travassos da Rosa, A.P., Estrada-Franco, J.G., Navarro-Lopez, R., Tesh, R.B. and Weaver, S.C. 2009.

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the others and the western and eastern subclades differed by 4.41% on average. Data from both of the slower-evolving nuclear markers

425:

is common and of no conservation concern, and it is even considered a plague species in places, but some populations are threatened.

6307: 5997: 610:

to a subspecies of the marsh rice rat. Other workers continued this lumping and by 1971 all other species Goldman had placed in the

3381:

is an excellent swimmer, diving well and using its tail to propel itself. It is probably able to forage underwater, which may help

3170:, which anchor some of the chewing muscles, join at a point below the first molar and do not extend forward beyond that point. The 6501: 8473: 6397:

Milazzo, M.L., Cajimat, M.N., Hanson, J.D., Bradley, R.D., Quintana, M., Sherman, C., Velásquez, R.T. and Fulhorst, C.F. 2006.

6095:

Fuentes-Montemayor, E., Cuarón, A.D., Vázquez-Domínguez, E., Benítez-Malvido, J., Valenzuela-Galván, D. and Andresen, E. 2009.

357:

The species was first described in 1877, the first of many related species from the region described until the 1910s. In 1918,

5976:

The Proceedings of the Zoological Society of London, 1859–1900: an exploration of breaks between calendar years of publication

8491: 1858:

fossils of this form have been found in Cueva de Abre, Tamaulipas. According to Goldman, it is intermediate in color between

8452: 3352:

with sufficient cover. In addition, it is found in sugarcane and rice fields. In Texas, it occurs in marsh vegetation along

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in northwestern Costa Rica and southeastern Nicaragua. In a 1966 review of Panamanian mammals, Charles Handley reduced both

1839: 709:

has since been regarded as a species distinct from the marsh rice rat. Afterward, some of the other forms synonymized under

548:, Guatemala, and two from Mexico) used by Alston in fact belonged to two or three different species. He restricted the name 5459:

Woodman, 1995, table 1, p. 225; Woodman and Croft, 2005, table 4; Dalquest and Roth, 1970, pp. 220, 226

6463:

Catalog of the type specimens of mammals in the United States National Museum, including the Biological Surveys collection

5771: 3166:, an opening just before the first molar, opens sidewards, not upwards as in a few other oryzomyines. The upper and lower 1819:

is larger and paler and has a more robust skull. Specimens from Tamaulipas are slightly darker than those from Texas. The

8578: 6715: 6438:

Musser, G.G. and Carleton, M.D. 2005. Superfamily Muroidea. pp. 894–1531 in Wilson, D.E. and Reeder, D.M. (eds.).

8486: 8369: 3116: 606:

reviewed specimens from this contact zone and found no grounds on which to separate the two species; thus, he reduced

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for one of the Mexican animals. Several other related species were described from the early 1890s onwards and in 1901

3340:, but not the central plateau of Mexico, through Central America south and east to northwestern Colombia; see under " 904:

should be recognized as distinct species. If this suggestion is followed, the eastern subclade would retain the name

8583: 8517: 7921: 6678: 5902:

Carleton, M.D. and Musser, G.G. 1984. Muroid rodents. pp. 289–379 in Anderson. S. and Jones, J.K. Jr. (eds.).

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group with the marsh rice rat and several species with more limited distributions, which he regarded as related to

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Platt, S.G., Rainwater, T.R., Miller, B.W. and Miller, C.M. 2000. Notes on the mammals of Turneffe Atoll, Belize.

5888: 3104:

extend to the sides of the braincase. The interparietal bone is narrow and wedge-shaped, so that the parietal and

1213:, but with the zygomatic arches broadly spreading and curved downward. Goldman, who reduced it to a subspecies of 8252: 7928: 6652:

Fossil shrews from Honduras and their significance for late glacial evolution in body size (Mammalia: Soricidae:

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differed from the marsh rice rat (11.30%). They suggested that these animals represented a species distinct from

1473: 1039:

and El Salvador) groups form distinct subclades; Hanson and colleagues recognized these as different subspecies,

752: 455: 398: 95: 6518:

Schmidly, D.J. and Davis, W.B. 2004. The mammals of Texas. 2nd edition. University of Texas Press, 501 pp.

3498: 8245: 7034: 6919: 6798: 3541: 3479: 3473: 2065: 1695:

clade, within which the average sequence divergence is 1.28%, and Hanson and colleagues proposed that the name

471:("rice rats"), a diverse assemblage of over a hundred species, and on higher taxonomic levels in the subfamily 3529: 2304:, which suggests that they may represent a separate species. However, Hanson and colleagues did not reinstate 839:

between the two groups was 11.30%, much larger than the distance between sister species in the related genera

6078: 5250:

Weksler, 2006, p. 37; Carleton and Arroyo-Cabrales, 2009, p. 117; Sánchez et al., 2001, p. 210

897: 409:

data, found evidence to recognize separate species from the Pacific and eastern sides of the distribution of

8274: 8201: 7963: 7907: 7893: 7886: 7879: 7678: 7671: 7598: 7377: 7141: 6708: 6261:

The glans penis in Neotropical cricetines (Family Muridae) with comments on classification of muroid rodents

3019: 2110: 1882: 1723:, that was described after Goldman's paper occur in the same region, but have not been studied genetically. 1403: 1296: 567: 519: 358: 4898:

Eliot, 1905, p. 186; Goldman, 1918, p. 29; Linzey et al., 2008; Duff and Lawson, 2004, p. 54

3504: 8322: 7956: 7914: 7664: 7549: 7191: 7119: 6805: 5844:

Bole, B.J. Jr. 1937. Annotated list of the mammals of the Mariato River District of the Azuero Peninsula.

3547: 3517: 3317: 3289: 3093: 1176:, but that further research is needed to determine whether it should be recognized as a subspecies. Large 2069: 8538: 8374: 8281: 7949: 7935: 7865: 7626: 7605: 7556: 7319: 7281: 7112: 7098: 7048: 7013: 6594:

Unexpected high levels of genetic variability and the population structure of an island endemic rodent (

3425: 3382: 3277: 3261: 3136: 2292:

was reinstated as a separate species these forms went with it. Specimens from near the type locality of

2145:, but were unable to resolve the correct name for the species because they could not examine samples of 1796: 1094: 812: 487: 217: 200: 3616:", because it is a widely distributed, common species with broad habitat tolerance that occurs in many 3455: 2327: 2220:, but distinctive in the well-developed ridges along the margins of the interorbital region, the short 2042:

as separate, because they thought the difference in color too small for the recognition of subspecies.

594:

but distinctive enough to be classified as separate species. In the 1930s, a few more forms related to

6124: 5739:

Further notes on mammals collected in Mexico by Dr. Audley C. Buller, with descriptions of new species

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Benson and Gehlbach, 1979, p. 227; Reid, 2009, p. 207; Schmidly and Davis, 2004, p. 281

3167: 3124: 3014: 3006: 1740: 734: 8395: 8360: 8194: 8172: 7970: 7942: 7900: 7872: 7858: 7829: 7633: 7563: 7384: 7326: 7273: 7205: 7148: 7105: 7091: 7084: 7077: 7041: 7027: 6940: 6912: 6692: 5950:

Experimental infection of potential reservoir hosts with Venezuelan equine encephalitis virus, Mexico

3535: 3017:) is present; it is absent in all other oryzomyines with studied penes except the marsh rice rat and 2921: 2898: 1676: 1665: 1654: 1624: 1602: 1591: 1274: 1263: 1217:, recorded it from southwestern Guatemala and nearby Chiapas and described it as slightly paler than 1088: 1076: 975: 964: 557: 5910: 1731: 1389: 1241:

from the country, while noting that specimens from some localities were slightly paler than others.

8073: 8044: 8030: 8022: 7978: 7685: 7656: 7619: 7289: 7155: 7070: 6926: 3621: 3246: 2044: 1874: 1699:

be restricted to this clade. These populations correspond to two subspecies recognized by Goldman (

1468: 1422: 1380: 1189: 1020: 757: 747: 729: 687: 681: 659: 598:

were described. As then recognized, the ranges of the marsh rice rat, a United States species, and

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and at least six more narrowly distributed species with peripheral distributions together form the

393: 387: 60: 8504: 6077:

Estébanes-González, M.L., Sánchez-Hernández, C., Romero-Almaraz, M. de L. and Schnell, G.D. 2011.

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as a full species in 1897 from specimens from Jalisco. In the same publication, he also described

603: 366: 282: 7836: 7745: 7731: 7591: 7527: 7265: 7219: 7184: 7020: 6905: 3569: 3523: 3432: 3407: 3301: 3245:, the front cusp of the upper first molar, is not divided in two by an indentation at its front ( 2917: 2319: 2221: 2089: 1754: 1548: 1250: 1152:

in 1918 and in 2009 Carleton and Arroyo-Cabrales concurred, arguing that the differences between

921: 766: 248: 90: 8543: 6026:

A checklist of mammals of the North American continent, the West Indies and the neighboring seas

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Descriptions of apparently new species and subspecies of mammals and a new generic name proposed

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Chu, Y.-K., Owen, R.D., Sánchez-Hernández, C., Romero-Almarez, M. de L. and Jonsson, C.B. 2008.

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Goldman, 1918, p. 39; Jones et al., 1983, p. 379; Hall and Dalquest, 1963, p. 289

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On the first and second upper molars, the flexi do not extend to the midline of the molars. The

3036: 621: 8439: 1140:, but he did not compare the two with each other. Merriam added a second species from Nayarit, 8593: 8525: 8382: 8165: 7815: 7766: 7612: 7584: 7577: 7570: 7513: 7499: 7477: 7470: 7333: 7198: 7177: 6933: 6519: 6491: 6477: 6462: 6445: 6372: 6362: 6303: 6135: 6025: 6014: 5993: 5760: 3593: 3387: 3297: 3293: 3276:, that form the front edge of the molar in some other oryzomyines. There is a distinct ridge ( 3120: 3010: 2241: 2213: 2170: 777: 724: 5749: 5738: 1970:, which also occurs in Belize, and has a more delicate skull. In 1901, Merriam described the 1160:

were age-related and within the normal range of variation of the animal. Another subspecies,

8530: 8059: 8051: 7822: 7787: 7780: 7759: 7506: 7456: 7449: 7442: 7435: 7428: 7355: 7345: 7297: 6947: 3929:

Musser and Carleton, 2005, p. 1152; Weksler et al., 2006, table 1, footnote e

3354: 3235: 3171: 3112: 3059: 2264:

as a species because it did not seem intermediate between the geographically closest forms,

2161: 1931: 1812: 1643: 1613: 1569: 1493: 1125: 953: 934: 836: 719: 693: 8465: 6004:

Fleas (Siphonaptera) of the Yucatan Peninsula (Campeche, Quintana Roo, and Yucatan), Mexico

2216:. In 1918, Goldman kept the animal as a separate species, remarking that it was similar to 1922: 7808: 7801: 7773: 7752: 7738: 7520: 7463: 7212: 6700: 6085: 3881:

Alston, 1877, pp. 756–757; Musser and Carleton, 2005, p. 1189, for placement in

3565: 3556: 3437: 3309: 3305: 3148: 3089: 1847: 1843: 1285: 1074:. This distributional pattern is similar to that of other western Mexican rodents such as 880:

placed the Costa Rican population within the marsh rice rat clade and placed some western

531: 270: 5687:

Medellín and Medellín, 2006, p. 710; Carleton and Arroyo-Cabrales, 2009, p. 113

2056:

in southeastern Nicaragua. According to Jones and Engstrom, rice rats from the island of

6217: 8292: 8235: 8065: 6278: 3625: 3617: 3214:, with the chewing edge located behind the vertical plane of the teeth. The molars are 3187: 3163: 3144: 3140: 3105: 3085: 2969: 2953: 2061: 2012: 1792: 1082: 1016: 912:, and the appropriate names for the Panamanian and Costa Rican species remain unclear. 505: 447: 405:

as currently constituted is likely a composite of several species; a 2010 study, using

374: 6398: 6328: 6096: 1488:, but could not exclude the possibility that they represent a different species. That 892:

and placed the Costa Rican population marginally closer to the marsh rice rat than to

544:, had caused much confusion about its identity, because the three specimens (one from 337:. There is much geographic variation in size, proportions, color, and skull features. 227: 8562: 8460: 8184: 8119: 8109: 6859: 6849: 6777: 6580:

Population and community ecology of small rodents in montane forest of western Mexico

6337: 6279:

Mammals from Isla Cozumel, México, with description of a new species of harvest mouse

3613: 3513: 3363: 3203: 3101: 3077: 2992: 1788: 1580: 1431: 1012: 773:

and related species would certainly result in the recognition of additional species.

527: 472: 329:

below. The hindfeet show some specializations for life in the water, such as reduced

325:

is a medium-sized to large rat. The coarse fur is buff to reddish above and white to

167: 80: 75: 17: 6547: 6110: 5795: 5606:

Medellín and Medellín, 2006, p. 710; Reid, 2006, p. 303; 2009, p. 207

4517:

Goldman, 1918, fig. 3; Murie, 1932, p. 1; Handon et al., 2010, fig. 1

3174:, a raising of the bone of the back of the mandible that houses the back end of the 8387: 8264: 7848: 7367: 7003: 6991: 6981: 6883: 6872: 6837: 6829: 6818: 6651: 6206: 6199: 5875:

Getting warmer: Effect of global climate change on distribution of rodents in Texas

3412: 3265: 3207: 3152: 3132: 3127:

that in some oryzomyines separates two foramina in the skull. The condition of the

3081: 3073: 3001: 2957: 2929: 2925: 1099: 781: 406: 330: 6593: 3222:, low-crowned, as in most other oryzomyines. Many accessory crests, including the 1353: 1059: 6383: 6150: 5785: 1535:, which could establish it as an older name for one of the other central Mexican 8499: 8478: 8421: 8354: 8000: 7990: 7794: 7645: 7060: 6969: 6959: 5960: 5761:

On a collection of mammals from Jalapa and Las Vigas, state of Vera Cruz, Mexico

3573: 3552: 3489: 3374: 3345: 3257: 3211: 2961: 2949: 2948:

arms (2n = 56, FNa = 56). The autosomes include 26 pairs of

2890: 2351: 2339: 2323: 1855: 1337: 856: 792: 307: 41: 8345: 6687: 6064: 5975: 4152:

Carleton and Arroyo-Cabrales, 2009, pp. 118–120; Goldman, 1918, p. 34

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in Veracruz include unidentified ticks, mites, fleas, and fly larvae. The flea

1122:—which are larger and different in some proportions and details of coloration. 8303: 8223: 8213: 8097: 8086: 7709: 7698: 7539: 7131: 6895: 6788: 6771: 6732: 6274:) of Nicaragua. Occasional Papers, The Museum, Texas Tech University 103:1–23. 6003: 5103:

Schmidt and Engstrom, 1994, p. 916; Hanson et al., 2010, pp. 342–343

3561: 3415:

and 1 per ha (0.4 per acre) in a disturbed area. In 24 hours, male Texas

3359: 3313: 3269: 3250: 3242: 3227: 3219: 2965: 2937: 2331: 2229: 1772: 1169: 1008: 579: 500: 494:

in 1877, using three specimens from Mexico and Guatemala. He named the animal

476: 468: 418: 351: 334: 314: 157: 51: 5477:

Schmidly and Davis, 2004, p. 381; Benson and Gehlbach, 1979, p. 228

776:

A 2010 study by Delton Hanson and colleagues used DNA sequence data from the

7309: 7254: 7167: 6327:

Linzey, A. V.; Timm, R.; Woodman, N.; Matson, J. & Samudio, R. (2017) .

5949: 3585: 3396: 3349: 3273: 2988: 2941: 2933: 2225: 1808: 1800: 1199:

and the underparts are usually white, but may be buffy, the normal color in

888:

group. The combined dataset supported the western and eastern clades within

841: 318: 107: 6433:

Miscellaneous Publications of the Museum of Zoology, University of Michigan

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Miscellaneous Publications of the Museum of Zoology, University of Michigan

2016: 6673: 6281:. University of Kansas Publications, Museum of Natural History 16:409–419. 6270:

Jones, J.K. Jr. and Engstrom, M.D. 1986. Synopsis of the rice rats (genus

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Miscellaneous Publications of the University of Michigan Museum of Zoology

6202:. University of Kansas Publications, Museum of Natural History 14:165–362. 6037:

Records of a collection of Mallophaga and Anoplura from Nicaraguan mammals

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Genetic characterization and phylogeny of a hantavirus from Western Mexico

5067:

Benson and Gehlbach, 1979, p. 226; Burt and Stirton, 1961, p. 60

3196: 3193: 3139:, an opening in the back part of the skull determined by the shape of the 2026:

of the eastern lowlands of Nicaragua was described as a separate species,

872:

in two main clades, but did not recover the western and eastern groups of

8339: 8155: 8143: 8132: 8012: 7489: 7418: 7406: 7396: 7241: 7231: 6753: 6441:

Mammal Species of the World: a taxonomic and geographic reference. 3rd ed

5798:. University of Kansas Publications, Museum of Natural History 5:207–215. 3468: 3446: 3223: 3215: 3159: 3143:, is present. The squamosal lacks a suspensory process that contacts the 2945: 2886: 2272:, and was separated by a large gap from the nearest known populations of 2209: 2184: 1962:

was described in 1932 from a pine ridge in western British Honduras (now

1881:. He placed six other names as full synonyms of this form, which has its 1851: 1520: 1512:, but Hanson and colleagues did not have data for other interior Mexican 1458:

were at the upper end of the range of variation in their large series of

1447: 847: 545: 536: 442: 347: 343: 177: 127: 6592:

Vega, R., Vázquez-Domínguez, E., Mejía-Puente, A. and Cuaro, A.D. 2004.

6125:

Descriptions of twelve new species and subspecies of mammals from Panama

6036: 1209:

was first described by Merriam in 1901 as a separate species similar to

999:

clade, which Hanson and colleagues proposed to recognize as the species

8413: 7721: 6633: 6579: 6561: 6529: 6313: 6285: 6243: 6179: 6161: 5935: 5874: 5827: 5802: 3484: 3321: 3175: 3097: 2984: 2909: 2057: 1975: 1945: 1898: 1348:) were described by Merriam in 1901, and Goldman had himself described 1345: 1181: 1137: 1071: 1063: 1036: 1011:, and according to Merriam (1901) have more robust skulls, with larger 992: 514: 6428: 6413: 6260: 6232: 6097:

Living on the edge: roads and edge effects on small mammal populations

5863: 3151:, a defining character of oryzomyines. There are some openings in the 454:) is the only member of its own group (unless western populations are 8426: 6765: 6759: 6747: 3672:(July 26); thus, the former is the original publication of this name. 3337: 3128: 2913: 2902: 2073: 1963: 1067: 1055: 1032: 819:) to study relationships among populations of the marsh rice rat and 808: 414: 310: 147: 137: 117: 8316: 6534:

species group) in southern Texas and northeastern Tamaulipas, Mexico

6488:

A Field Guide to the Mammals of Central America and Southeast Mexico

6050: 6049:

Engstrom, M.D., Schmidt, C.A., Morales, J.C. and Dowler, R.C. 1989.

5040:

Carleton and Musser, 1989, p. 24; Weksler, 2006, pp. 23–25

2952:

chromosomes, with a long and a very short arm, and one medium-sized

1492:, which occurs at over 2,000 m (6,600 ft) altitude in the 1450:

of the three forms in their morphometric analyses. The holotypes of

1195:, but usually with paler fur; the upperparts are more buffy than in 855:

clade, two populations from Panama and Costa Rica were successively

326: 6444:. Baltimore: The Johns Hopkins University Press, 2 vols., 2142 pp. 4526:

Goldman, 1918, pp. 39–40; Hanson et al., 2010, appendix A

3483:, with about 25 worms per rat, and two were infected by one or two 1054:

occurs along the Pacific coast from central Sonora to southeastern

1050:

As defined by Carleton and Arroyo-Cabrales in 2009, the subspecies

1019:(cheekbones), and better developed ridges along the margins of the 6634:

Morphological variation between Pleistocene and Recent samples of

6622: 6611: 6422:

Occasional Papers of the Museum of Zoology, University of Michigan

6237:

Occasional Papers of the Museum of Zoology, University of Michigan

5846:

Scientific Publications of the Cleveland Museum of Natural History

5727: 832: 769:, and Carleton and Arroyo-Cabrales wrote that further research on 620: 583: 370: 3304:, processes at the back of a vertebra, are absent from the fifth 3202:(one upper and one lower incisor and three upper and three lower 2204:

were first described by Goldman in 1912, who introduced the name

2034:

on the basis of its distinctly dark fur. In reviewing Nicaraguan

2030:, by Merriam in 1901, and Goldman retained it as a subspecies of 1799:

counties, Texas, and in far northeastern Tamaulipas, the two are

6623:

Ten new genera of oryzomyine rodents (Cricetidae: Sigmodontinae)

6399:

Catacamas virus, a hantaviral species naturally associated with

6216:

Hanson, J.D., Indorf, J.L., Swier, V.J. and Bradley, R.D. 2010.

3605: 3494: 3256:

On the first lower molar, the labial and lingual conules of the

2897:

on the hindfeet are reduced, as are the ungual tufts, and small

796: 479:, along with hundreds of other species of mainly small rodents. 8400: 8320: 6704: 6439: 5893:

complex (Rodentia: Cricetidae: Sigmodontinae) in Western Mexico

4113:

Hanson et al., 2010, fig. 2, table 1, appendix I

900:, the authors suggested that the four groups they found within 6506:(Rodentia: Muridae) with notes on taxonomy and natural history 6213:. Chicago: Field Museum of Natural History, xii + 861 pp. 5130:

Hooper and Musser, 1964, p. 13; Weksler, 2006, p. 57

4083:

Hanson et al., 2010, fig. 2, p. 343, appendix I

3288:

As usual in oryzomyines, there are twelve ribs. The first rib

2894: 614:

group were classified under the marsh rice rat, together with

5468:

Reid, 2009, p. 207; Hall and Dalquest, 1963, p. 287

530:, who had done much work on North American rodents. In 1893, 6302:. Princeton and Oxford: Princeton University Press, 240 pp. 6209:. pp. 753–795 in Wenzel, R.L. and Tipton, V.J. (eds.). 6079:Ácaros parásitos de roedores de Playa de Oro, Colima, México 6065:

Mites and ticks associated with some small mammals in Mexico

3668:

article was published earlier (July 19) than the one in the

3391:, which also swims well, but does not dive. When disturbed, 2991:

digits at its tip. The outer surface of the penis is mostly

1144:, in 1901, noting that it was smaller and more reddish than 6638:(Insectivora: Soricidae) from the Yucatan Peninsula, Mexico 2976:, but there are no consistent differences between the two. 1815:

in the wild. Compared to populations further to the south,

1434:

differentiation and offered some comments on the status of

1172:. Carleton and Arroyo-Cabrales wrote that it is similar to 1031:

sequence differences average 2.06% and western (Jalisco to

586:

and western Mexico south to Costa Rica. He placed it in an

6474:

A Field Guide to Mammals of North America, North of Mexico

6051:

Records of mammals from Isla Cozumel, Quintana Roo, Mexico

5936:

Late Pleistocene mammals from a cave in Tamaulipas, Mexico

3316:(small bones) are present with a spinous back border. The 3009:, located in the crater at the end of the penis, a fleshy 3005:

and the marsh rice rat among oryzomyines examined. On the

2080:

occurs up to an altitude of 1,250 m (4,100 ft).

574:

in 1918 and consolidated many forms into a single species

564:

group of species, which also included the marsh rice rat.

552:

to the animal from Guatemala, and introduced the new name

4143:

Carleton and Arroyo-Cabrales, 2009, pp. 118, 121–122

6490:. 2nd edition. Oxford University Press US, 346 pp. 6183:

only subspecifically different from the marsh rice rat,

5925:

Swimming ability in three Costa Rican dry forest rodents

4436:

Merriam, 1901a, p. 103; Merriam, 1901b, p. 280

3300:(chest) vertebrae, a synapomorphy of the Sigmodontinae. 3218:, with the cusps higher than the connecting crests, and 2308:

as a species, because they could not examine samples of

2232:. In 1937, Bole described another species of Panamanian 1484:

may represent no more than robust upland populations of

5810:

sp. n. (Coccidia: Eimeriidae) from the marsh rice rat,

3344:" for details. The species has also been found in late 2916:-hemiglandular): it is not split in two chambers by an 2893:(tufts of hair on the digits) are present. Many of the 321:. Weighing about 43 to 82 g (1.5 to 2.9 oz), 6578:

Vázquez, L.B., Medellín, R.A. and Cameron, G.N. 2000.

6284:

Jones, J.K. Jr., Carter, D.C. and Webster, W.D. 1983.

6244:

First South American record of Coues' marsh rice rat,

2338:

may also be discovered in the Pacific lowlands of the

2003:

sequence data, it falls among populations of nominate

1827:

form a separate group, but cluster among specimens of

1312:

from the interior plateaus of central Mexico together—

1237:(1961), Burt and Stirton recorded only the subspecies 6560:

Underwood, H.T., Owen, J.G. and Engstrom, M.D. 1986.

6160:

Haiduk, M.W., Bickham, J.W. and Schmidly, D.J. 1979.

5678:

Deardoff et al., 2009, p. 519; 2010, p. 350

3362:) and in Veracruz, it has even been found on the dry 2972:

has been used to distinguish the marsh rice rat from

737:

and Joaquin Arroyo-Cabrales reviewed western Mexican

6621:

Weksler, M., Percequillo, A.R. and Voss, R.S. 2006.

4373:

Carleton and Arroyo-Cabrales, 2009, pp. 113–114

4206:

Hanson et al., 2010, figs. 1–2, appendix I

3395:

will enter the water and swim away. It is primarily

3076:

bones do not extend back beyond the point where the

995:, western Mexico, east to El Salvador form a single 629:

and related species according to Goldman (1918): 1,

534:

wrote that the species, by then placed in the genus

8329: 8291: 8262: 8233: 8211: 8182: 8153: 8129: 8107: 8083: 8010: 7988: 7846: 7719: 7695: 7643: 7537: 7487: 7416: 7394: 7365: 7343: 7307: 7251: 7229: 7165: 7129: 7058: 7001: 6979: 6957: 6893: 6869: 6847: 6815: 6786: 6566:(Rodentia: Cricetidae) from San Luis Potosi, Mexico 6377:

Proceedings of the Biological Society of Washington

6144:

Proceedings of the Biological Society of Washington

6063:Estébanes-González, M.L. and Cervantes, F.A. 2005. 5857:

Proceedings of the Biological Society of Washington

5801:Barnard, W.P., Ernst, J.V. and Stevens, R.O. 1971. 4787:

Goldman, 1918, p. 31; Harris, 1943, p. 12

3666:

Proceedings of the Biological Society of Washington

3584:easily survive experimental infection with several 3467:in San Luis Potosí, five each were infected by the 2968:, with a long and a vestigial arm. The form of the 2330:, northwestern Colombia. The Colombian specimen is 2125:from Costa Rica have historically been referred to 884:specimens closer to the marsh rice rat than to the 6616:Bulletin of the American Museum of Natural History 6500:Sánchez H., J., Ochoa G., J. and Voss, R.S. 2001. 6476:. 4th ed. Houghton Mifflin Harcourt, 579 pp. 5904:Orders and families of Recent mammals of the world 5897:Bulletin of the American Museum of Natural History 5765:Bulletin of the American Museum of Natural History 5754:Bulletin of the American Museum of Natural History 5743:Bulletin of the American Museum of Natural History 5732:Bulletin of the American Museum of Natural History 5633:Estébanes-González and Cervantes, 2005, p. 27 5010:Goldman, 1918, p. 29; Reid, 2009, p. 206 4649:Murie, 1932, pp. 1–2; Murie, 1935, p. 26 1691:populations from Texas to Nicaragua form a single 6407:American Journal of Tropical Medicine and Hygiene 6392:Proceedings of the Washington Academy of Sciences 5969:American Journal of Tropical Medicine and Hygiene 4796:Hanson et al., 2010, p. 343, appendix A 4352:Hanson et al., 2010, fig. 1, appendix I 3977:Hanson et al., 2010, figs. 1–2, table 1 3670:Proceedings of the Washington Academy of Sciences 3422:Mammals of Central America & Southeast Mexico 3385:from that of the ecologically similar cotton rat 2912:has the characteristic pattern of sigmodontines ( 1803:(occur in the same places). In the contact zone, 1184:may also belong to this form. Goldman wrote that 401:, have been reinstated as species. Nevertheless, 5828:Ecological and taxonomic notes on the rice rat ( 5528:Murie, 1935, p. 26; Reid, 2009, p. 207 4910:Carleton and Arroyo-Cabrales, 2009, table 2 3230:on the lower molars, are present, another trait 2956:pair, with one arm shorter than the other. The 1225:. Three specimens from central El Salvador have 5779:Proceedings of the Zoological Society of London 4400:Carleton and Arroyo-Cabrales, 2009, p. 114 4343:Carleton and Arroyo-Cabrales, 2009, p. 113 4334:Carleton and Arroyo-Cabrales, 2009, p. 118 4134:Carleton and Arroyo-Cabrales, 2009, p. 120 4122:Carleton and Arroyo-Cabrales, 2009, p. 119 3956:Carleton and Arroyo-Cabrales, 2009, p. 107 3947:Carleton and Arroyo-Cabrales, 2009, p. 116 3836:Carleton and Arroyo-Cabrales, 2009, p. 117 3648:Proceedings of the Zoological Society of London 2318:was first reported from Colombia in 1987, when 1707:) and an island form he retained as a species ( 6530:Genic variation and systematics of rice rats ( 5923:Cook, W.M., Timm, R.M. and Hyman, D.E. 2001. 4508:Hanson et al., 2010, fig. 1, table 1 3938:Carleton and Arroyo-Cabrales, 2009, p. 94 2932:(shared-derived character) of Oryzomyini. The 2284:to subspecies of the marsh rice rat (in which 1994:was inside the range of variation of mainland 1838:is known from further south in Mexico, in the 1787:meets that of the marsh rice rat; in parts of 6716: 6467:Bulletin of the United States National Museum 6314:The zoogeographical limits of the West Indies 5906:. John Wiley and Sons, New York, 686 pp. 5887:Carleton, M.D. and Arroyo-Cabrales, J. 2009. 5651:Estébanes-González et al., 2011, table 1 3708:Head and body length instead of total length. 3592:, suggesting that the species may serve as a 1779:, which was described as a separate species, 1023:of the skull (between the eyes). Within the " 618:from Panama, described as a species in 1937. 8: 6373:Six new mammals from Cozumel Island, Yucatan 5498:Fuentes-Montemayor et al., 2009, p. 865 5031:Weksler, 2006, pp. 19, 23, table 5 4553:Schmidt and Engstrom, 1994, pp. 915–916 4544:Schmidt and Engstrom, 1994, pp. 916–917 3973: 3971: 3264:. The second lower molar bears a crest, the 2920:and the front part (antrum) is covered by a 2363:, but "Coues' Oryzomys" has also been used. 1711:). Two other subspecies Goldman recognized, 851:(7.48% and 7.52%, respectively). Within the 6429:Mammals from Guatemala and British Honduras 6041:Journal of the Kansas Entomological Society 4015: 4013: 3664:as new in both of his 1901 papers, but his 3572:in western Mexico—which are related to the 2248:. Although Goldman recommended to him that 1775:, Mexico, are classified as the subspecies 801:interphotoreceptor retinoid-binding protein 27:Semiaquatic rodent in the family Cricetidae 8317: 6723: 6709: 6701: 5575: 5573: 5506: 5504: 4610:Dalquest and Roth, 1970, pp. 220, 226 3560:in Mexico. The species is infected by two 3260:, the frontmost cusp, are separated by an 1966:); it is smaller and darker than nominate 1420:, Carleton and Arroyo-Cabrales classified 361:consolidated most into the single species 333:of hair around the digits. It has 56 226: 69: 50: 31: 6109:Genoways, H.H. and Jones, J.K. Jr. 1971. 5992:New Haven: A & C Black, 312 pp. 5961:Short report: Experimental infections of 5563: 5561: 5485: 5483: 4997: 4995: 4993: 4991: 4989: 4987: 4906: 4904: 4745:Jones and Engstrom, 1986, pp. 10, 12 4130: 4128: 4100: 4098: 3985: 3983: 3832: 3830: 2372:Measurements of different populations of 2038:in 1986, Jones and Engstrom did not keep 1771:, those in southernmost Texas and nearby 6357:Medellín, X.L. and Medellín, R.A. 2006. 6030:Field Columbian Museum Zoological Series 6019:Field Columbian Museum Zoological Series 5018: 5016: 5001:Medellín and Medellín, 2006, p. 710 4968: 4966: 4964: 4831: 4829: 4810: 4808: 4806: 4804: 4802: 4762: 4760: 4360: 4358: 3761: 3759: 3757: 3755: 3753: 3751: 3749: 3747: 3336:extends from southern Texas and central 2370: 2087: 1889:Allen and Chapman, 1897, from Veracruz; 1546: 1416:In their 2009 review of western Mexican 1248: 919: 761:from interior Mexico as species. Still, 6528:Schmidt, C.A. and Engstrom, M.D. 1994. 6286:Records of mammals from Hidalgo, Mexico 6277:Jones, J.K. Jr. and Lawlor, T.E. 1965. 5441:Weksler, 2006, p. 53; fig. 28 5423:Weksler, 2006, p. 52, table 5 5392: 5390: 5371: 5369: 5341: 5339: 5337: 5304:Weksler, 2006, p. 41, table 5 5259:Weksler, 2006, p. 38, table 5 5205:Weksler, 2006, p. 28, table 5 5196:Schmidt and Engstrom, 1994, p. 917 5187:Weksler, 2006, p. 32, table 5 5174: 5172: 5157:Weksler, 2006, p. 27, table 5 5058:Weksler, 2006, p. 17, table 5 4945: 4943: 4714: 4712: 4679:Schmidt and Engstrom, 1989, p. 414 4562:Schmidt and Engstrom, 1994, p. 920 4535:Schmidt and Engstrom, 1994, p. 914 4321: 4319: 4317: 4315: 4313: 4247: 4245: 4235: 4233: 4223: 4221: 4181: 4179: 4070: 4068: 4066: 4038: 4036: 4034: 3765:Musser and Carleton, 2005, p. 1147 3720: 3638: 2979:As is characteristic of Sigmodontinae, 2885:the moderately large eyes show reddish 1504:. A specimen from inland Michoacán has 835:sister to the marsh rice rat; the mean 446:. The eighth species of the genus, the 6555:Annals and Magazine of Natural History 6222:complex: evidence for multiple species 5965:with sympatric arboviruses from Mexico 5826:Benson, D.E. and Gehlbach, F.R. 1979. 5705:Benson and Gehlbach, 1979, p. 228 5669:Chu et al., 2008; Milazzo et al., 2006 5597:Benson and Gehlbach, 1979, p. 226 5555:Benson and Gehlbach, 1979, p. 227 5331:Carleton and Musser, 1984, p. 292 4774: 4772: 4657: 4655: 4618: 4616: 4570: 4568: 4294: 4292: 4282: 4280: 4261: 4259: 4257: 4193: 4191: 4169: 4167: 3998:Hanson et al., 2010, figs. 1, 3–4 3811: 3809: 3807: 3805: 3795: 3793: 3791: 3789: 3728: 3726: 3724: 1098:and has been recognized as a distinct 498:, placing it in the now-defunct genus 6418:from a pine ridge in British Honduras 6129:Smithsonian Miscellaneous Collections 5714:Cameron and Scheel, 2001, p. 676 5696:Schmidly and Davis, 2004, p. 381 4736:Genoways and Jones, 1971, p. 834 4019:Hanson et al., 2010, pp. 342–343 2072:, the gap behind the end of the bony 1866:, but has a skull similar to that of 1058:and inland along rivers into central 827:data placed all studied specimens of 717:were resurrected as separate species— 373:with its United States relative, the 7: 6461:Poole, A.J. and Schantz, V.S. 1942. 6259:Hooper, E.T. and Musser, G.G. 1964. 6198:Hall, E.R. and Dalquest, W.W. 1963. 5759:Allen, J.A. and Chapman, F.M. 1897. 5567:Hall and Dalquest, 1963, p. 288 5489:Hall and Dalquest, 1963, p. 287 4972:Jones and Engstrom, 1986, p. 12 4727:Jones and Engstrom, 1986, p. 10 4463:Allen and Chapman, 1897, p. 206 4445:Poole and Schantz, 1942, p. 306 3590:Venezuelan equine encephalitis virus 2288:was included at the time), and when 1168:, and has a shorter tail and weaker 1003:. These animals differ by 4.4% from 8604:Taxa named by Edward Richard Alston 8569:IUCN Red List least concern species 6338:IUCN Red List of Threatened Species 5862:Burt, W.H. and Stirton, R.A. 1961. 4754:Jones and Engstrom, 1986, p. 7 4670:Jones and Lawlor, 1965, p. 413 1426:as a separate species from lowland 1308:Goldman grouped four subspecies of 1148:. Goldman synonymized the three as 908:, the western group would be named 6562:Endohelminths of three species of 6298:Kays, R.W. and Wilson, D.E. 2000. 6207:Checklist of the mammals of Panama 6069:International Journal of Acarology 5990:Mammals of the World: A checklist. 5934:Dalquest, W.W. and Roth, E. 1970. 5660:Barnard et al., 1971, p. 1294 5588:Vázquez et al., 2000, table 1 5139:Hooper and Musser, 1964, p. 7 4919:Burt and Stirton, 1961, p. 60 4215:Burt and Stirton, 1961, p. 61 2964:, with a long and a short arm, or 2052:with gray underparts, occurs with 1364:is large, dark and large-toothed, 25: 6388:) of the United States and Mexico 6384:Synopsis of the rice rats (genus 6233:A list of mammals from Costa Rica 5851:Burt, W.H. 1934. A new rice rat ( 5750:Additional mammals from Nicaragua 4092:Hanson et al., 2010, table 1 3902:Goldman, 1918, pp. 16, 28–29 2826:Costa Rica, Panama, and Colombia 2256:both be treated as subspecies of 876:as separate clades. In addition, 741:, reaffirmed the distinctness of 602:meet in southern Texas. In 1960, 6686: 6672: 6650:Woodman, N. and Croft, D. 2005. 6218:Molecular divergence within the 6136:Five new rice rats of the genus 5094:Haiduk et al., 1979, p. 612 4592:Hanson et al., 2010, fig. 2 4364:Hanson et al., 2010, fig. 1 4104:Hanson et al., 2010, p. 343 4007:Hanson et al., 2010, fig. 5 3989:Hanson et al., 2010, p. 341 3965:Hanson et al., 2010, p. 337 3845:Hanson et al., 2010, p. 342 3741:Hanson et al., 2010, p. 336 3044: 3035: 2169: 2160: 2084:Costa Rica, Panama, and Colombia 1930: 1921: 1909:Eliot, 1904, from Veracruz; and 1767:The northernmost populations of 1739: 1730: 1388: 1379: 512:) and two species now placed in 504:, and noted similarities to the 413:and two additional species from 94: 8589:Fauna of the Rio Grande valleys 6166:from Mexico and Central America 5796:Mammals from Tamaulipas, Mexico 4697:Platt et al., 2000, p. 167 3580:hantaviruses, however. Chiapas 3308:. Between the second and third 1462:from the western lowlands, and 1356:in 1915. According to Goldman, 6151:The rice rats of North America 5988:Duff, A. and Lawson, A. 2004. 5579:Vega et al., 2004, p. 218 5510:Vega et al., 2004, p. 217 5049:Weksler, 2006, pp. 79, 81 4871:Hershkovitz, 1987, p. 154 4862:Hershkovitz, 1987, p. 152 4688:Vega et al., 2004, p. 210 3206:on each side of the jaws), as 2983:has a complex penis, with the 2399:Western Mexico to El Salvador 2322:reported on its occurrence at 2200:is rare in Panama. Panamanian 1905:Merriam, 1901, from Veracruz; 1893:Merriam, 1901, from Veracruz; 1372:is large and relatively pale. 1229:sequences similar to those of 239:(in red) and other species of 1: 6403:(Coues' oryzomys) in Honduras 6162:Karyotypes of six species of 5873:Cameron, G.N. and Scheel, D. 5786:Biologia centrali-americana. 5432:Weksler, 2006, pp. 52–53 5384:Weksler, 2006, pp. 44–45 5363:Weksler, 2006, pp. 44–49 5354:Weksler, 2006, pp. 43–44 5322:Weksler, 2006, pp. 41–42 5295:Weksler, 2006, pp. 40–41 5277:Weksler, 2006, pp. 38–39 5232:Weksler, 2006, pp. 34–35 5166:Weksler, 2006, pp. 31–32 5121:Weksler, 2006, pp. 56–57 5112:Weksler, 2006, pp. 55–56 5085:Weksler, 2006, pp. 58–59 4937:Goldman, 1918, pp. 30–31 4889:Goldman, 1918, pp. 29–43 4880:Eliot, 1905, pp. 182–186 4823:Goldman, 1918, pp. 42–43 4631:Goldman, 1918, pp. 29–30 3854:Weksler et al., 2006, table 1 3783:Dickinson, 2005, table 1 3681:Number of specimens measured. 3341: 2987:(penis bone) ending in three 2404:San José de Guaymas, Sonora ( 2068:(openings in the roof of the 1783:, in 1891. Here the range of 916:Western Mexico to El Salvador 526:, honors American naturalist 6456:Caribbean Journal of Science 6008:Caribbean Journal of Science 5954:Emerging Infectious Diseases 5929:Revista de Biología Tropical 3911:Hall, 1960, pp. 172–173 3774:Dickinson, 2005, p. 427 3268:, before the two cusps, the 3226:on the upper molars and the 2641:Rio Verde, San Luis Potosí ( 2296:differ by about 7% in their 2015:off the coast of Belize and 1466:clustered with specimens of 1106:occurs close to three other 560:united many of those into a 6570:The Southwestern Naturalist 6290:The Southwestern Naturalist 6190:The Southwestern Naturalist 6092:27(1):169–176 (in Spanish). 6055:The Southwestern Naturalist 5940:The Southwestern Naturalist 5818:The Journal of Parasitology 5791:. R.H. Porter, 220 pp. 5615:Eckerlin, 2005, p. 155 4499:Merriam, 1901b, p. 284 4472:Merriam, 1901b, p. 283 4454:Merriam, 1901b, p. 288 4427:Merriam, 1901a, p. 103 4325:Goldman, 1918, plate I 4274:Merriam, 1901b, p. 279 4239:Merriam, 1901b, p. 281 4227:Merriam, 1901b, p. 282 4185:Merriam, 1901b, p. 286 4074:Merriam, 1901b, p. 285 4060:Merriam, 1901b, p. 287 3893:Merriam, 1901b, p. 275 3373:lives on the ground and is 3117:posterolateral palatal pits 1621:Oryzomys mexicanus peragrus 1360:is pale and large-toothed, 639:O. c. zygomaticus 8620: 8574:Rodents of Central America 5864:The mammals of El Salvador 5642:Emerson, 1971, p. 334 4853:Handley, 1966, p. 781 4706:Koopman, 1959, p. 237 4391:Goldman, 1918, p. 282 4251:Goldman, 1915, p. 129 3920:Goldman, 1918, fig. 3 3499:Eubrachylaelaps circularis 2809:Isla de Omotepe, Nicaragua 1907:Oryzomys jalapae apatelius 1823:sequences of specimens of 1558:Oryzomys jalapae apatelius 1496:, was the same species as 1235:The Mammals of El Salvador 701:Additional studies of the 660:O. c. albiventer 582:distributed from southern 8599:Mammals described in 1877 6742: 6627:American Museum Novitates 6596:Oryzomys couesi cozumelae 6101:Journal of Animal Ecology 5519:Alston, 1877, p. 757 5450:Weksler, 2006, p. 54 5414:Weksler, 2006, p. 53 5405:Weksler, 2006, p. 52 5396:Weksler, 2006, p. 49 5375:Weksler, 2006, p. 44 5345:Weksler, 2006, p. 43 5313:Weksler, 2006, p. 42 5286:Weksler, 2006, p. 40 5268:Weksler, 2006, p. 37 5241:Weksler, 2006, p. 35 5223:Weksler, 2006, p. 31 5214:Weksler, 2006, p. 30 5178:Weksler, 2006, p. 32 5148:Weksler, 2006, p. 57 5076:Weksler, 2006, p. 59 4981:Goldman, 1918, p. 42 4958:Murie, 1932, table I 4949:Goldman, 1918, p. 30 4928:Goldman, 1918, p. 41 4718:Goldman, 1918, p. 32 4661:Goldman, 1918, p. 43 4640:Goldman, 1918, p. 31 4622:Goldman, 1918, p. 39 4574:Goldman, 1918, p. 40 4382:Thomas, 1893, p. 404 4307:Goldman, 1918, p. 38 4298:Goldman, 1918, p. 36 4286:Goldman, 1918, p. 35 4265:Goldman, 1918, p. 37 4197:Goldman, 1918, p. 33 4173:Goldman, 1918, p. 34 3872:Musser and Carleton, 2005 3815:Thomas, 1893, p. 403 3799:Alston, 1877, p. 756 3542:Prolistrophorus frontalis 3463:from Cozumel. Out of ten 3210:. The upper incisors are 2872: 2825: 2614: 2525: 2398: 2224:(part of the roof of the 1860:O. c. aquaticus 1713:O. c. richmondi 1701:O. c. aquaticus 1474:Baja California Peninsula 1260:Oryzomys crinitus aztecus 1223:O. c. mexicanus 1104:O. c. mexicanus 1052:Oryzomys couesi mexicanus 753:Baja California Peninsula 669:O. c. aquaticus 643:O. c. mexicanus 635:O. c. richmondi 399:Baja California Peninsula 278:Oryzomys palustris couesi 254: 247: 234: 225: 206: 199: 91:Scientific classification 89: 67: 58: 49: 34: 6300:Mammals of North America 6205:Handley, C.O. Jr. 1966. 4778:Goldman, 1912, p. 7 4583:Baker, 1951, p. 215 4418:Allen, 1891, p. 289 4409:Eliot, 1904, p. 266 3863:Weksler, 2006, p. 3 3690:Only averages available. 3530:Echinonyssus microchelae 3521:in Nicaragua, the mites 3474:Hassalstrongylus musculi 2788:Southeastern Nicaragua ( 2767:San Antonio, Nicaragua ( 2137:sequence divergence) as 2066:sphenopalatine vacuities 1960:Oryzomys couesi pinicola 1891:Oryzomys jalapae rufinus 1721:O. c. pinicola 1717:O. c. peragrus 1662:Oryzomys jalapae rufinus 1632:Oryzomys couesi pinicola 1293:Oryzomys couesi regillus 1150:O. couesi mexicanus 868:also placed examples of 665:O. c. peragrus 655:O. c. regillus 651:O. c. crinitus 6646:(subscription required) 6606:(subscription required) 6602:Biological Conservation 6588:(subscription required) 6574:(subscription required) 6542:(subscription required) 6514:(subscription required) 6322:(subscription required) 6294:(subscription required) 6255:(subscription required) 6231:Harris, W.P. Jr. 1943. 6211:Ectoparasites of Panama 6200:The mammals of Veracruz 6194:(subscription required) 6174:(subscription required) 6105:(subscription required) 6090:Acta Zoológica Mexicana 6073:(subscription required) 6059:(subscription required) 6045:(subscription required) 5984:(subscription required) 5944:(subscription required) 5919:(subscription required) 5883:(subscription required) 5855:) from Sonora, Mexico. 5840:(subscription required) 5822:(subscription required) 5022:Reid, 2009, p. 206 4844:Bole, 1937, p. 167 4835:Bole, 1937, p. 166 4814:Reid, 2009, p. 207 4766:Bole, 1937, p. 165 4490:Allen, 1910, p. 99 4161:Burt, 1934, p. 108 4051:Allen, 1897, p. 52 4042:Burt, 1934, p. 107 4028:Allen, 1897, p. 53 3824:Hall, 1960, p. 173 3397:active during the night 3383:differentiate its niche 3208:usual in muroid rodents 3020:Holochilus brasiliensis 2993:covered by small spines 2354:have been proposed for 1990:trivial and found that 1978:as a separate species, 1508:data characteristic of 1368:is large and dark, and 898:genetic species concept 733:from Jamaica. In 2009, 647:O. c. aztecus 570:revised North American 568:Edward Alphonso Goldman 458:as a separate species, 440:group within the genus 359:Edward Alphonso Goldman 6548:Notes on some Mexican 6242:Hershkovitz, P. 1987. 5974:Dickinson, E.C. 2005. 5772:On two new species of 5624:Underwood et al., 1986 4481:Murie, 1932, p. 1 3548:Prolistrophorus bakeri 3518:Hoplopleura oryzomydis 3318:entepicondylar foramen 3131:in the head is highly 3123:, an extension of the 3094:sphenopalatine foramen 2127:O. c. couesi 1879:Oryzomys couesi couesi 1864:O. c. couesi 1829:O. c. couesi 1705:O. c. couesi 1476:. They suggested that 1430:on the basis of clear 1219:O. c. couesi 1102:zone in some reviews. 727:, western Mexico, and 698: 631:Oryzomys couesi couesi 8539:Paleobiology Database 6382:Merriam, C.H. 1901b. 6371:Merriam, C.H. 1901a. 6002:Eckerlin, R.P. 2005. 3505:Gigantolaelaps boneti 3278:anterolabial cingulum 3262:anteromedian fossette 3137:subsquamosal fenestra 2342:in western Colombia. 2300:sequences from other 1162:Oryzomys couesi lambi 1095:Osgoodomys banderanus 942:Oryzomys couesi lambi 813:alcohol dehydrogenase 624: 40:Temporal range: Late 18:Coues's rice rat 8304:Ekbletomys hypenemus 6838:A. praeuniversitatis 6683:at Wikimedia Commons 6642:Journal of Mammalogy 6584:Journal of Mammalogy 6538:Journal of Mammalogy 6345:: e.T15592A115128044 6318:Journal of Mammalogy 6312:Koopman, K.F. 1959. 6251:Journal of Mammalogy 6226:Journal of Mammalogy 6170:Journal of Mammalogy 6155:North American Fauna 6149:Goldman, E.A. 1918. 6134:Goldman, E.A. 1915. 6123:Goldman, E.A. 1912. 6118:Journal of Mammalogy 6035:Emerson, K.C. 1971. 5879:Journal of Mammalogy 5836:Journal of Mammalogy 5770:Alston, E.R. 1877 . 3536:Ornithonyssus bacoti 3408:Population densities 3332:The distribution of 3328:Ecology and behavior 3284:Postcranial skeleton 2922:glandular epithelium 2620:Brownsville, Texas ( 2425:Escuinapa, Sinaloa ( 2208:for a specimen from 1911:Oryzomys richardsoni 1897:Merriam, 1901, from 1831:from further south. 1640:Oryzomys richardsoni 1472:from the tip of the 1207:Oryzomys zygomaticus 1188:was very similar to 1089:Peromyscus perfulvus 1077:Sigmodon mascotensis 972:Oryzomys zygomaticus 751:from the tip of the 558:Clinton Hart Merriam 510:Hesperomys palustris 8579:Mammals of Colombia 6140:from Middle America 6113:Oryzomys dimidiatus 6111:Second specimen of 5784:Alston, E.R. 1882. 3732:Linzey et al., 2016 3622:Habitat destruction 3600:Conservation status 3551:in Colima, and the 3508:have been found on 3292:with both the last 3247:anteromedian flexus 2662:Orizaba, Veracruz ( 2615:Texas to Nicaragua 2488:Nenton, Guatemala ( 2376: 2238:Oryzomys azuerensis 2206:Oryzomys gatunensis 2107:Oryzomys gatunensis 2099:Oryzomys azuerensis 2092: 2070:mesopterygoid fossa 2045:Oryzomys dimidiatus 1875:nominate subspecies 1850:, and far northern 1551: 1423:Oryzomys albiventer 1253: 1021:interorbital region 924: 730:Oryzomys antillarum 616:Oryzomys azuerensis 562:palustris-mexicanus 394:Oryzomys peninsulae 388:Oryzomys antillarum 61:Conservation status 6632:Woodman, N. 1995. 6610:Weksler, M. 2006. 6532:Oryzomys palustris 6220:Oryzomys palustris 6185:Oryzomys palustris 6084:2016-03-03 at the 6024:Eliot, D.G. 1905. 6013:Eliot, D.G. 1904. 5980:Journal of Zoology 5931:49(3–4):1177–1181. 5812:Oryzomys palustris 5794:Baker, R.H. 1951. 5748:Allen, J.A. 1910. 5737:Allen, J.A. 1897. 5726:Allen, J.A. 1891. 3662:Oryzomys cozumelae 3660:Merriam described 3570:Playa de Oro virus 3524:Laelaps oryzomydis 3324:(upper arm bone). 3147:, the roof of the 3108:meet extensively. 2942:fundamental number 2918:incisura angularis 2746:Yaruca, Honduras ( 2683:Tumbala, Chiapas ( 2371: 2328:Córdoba Department 2320:Philip Hershkovitz 2222:interparietal bone 2090:Taxonomic synonyms 2088: 2028:Oryzomys richmondi 1980:Oryzomys cozumelae 1895:Oryzomys teapensis 1781:Oryzomys aquaticus 1673:Oryzomys teapensis 1651:Oryzomys richmondi 1588:Oryzomys cozumelae 1566:Oryzomys aquaticus 1549:Taxonomic synonyms 1547: 1543:Texas to Nicaragua 1469:O. peninsulae 1328:. Three of those ( 1251:Taxonomic synonyms 1249: 1130:Oryzomys mexicanus 1116:O. peninsulae 1112:O. albiventer 1025:Oryzomys mexicanus 1001:Oryzomys mexicanus 950:Oryzomys mexicanus 922:Taxonomic synonyms 920: 910:Oryzomys mexicanus 758:O. albiventer 748:O. peninsulae 699: 688:O. peninsulae 682:O. antillarum 673:O. gatunensis 588:Oryzomys palustris 8584:Rodents of Mexico 8554: 8553: 8526:Open Tree of Life 8323:Taxon identifiers 8314: 8313: 6736: 6731:Species of tribe 6677:Media related to 6660:Fieldiana Geology 6546:Thomas, O. 1893. 6524:978-0-292-70241-7 6496:978-0-19-534322-9 6482:978-0-395-93596-5 6450:978-0-8018-8221-0 6367:978-970-9000-30-6 6224:(abstract only). 6178:Hall, E.R. 1960. 5820:57(6):1293–1296. 5808:Isospora hammondi 5804:Eimeria palustris 5537:Cook et al., 2001 3456:Polygenis odiosus 3388:Sigmodon hirsutus 3296:(neck) and first 3168:masseteric ridges 3162:(lower jaw), the 3121:alisphenoid strut 3113:incisive foramina 2999:shares only with 2936:includes 56 2899:interdigital webs 2879: 2878: 2875: 2704:El Cayo, Belize ( 2260:, Bole described 2242:Veraguas Province 2120: 2119: 2114: 2103: 2011:is also found on 1974:of the island of 1903:Oryzomys goldmani 1709:O. cozumelae 1686: 1685: 1680: 1669: 1658: 1647: 1636: 1628: 1617: 1616:and Chapman, 1897 1606: 1599:Oryzomys goldmani 1595: 1584: 1577:Hesperomys couesi 1573: 1562: 1306: 1305: 1300: 1289: 1278: 1271:Oryzomys crinitus 1267: 985: 984: 979: 968: 957: 946: 938: 745:, and reinstated 715:O. palustris 677:O. cozumelae 612:O. palustris 496:Hesperomys couesi 452:O. palustris 391:from Jamaica and 379:O. palustris 295: 294: 286: 274: 262: 258:Hesperomys couesi 84: 16:(Redirected from 8611: 8547: 8546: 8534: 8533: 8521: 8520: 8508: 8507: 8495: 8494: 8482: 8481: 8469: 8468: 8456: 8455: 8443: 8442: 8430: 8429: 8417: 8416: 8404: 8403: 8391: 8390: 8378: 8377: 8365: 8364: 8363: 8350: 8349: 8348: 8318: 7922:O. longicaudatus 7346:Microakodontomys 6970:D. albimaculatus 6920:C. maracajuensis 6799:A. galapagoensis 6734: 6725: 6718: 6711: 6702: 6691:Data related to 6690: 6676: 6647: 6607: 6589: 6575: 6543: 6515: 6504:Oryzomys gorgasi 6458:36(1–2):166–168. 6427:Murie, A. 1935. 6412:Murie, A. 1932. 6409:75(5):1003–1010. 6354: 6352: 6350: 6323: 6295: 6256: 6195: 6175: 6106: 6074: 6060: 6046: 5985: 5982:266(4):427–430. 5945: 5920: 5884: 5841: 5823: 5781:1876(4):755–757. 5721:Literature cited 5715: 5712: 5706: 5703: 5697: 5694: 5688: 5685: 5679: 5676: 5670: 5667: 5661: 5658: 5652: 5649: 5643: 5640: 5634: 5631: 5625: 5622: 5616: 5613: 5607: 5604: 5598: 5595: 5589: 5586: 5580: 5577: 5568: 5565: 5556: 5553: 5547: 5544: 5538: 5535: 5529: 5526: 5520: 5517: 5511: 5508: 5499: 5496: 5490: 5487: 5478: 5475: 5469: 5466: 5460: 5457: 5451: 5448: 5442: 5439: 5433: 5430: 5424: 5421: 5415: 5412: 5406: 5403: 5397: 5394: 5385: 5382: 5376: 5373: 5364: 5361: 5355: 5352: 5346: 5343: 5332: 5329: 5323: 5320: 5314: 5311: 5305: 5302: 5296: 5293: 5287: 5284: 5278: 5275: 5269: 5266: 5260: 5257: 5251: 5248: 5242: 5239: 5233: 5230: 5224: 5221: 5215: 5212: 5206: 5203: 5197: 5194: 5188: 5185: 5179: 5176: 5167: 5164: 5158: 5155: 5149: 5146: 5140: 5137: 5131: 5128: 5122: 5119: 5113: 5110: 5104: 5101: 5095: 5092: 5086: 5083: 5077: 5074: 5068: 5065: 5059: 5056: 5050: 5047: 5041: 5038: 5032: 5029: 5023: 5020: 5011: 5008: 5002: 4999: 4982: 4979: 4973: 4970: 4959: 4956: 4950: 4947: 4938: 4935: 4929: 4926: 4920: 4917: 4911: 4908: 4899: 4896: 4890: 4887: 4881: 4878: 4872: 4869: 4863: 4860: 4854: 4851: 4845: 4842: 4836: 4833: 4824: 4821: 4815: 4812: 4797: 4794: 4788: 4785: 4779: 4776: 4767: 4764: 4755: 4752: 4746: 4743: 4737: 4734: 4728: 4725: 4719: 4716: 4707: 4704: 4698: 4695: 4689: 4686: 4680: 4677: 4671: 4668: 4662: 4659: 4650: 4647: 4641: 4638: 4632: 4629: 4623: 4620: 4611: 4608: 4602: 4599: 4593: 4590: 4584: 4581: 4575: 4572: 4563: 4560: 4554: 4551: 4545: 4542: 4536: 4533: 4527: 4524: 4518: 4515: 4509: 4506: 4500: 4497: 4491: 4488: 4482: 4479: 4473: 4470: 4464: 4461: 4455: 4452: 4446: 4443: 4437: 4434: 4428: 4425: 4419: 4416: 4410: 4407: 4401: 4398: 4392: 4389: 4383: 4380: 4374: 4371: 4365: 4362: 4353: 4350: 4344: 4341: 4335: 4332: 4326: 4323: 4308: 4305: 4299: 4296: 4287: 4284: 4275: 4272: 4266: 4263: 4252: 4249: 4240: 4237: 4228: 4225: 4216: 4213: 4207: 4204: 4198: 4195: 4186: 4183: 4174: 4171: 4162: 4159: 4153: 4150: 4144: 4141: 4135: 4132: 4123: 4120: 4114: 4111: 4105: 4102: 4093: 4090: 4084: 4081: 4075: 4072: 4061: 4058: 4052: 4049: 4043: 4040: 4029: 4026: 4020: 4017: 4008: 4005: 3999: 3996: 3990: 3987: 3978: 3975: 3966: 3963: 3957: 3954: 3948: 3945: 3939: 3936: 3930: 3927: 3921: 3918: 3912: 3909: 3903: 3900: 3894: 3891: 3885: 3879: 3873: 3870: 3864: 3861: 3855: 3852: 3846: 3843: 3837: 3834: 3825: 3822: 3816: 3813: 3800: 3797: 3784: 3781: 3775: 3772: 3766: 3763: 3742: 3739: 3733: 3730: 3709: 3706: 3700: 3697: 3691: 3688: 3682: 3679: 3673: 3658: 3652: 3643: 3596:for that virus. 3588:, including the 3568:in Honduras and 3480:H. bocqueti 3459:was found on an 3310:caudal vertebrae 3249:). A crest, the 3201: 3199: 3198: 3195: 3172:capsular process 3125:alisphenoid bone 3048: 3039: 3015:subapical lobule 3007:urethral process 2873: 2831:Azuero, Panama ( 2526:Interior Mexico 2377: 2228:), and the long 2173: 2164: 2109: 2101: 2093: 2048:, a small, dark 1934: 1925: 1887:Oryzomys jalapae 1743: 1734: 1675: 1664: 1653: 1642: 1634: 1623: 1612: 1610:Oryzomys jalapae 1601: 1590: 1579: 1568: 1560: 1552: 1525:Oryzomys fulgens 1494:Valley of Mexico 1446:, including the 1392: 1383: 1338:Distrito Federal 1295: 1284: 1282:Oryzomys fulgens 1273: 1262: 1254: 1221:but darker than 1134:Oryzomys bulleri 1128:first described 1126:Joel Asaph Allen 1043:in the west and 1017:zygomatic arches 974: 963: 952: 944: 933: 931:Oryzomys bulleri 925: 837:genetic distance 735:Michael Carleton 720:Oryzomys nelsoni 703:palustris–couesi 625:Distribution of 554:Oryzomys fulgens 490:first described 460:O. texensis 304:Coues's rice rat 302:, also known as 281: 269: 260: 235:Distribution of 230: 212: 99: 98: 78: 73: 72: 54: 32: 21: 8619: 8618: 8614: 8613: 8612: 8610: 8609: 8608: 8559: 8558: 8555: 8550: 8542: 8537: 8529: 8524: 8516: 8511: 8503: 8498: 8490: 8485: 8477: 8472: 8464: 8459: 8451: 8446: 8438: 8433: 8425: 8420: 8412: 8407: 8399: 8394: 8386: 8381: 8373: 8368: 8361:Oryzomys couesi 8359: 8358: 8353: 8344: 8343: 8338: 8331:Oryzomys couesi 8325: 8315: 8310: 8287: 8258: 8229: 8207: 8178: 8149: 8125: 8103: 8079: 8006: 7984: 7929:O. magellanicus 7842: 7830:O. sydandersoni 7715: 7691: 7639: 7599:N. maculiventer 7533: 7483: 7412: 7390: 7361: 7356:M. transitorius 7339: 7303: 7247: 7225: 7192:H. megacephalus 7161: 7142:H. brasiliensis 7125: 7054: 6997: 6975: 6953: 6889: 6865: 6843: 6811: 6782: 6738: 6729: 6694:Oryzomys couesi 6680:Oryzomys couesi 6669: 6645: 6644:76(1):223–231. 6605: 6587: 6573: 6572:31(3):410–411. 6541: 6540:75(4):914–928. 6513: 6502:Rediscovery of 6486:Reid, F. 2009. 6472:Reid, F. 2006. 6401:Oryzomys couesi 6359:Oryzomys couesi 6348: 6346: 6331:Oryzomys couesi 6326: 6321: 6320:40(2):236–240. 6293: 6292:28(3):378–380. 6254: 6253:68(1):152–154. 6246:Oryzomys couesi 6193: 6181:Oryzomys couesi 6173: 6172:60(3):610–615. 6104: 6086:Wayback Machine 6072: 6058: 6057:34(3):413–415. 6044: 6043:44(3):332–334. 5983: 5963:Oryzomys couesi 5943: 5942:15(2):217–230. 5918: 5891:Oryzomys couesi 5882: 5881:82(3):652–680. 5839: 5838:60(1):225–228. 5830:Oryzomys couesi 5821: 5723: 5718: 5713: 5709: 5704: 5700: 5695: 5691: 5686: 5682: 5677: 5673: 5668: 5664: 5659: 5655: 5650: 5646: 5641: 5637: 5632: 5628: 5623: 5619: 5614: 5610: 5605: 5601: 5596: 5592: 5587: 5583: 5578: 5571: 5566: 5559: 5554: 5550: 5545: 5541: 5536: 5532: 5527: 5523: 5518: 5514: 5509: 5502: 5497: 5493: 5488: 5481: 5476: 5472: 5467: 5463: 5458: 5454: 5449: 5445: 5440: 5436: 5431: 5427: 5422: 5418: 5413: 5409: 5404: 5400: 5395: 5388: 5383: 5379: 5374: 5367: 5362: 5358: 5353: 5349: 5344: 5335: 5330: 5326: 5321: 5317: 5312: 5308: 5303: 5299: 5294: 5290: 5285: 5281: 5276: 5272: 5267: 5263: 5258: 5254: 5249: 5245: 5240: 5236: 5231: 5227: 5222: 5218: 5213: 5209: 5204: 5200: 5195: 5191: 5186: 5182: 5177: 5170: 5165: 5161: 5156: 5152: 5147: 5143: 5138: 5134: 5129: 5125: 5120: 5116: 5111: 5107: 5102: 5098: 5093: 5089: 5084: 5080: 5075: 5071: 5066: 5062: 5057: 5053: 5048: 5044: 5039: 5035: 5030: 5026: 5021: 5014: 5009: 5005: 5000: 4985: 4980: 4976: 4971: 4962: 4957: 4953: 4948: 4941: 4936: 4932: 4927: 4923: 4918: 4914: 4909: 4902: 4897: 4893: 4888: 4884: 4879: 4875: 4870: 4866: 4861: 4857: 4852: 4848: 4843: 4839: 4834: 4827: 4822: 4818: 4813: 4800: 4795: 4791: 4786: 4782: 4777: 4770: 4765: 4758: 4753: 4749: 4744: 4740: 4735: 4731: 4726: 4722: 4717: 4710: 4705: 4701: 4696: 4692: 4687: 4683: 4678: 4674: 4669: 4665: 4660: 4653: 4648: 4644: 4639: 4635: 4630: 4626: 4621: 4614: 4609: 4605: 4600: 4596: 4591: 4587: 4582: 4578: 4573: 4566: 4561: 4557: 4552: 4548: 4543: 4539: 4534: 4530: 4525: 4521: 4516: 4512: 4507: 4503: 4498: 4494: 4489: 4485: 4480: 4476: 4471: 4467: 4462: 4458: 4453: 4449: 4444: 4440: 4435: 4431: 4426: 4422: 4417: 4413: 4408: 4404: 4399: 4395: 4390: 4386: 4381: 4377: 4372: 4368: 4363: 4356: 4351: 4347: 4342: 4338: 4333: 4329: 4324: 4311: 4306: 4302: 4297: 4290: 4285: 4278: 4273: 4269: 4264: 4255: 4250: 4243: 4238: 4231: 4226: 4219: 4214: 4210: 4205: 4201: 4196: 4189: 4184: 4177: 4172: 4165: 4160: 4156: 4151: 4147: 4142: 4138: 4133: 4126: 4121: 4117: 4112: 4108: 4103: 4096: 4091: 4087: 4082: 4078: 4073: 4064: 4059: 4055: 4050: 4046: 4041: 4032: 4027: 4023: 4018: 4011: 4006: 4002: 3997: 3993: 3988: 3981: 3976: 3969: 3964: 3960: 3955: 3951: 3946: 3942: 3937: 3933: 3928: 3924: 3919: 3915: 3910: 3906: 3901: 3897: 3892: 3888: 3880: 3876: 3871: 3867: 3862: 3858: 3853: 3849: 3844: 3840: 3835: 3828: 3823: 3819: 3814: 3803: 3798: 3787: 3782: 3778: 3773: 3769: 3764: 3745: 3740: 3736: 3731: 3722: 3718: 3713: 3712: 3707: 3703: 3698: 3694: 3689: 3685: 3680: 3676: 3659: 3655: 3644: 3640: 3635: 3618:protected areas 3610:Oryzomys couesi 3602: 3566:Catacamas virus 3557:Eimeria couesii 3512:in Oaxaca, the 3510:Oryzomys couesi 3461:Oryzomys couesi 3438:Boa constrictor 3401:Oryzomys couesi 3371:Oryzomys couesi 3334:Oryzomys couesi 3330: 3286: 3192: 3191: 3184: 3149:tympanic cavity 3106:squamosal bones 3090:zygomatic plate 3086:maxillary bones 3070: 3069: 3068: 3067: 3051: 3050: 3049: 3041: 3040: 3029: 2997:Oryzomys couesi 2981:Oryzomys couesi 2974:Oryzomys couesi 2970:sex chromosomes 2882:Oryzomys couesi 2852:Gatún, Panama ( 2804:30.9 (29–33.5) 2798:275.8 (255–295) 2780:135.6 (127–150) 2777:264.9 (242–292) 2756:267.5 (255–280) 2741:34.3 (33–35.5) 2678:33.1 (32–34.5) 2480:125.1 (105–150) 2477:244.8 (210–288) 2459:132.3 (102–160) 2456:245.7 (195–301) 2438:137.4 (127–165) 2435:251.4 (239–273) 2374:Oryzomys couesi 2369: 2356:Oryzomys couesi 2348: 2316:Oryzomys couesi 2240:from Paracoté, 2195: 2194: 2193: 2192: 2176: 2175: 2174: 2166: 2165: 2086: 2009:Oryzomys couesi 1958:The subspecies 1956: 1955: 1954: 1953: 1937: 1936: 1935: 1927: 1926: 1846:, the state of 1844:San Luis Potosi 1769:Oryzomys couesi 1765: 1764: 1763: 1762: 1746: 1745: 1744: 1736: 1735: 1719:, and a third, 1697:Oryzomys couesi 1644:J.A. Allen 1614:J.A. Allen 1570:J.A. Allen 1545: 1533:O. fulgens 1523:of the species 1414: 1413: 1412: 1411: 1395: 1394: 1393: 1385: 1384: 1247: 1245:Interior Mexico 1120:O. nelsoni 1035:) and eastern ( 1005:Oryzomys couesi 989:Oryzomys couesi 987:Populations of 954:J.A. Allen 935:J.A. Allen 918: 906:Oryzomys couesi 743:O. nelsoni 694:O. nelsoni 627:Oryzomys couesi 600:Oryzomys couesi 576:Oryzomys couesi 542:Oryzomys couesi 532:Oldfield Thomas 492:Oryzomys couesi 485: 434:Oryzomys couesi 431: 423:Oryzomys couesi 363:Oryzomys couesi 339:Oryzomys couesi 299:Oryzomys couesi 290:and see below. 266:Oryzomys couesi 237:Oryzomys couesi 221: 214: 210:Oryzomys couesi 208: 195: 93: 85: 74: 70: 63: 45: 38: 36:Oryzomys couesi 28: 23: 22: 15: 12: 11: 5: 8617: 8615: 8607: 8606: 8601: 8596: 8591: 8586: 8581: 8576: 8571: 8561: 8560: 8552: 8551: 8549: 8548: 8535: 8522: 8509: 8496: 8483: 8470: 8457: 8444: 8431: 8418: 8405: 8392: 8379: 8366: 8351: 8335: 8333: 8327: 8326: 8321: 8312: 8311: 8309: 8308: 8297: 8295: 8293:Incertae sedis 8289: 8288: 8286: 8285: 8278: 8270: 8268: 8260: 8259: 8257: 8256: 8249: 8241: 8239: 8236:Transandinomys 8231: 8230: 8228: 8227: 8219: 8217: 8209: 8208: 8206: 8205: 8198: 8190: 8188: 8180: 8179: 8177: 8176: 8173:S. ucayalensis 8169: 8161: 8159: 8151: 8150: 8148: 8147: 8138: 8136: 8127: 8126: 8124: 8123: 8115: 8113: 8105: 8104: 8102: 8101: 8092: 8090: 8081: 8080: 8078: 8077: 8070: 8063: 8055: 8048: 8041: 8034: 8026: 8018: 8016: 8008: 8007: 8005: 8004: 7996: 7994: 7986: 7985: 7983: 7982: 7974: 7967: 7960: 7953: 7946: 7939: 7932: 7925: 7918: 7911: 7904: 7897: 7890: 7883: 7876: 7869: 7862: 7854: 7852: 7844: 7843: 7841: 7840: 7833: 7826: 7819: 7812: 7805: 7798: 7791: 7784: 7777: 7770: 7763: 7756: 7749: 7742: 7735: 7727: 7725: 7717: 7716: 7714: 7713: 7704: 7702: 7693: 7692: 7690: 7689: 7682: 7679:N. narboroughi 7675: 7672:N. fernandinae 7668: 7660: 7651: 7649: 7641: 7640: 7638: 7637: 7630: 7623: 7616: 7609: 7602: 7595: 7588: 7581: 7574: 7567: 7560: 7553: 7550:N. albigularis 7545: 7543: 7535: 7534: 7532: 7531: 7524: 7517: 7510: 7503: 7495: 7493: 7485: 7484: 7482: 7481: 7474: 7467: 7460: 7453: 7446: 7439: 7432: 7424: 7422: 7414: 7413: 7411: 7410: 7402: 7400: 7392: 7391: 7389: 7388: 7381: 7373: 7371: 7363: 7362: 7360: 7359: 7351: 7349: 7341: 7340: 7338: 7337: 7330: 7323: 7320:M. caliginosus 7315: 7313: 7305: 7304: 7302: 7301: 7293: 7285: 7282:M. desmarestii 7277: 7269: 7260: 7258: 7249: 7248: 7246: 7245: 7237: 7235: 7227: 7226: 7224: 7223: 7216: 7209: 7202: 7195: 7188: 7181: 7173: 7171: 7163: 7162: 7160: 7159: 7152: 7145: 7137: 7135: 7127: 7126: 7124: 7123: 7116: 7109: 7102: 7095: 7088: 7081: 7074: 7066: 7064: 7056: 7055: 7053: 7052: 7045: 7038: 7035:E. macconnelli 7031: 7024: 7017: 7009: 7007: 6999: 6998: 6996: 6995: 6987: 6985: 6977: 6976: 6974: 6973: 6965: 6963: 6955: 6954: 6952: 6951: 6944: 6937: 6930: 6923: 6916: 6909: 6901: 6899: 6891: 6890: 6888: 6887: 6878: 6876: 6867: 6866: 6864: 6863: 6855: 6853: 6845: 6844: 6842: 6841: 6833: 6824: 6822: 6813: 6812: 6810: 6809: 6806:A. xanthaeolus 6802: 6794: 6792: 6784: 6783: 6781: 6780: 6774: 6768: 6762: 6756: 6750: 6743: 6740: 6739: 6730: 6728: 6727: 6720: 6713: 6705: 6699: 6698: 6697:at Wikispecies 6684: 6668: 6667:External links 6665: 6664: 6663: 6648: 6630: 6619: 6608: 6590: 6576: 6558: 6557:(6)11:402–405. 6544: 6526: 6516: 6498: 6484: 6470: 6459: 6452: 6436: 6425: 6410: 6395: 6380: 6369: 6355: 6324: 6310: 6296: 6282: 6275: 6268: 6257: 6240: 6229: 6228:91(2):336–347. 6214: 6203: 6196: 6192:5(3):171–173. 6176: 6158: 6147: 6132: 6121: 6107: 6093: 6075: 6061: 6047: 6033: 6022: 6011: 6010:41(1):152–157. 6000: 5986: 5972: 5971:82(2):350–353. 5957: 5956:15(4):519–525. 5946: 5932: 5921: 5915:Virus Research 5907: 5900: 5889:Review of the 5885: 5871: 5860: 5849: 5842: 5824: 5799: 5792: 5782: 5768: 5757: 5746: 5735: 5722: 5719: 5717: 5716: 5707: 5698: 5689: 5680: 5671: 5662: 5653: 5644: 5635: 5626: 5617: 5608: 5599: 5590: 5581: 5569: 5557: 5548: 5539: 5530: 5521: 5512: 5500: 5491: 5479: 5470: 5461: 5452: 5443: 5434: 5425: 5416: 5407: 5398: 5386: 5377: 5365: 5356: 5347: 5333: 5324: 5315: 5306: 5297: 5288: 5279: 5270: 5261: 5252: 5243: 5234: 5225: 5216: 5207: 5198: 5189: 5180: 5168: 5159: 5150: 5141: 5132: 5123: 5114: 5105: 5096: 5087: 5078: 5069: 5060: 5051: 5042: 5033: 5024: 5012: 5003: 4983: 4974: 4960: 4951: 4939: 4930: 4921: 4912: 4900: 4891: 4882: 4873: 4864: 4855: 4846: 4837: 4825: 4816: 4798: 4789: 4780: 4768: 4756: 4747: 4738: 4729: 4720: 4708: 4699: 4690: 4681: 4672: 4663: 4651: 4642: 4633: 4624: 4612: 4603: 4594: 4585: 4576: 4564: 4555: 4546: 4537: 4528: 4519: 4510: 4501: 4492: 4483: 4474: 4465: 4456: 4447: 4438: 4429: 4420: 4411: 4402: 4393: 4384: 4375: 4366: 4354: 4345: 4336: 4327: 4309: 4300: 4288: 4276: 4267: 4253: 4241: 4229: 4217: 4208: 4199: 4187: 4175: 4163: 4154: 4145: 4136: 4124: 4115: 4106: 4094: 4085: 4076: 4062: 4053: 4044: 4030: 4021: 4009: 4000: 3991: 3979: 3967: 3958: 3949: 3940: 3931: 3922: 3913: 3904: 3895: 3886: 3874: 3865: 3856: 3847: 3838: 3826: 3817: 3801: 3785: 3776: 3767: 3743: 3734: 3719: 3717: 3714: 3711: 3710: 3701: 3692: 3683: 3674: 3653: 3637: 3636: 3634: 3631: 3626:climate change 3601: 3598: 3582:O. couesi 3578:O. couesi 3465:O. couesi 3451:O. couesi 3443:O. couesi 3417:O. couesi 3393:O. couesi 3379:O. couesi 3329: 3326: 3285: 3282: 3232:O. couesi 3188:dental formula 3183: 3180: 3164:mental foramen 3145:tegmen tympani 3141:squamosal bone 3102:parietal bones 3072:The nasal and 3053: 3052: 3043: 3042: 3034: 3033: 3032: 3031: 3030: 3028: 3025: 2966:subtelocentric 2954:submetacentric 2877: 2876: 2870: 2869: 2866: 2863: 2860: 2857: 2849: 2848: 2845: 2842: 2839: 2836: 2828: 2827: 2823: 2822: 2819: 2816: 2813: 2810: 2806: 2805: 2802: 2799: 2796: 2793: 2785: 2784: 2781: 2778: 2775: 2772: 2764: 2763: 2760: 2757: 2754: 2751: 2743: 2742: 2739: 2736: 2733: 2730: 2722: 2721: 2718: 2715: 2712: 2709: 2701: 2700: 2697: 2694: 2691: 2688: 2680: 2679: 2676: 2673: 2670: 2667: 2659: 2658: 2655: 2652: 2649: 2646: 2638: 2637: 2634: 2631: 2628: 2625: 2617: 2616: 2612: 2611: 2608: 2605: 2602: 2599: 2591: 2590: 2587: 2584: 2581: 2578: 2570: 2569: 2566: 2563: 2560: 2557: 2549: 2548: 2545: 2542: 2539: 2536: 2528: 2527: 2523: 2522: 2519: 2516: 2513: 2510: 2506: 2505: 2502: 2499: 2496: 2493: 2485: 2484: 2481: 2478: 2475: 2472: 2464: 2463: 2460: 2457: 2454: 2451: 2443: 2442: 2439: 2436: 2433: 2430: 2422: 2421: 2418: 2415: 2412: 2409: 2401: 2400: 2396: 2395: 2392: 2389: 2386: 2381: 2368: 2365: 2361:O. couesi 2347: 2344: 2336:O. couesi 2302:O. couesi 2290:O. couesi 2286:O. couesi 2274:O. couesi 2191:Goldman, 1912) 2178: 2177: 2168: 2167: 2159: 2158: 2157: 2156: 2155: 2143:O. couesi 2139:O. couesi 2131:O. couesi 2118: 2117: 2116: 2115: 2104: 2085: 2082: 2078:O. couesi 2062:Lake Nicaragua 2054:O. couesi 2032:O. couesi 2019:off Honduras. 2013:Turneffe Atoll 1952:Merriam, 1901) 1939: 1938: 1929: 1928: 1920: 1919: 1918: 1917: 1916: 1785:O. couesi 1748: 1747: 1738: 1737: 1729: 1728: 1727: 1726: 1725: 1684: 1683: 1682: 1681: 1670: 1659: 1648: 1637: 1629: 1618: 1607: 1596: 1585: 1574: 1563: 1544: 1541: 1410:Goldman, 1915) 1397: 1396: 1387: 1386: 1378: 1377: 1376: 1375: 1374: 1332:from Jalisco, 1304: 1303: 1302: 1301: 1290: 1279: 1268: 1246: 1243: 1180:from northern 1178:O. couesi 1142:Oryzomys rufus 1083:Hodomys alleni 983: 982: 981: 980: 969: 961:Oryzomys rufus 958: 947: 939: 917: 914: 902:O. couesi 894:O. couesi 890:O. couesi 886:O. couesi 882:O. couesi 874:O. couesi 853:O. couesi 829:O. couesi 821:O. couesi 771:O. couesi 763:O. couesi 725:Marías Islands 711:O. couesi 707:O. couesi 608:O. couesi 596:O. couesi 592:O. couesi 506:marsh rice rat 484: 481: 475:of the family 448:marsh rice rat 438:O. couesi 430: 427: 411:O. couesi 403:O. couesi 383:O. couesi 375:marsh rice rat 323:O. couesi 313:in the family 293: 292: 288: 287: 275: 263: 252: 251: 245: 244: 232: 231: 223: 222: 215: 204: 203: 197: 196: 192:O. couesi 189: 187: 183: 182: 175: 171: 170: 165: 161: 160: 155: 151: 150: 145: 141: 140: 135: 131: 130: 125: 121: 120: 115: 111: 110: 105: 101: 100: 87: 86: 68: 65: 64: 59: 56: 55: 47: 46: 39: 26: 24: 14: 13: 10: 9: 6: 4: 3: 2: 8616: 8605: 8602: 8600: 8597: 8595: 8592: 8590: 8587: 8585: 8582: 8580: 8577: 8575: 8572: 8570: 8567: 8566: 8564: 8557: 8545: 8540: 8536: 8532: 8527: 8523: 8519: 8514: 8510: 8506: 8501: 8497: 8493: 8488: 8484: 8480: 8475: 8471: 8467: 8462: 8458: 8454: 8449: 8445: 8441: 8436: 8432: 8428: 8423: 8419: 8415: 8410: 8406: 8402: 8397: 8393: 8389: 8384: 8380: 8376: 8371: 8367: 8362: 8356: 8352: 8347: 8341: 8337: 8336: 8334: 8332: 8328: 8324: 8319: 8307: 8305: 8299: 8298: 8296: 8294: 8290: 8284: 8283: 8279: 8277: 8276: 8275:Z. brevicauda 8272: 8271: 8269: 8267: 8266: 8261: 8255: 8254: 8253:T. talamancae 8250: 8248: 8247: 8243: 8242: 8240: 8238: 8237: 8232: 8226: 8225: 8221: 8220: 8218: 8216: 8215: 8210: 8204: 8203: 8199: 8197: 8196: 8192: 8191: 8189: 8187: 8186: 8185:Sigmodontomys 8181: 8175: 8174: 8170: 8168: 8167: 8163: 8162: 8160: 8158: 8157: 8152: 8146: 8145: 8144:R. primigenus 8140: 8139: 8137: 8135: 8134: 8128: 8122: 8121: 8117: 8116: 8114: 8112: 8111: 8110:Pseudoryzomys 8106: 8100: 8099: 8094: 8093: 8091: 8089: 8088: 8082: 8076: 8075: 8074:O. peninsulae 8071: 8069: 8068: 8067:O. palustris 8064: 8062: 8061: 8056: 8054: 8053: 8049: 8047: 8046: 8045:O. dimidiatus 8042: 8040: 8039: 8035: 8033: 8032: 8031:O. antillarum 8027: 8025: 8024: 8023:O. albiventer 8020: 8019: 8017: 8015: 8014: 8009: 8003: 8002: 7998: 7997: 7995: 7993: 7992: 7987: 7981: 7980: 7975: 7973: 7972: 7968: 7966: 7965: 7964:O. stramineus 7961: 7959: 7958: 7954: 7952: 7951: 7947: 7945: 7944: 7940: 7938: 7937: 7933: 7931: 7930: 7926: 7924: 7923: 7919: 7917: 7916: 7912: 7910: 7909: 7908:O. fulvescens 7905: 7903: 7902: 7898: 7896: 7895: 7894:O. flavescens 7891: 7889: 7888: 7887:O. destructor 7884: 7882: 7881: 7880:O. chacoensis 7877: 7875: 7874: 7870: 7868: 7867: 7863: 7861: 7860: 7856: 7855: 7853: 7851: 7850: 7845: 7839: 7838: 7837:O. trinitatis 7834: 7832: 7831: 7827: 7825: 7824: 7820: 7818: 7817: 7813: 7811: 7810: 7806: 7804: 7803: 7799: 7797: 7796: 7792: 7790: 7789: 7785: 7783: 7782: 7778: 7776: 7775: 7771: 7769: 7768: 7764: 7762: 7761: 7757: 7755: 7754: 7750: 7748: 7747: 7746:O. catherinae 7743: 7741: 7740: 7736: 7734: 7733: 7732:O. auyantepui 7729: 7728: 7726: 7724: 7723: 7718: 7712: 7711: 7706: 7705: 7703: 7701: 7700: 7694: 7688: 7687: 7683: 7681: 7680: 7676: 7674: 7673: 7669: 7667: 7666: 7665:N. indefessus 7661: 7659: 7658: 7653: 7652: 7650: 7648: 7647: 7642: 7636: 7635: 7631: 7629: 7628: 7627:N. pectoralis 7624: 7622: 7621: 7617: 7615: 7614: 7610: 7608: 7607: 7606:N. meridensis 7603: 7601: 7600: 7596: 7594: 7593: 7589: 7587: 7586: 7582: 7580: 7579: 7575: 7573: 7572: 7568: 7566: 7565: 7561: 7559: 7558: 7557:N. auriventer 7554: 7552: 7551: 7547: 7546: 7544: 7542: 7541: 7536: 7530: 7529: 7525: 7523: 7522: 7518: 7516: 7515: 7511: 7509: 7508: 7504: 7502: 7501: 7497: 7496: 7494: 7492: 7491: 7486: 7480: 7479: 7475: 7473: 7472: 7468: 7466: 7465: 7461: 7459: 7458: 7454: 7452: 7451: 7447: 7445: 7444: 7440: 7438: 7437: 7433: 7431: 7430: 7426: 7425: 7423: 7421: 7420: 7415: 7409: 7408: 7404: 7403: 7401: 7399: 7398: 7393: 7387: 7386: 7382: 7380: 7379: 7378:M. altissimus 7375: 7374: 7372: 7370: 7369: 7364: 7358: 7357: 7353: 7352: 7350: 7348: 7347: 7342: 7336: 7335: 7331: 7329: 7328: 7327:M. robustulus 7324: 7322: 7321: 7317: 7316: 7314: 7312: 7311: 7306: 7300: 7299: 7294: 7292: 7291: 7286: 7284: 7283: 7278: 7276: 7275: 7274:M. curazensis 7270: 7268: 7267: 7262: 7261: 7259: 7257: 7256: 7250: 7244: 7243: 7239: 7238: 7236: 7234: 7233: 7228: 7222: 7221: 7217: 7215: 7214: 7210: 7208: 7207: 7206:H. perenensis 7203: 7201: 7200: 7196: 7194: 7193: 7189: 7187: 7186: 7182: 7180: 7179: 7175: 7174: 7172: 7170: 7169: 7164: 7158: 7157: 7153: 7151: 7150: 7146: 7144: 7143: 7139: 7138: 7136: 7134: 7133: 7128: 7122: 7121: 7120:H. saturatior 7117: 7115: 7114: 7110: 7108: 7107: 7103: 7101: 7100: 7096: 7094: 7093: 7089: 7087: 7086: 7082: 7080: 7079: 7075: 7073: 7072: 7068: 7067: 7065: 7063: 7062: 7057: 7051: 7050: 7046: 7044: 7043: 7039: 7037: 7036: 7032: 7030: 7029: 7025: 7023: 7022: 7018: 7016: 7015: 7011: 7010: 7008: 7006: 7005: 7000: 6994: 6993: 6989: 6988: 6986: 6984: 6983: 6978: 6972: 6971: 6967: 6966: 6964: 6962: 6961: 6956: 6950: 6949: 6945: 6943: 6942: 6938: 6936: 6935: 6931: 6929: 6928: 6924: 6922: 6921: 6917: 6915: 6914: 6910: 6908: 6907: 6903: 6902: 6900: 6898: 6897: 6892: 6886: 6885: 6880: 6879: 6877: 6875: 6874: 6868: 6862: 6861: 6857: 6856: 6854: 6852: 6851: 6850:Amphinectomys 6846: 6840: 6839: 6834: 6832: 6831: 6826: 6825: 6823: 6821: 6820: 6814: 6808: 6807: 6803: 6801: 6800: 6796: 6795: 6793: 6791: 6790: 6785: 6779: 6778:Sigmodontinae 6775: 6773: 6769: 6767: 6763: 6761: 6757: 6755: 6751: 6749: 6745: 6744: 6741: 6737: 6726: 6721: 6719: 6714: 6712: 6707: 6706: 6703: 6696: 6695: 6689: 6685: 6682: 6681: 6675: 6671: 6670: 6666: 6661: 6657: 6655: 6649: 6643: 6639: 6637: 6631: 6628: 6624: 6620: 6617: 6613: 6609: 6604:137:210–222. 6603: 6599: 6597: 6591: 6586:81(1):77–85. 6585: 6581: 6577: 6571: 6567: 6565: 6559: 6556: 6552: 6551: 6545: 6539: 6535: 6533: 6527: 6525: 6521: 6517: 6511: 6507: 6505: 6499: 6497: 6493: 6489: 6485: 6483: 6479: 6475: 6471: 6468: 6464: 6460: 6457: 6453: 6451: 6447: 6443: 6442: 6437: 6434: 6430: 6426: 6423: 6419: 6417: 6411: 6408: 6404: 6402: 6396: 6393: 6389: 6387: 6381: 6378: 6374: 6370: 6368: 6364: 6360: 6356: 6344: 6340: 6339: 6334: 6332: 6325: 6319: 6315: 6311: 6309: 6308:0-691-07012-1 6305: 6301: 6297: 6291: 6287: 6283: 6280: 6276: 6273: 6269: 6266: 6262: 6258: 6252: 6248: 6247: 6241: 6238: 6234: 6230: 6227: 6223: 6221: 6215: 6212: 6208: 6204: 6201: 6197: 6191: 6187: 6186: 6182: 6177: 6171: 6167: 6165: 6159: 6156: 6152: 6148: 6145: 6141: 6139: 6133: 6130: 6126: 6122: 6119: 6115: 6114: 6108: 6102: 6098: 6094: 6091: 6087: 6083: 6080: 6076: 6071:31(1):23–37. 6070: 6066: 6062: 6056: 6052: 6048: 6042: 6038: 6034: 6031: 6027: 6023: 6020: 6016: 6012: 6009: 6005: 6001: 5999: 5998:0-7136-6021-X 5995: 5991: 5987: 5981: 5977: 5973: 5970: 5966: 5964: 5958: 5955: 5951: 5947: 5941: 5937: 5933: 5930: 5926: 5922: 5917:131:180–188. 5916: 5912: 5908: 5905: 5901: 5898: 5894: 5892: 5886: 5880: 5876: 5872: 5869: 5865: 5861: 5858: 5854: 5850: 5847: 5843: 5837: 5833: 5831: 5825: 5819: 5815: 5813: 5809: 5805: 5800: 5797: 5793: 5790: 5789: 5783: 5780: 5776: 5775: 5769: 5766: 5762: 5758: 5755: 5751: 5747: 5744: 5740: 5736: 5733: 5729: 5725: 5724: 5720: 5711: 5708: 5702: 5699: 5693: 5690: 5684: 5681: 5675: 5672: 5666: 5663: 5657: 5654: 5648: 5645: 5639: 5636: 5630: 5627: 5621: 5618: 5612: 5609: 5603: 5600: 5594: 5591: 5585: 5582: 5576: 5574: 5570: 5564: 5562: 5558: 5552: 5549: 5543: 5540: 5534: 5531: 5525: 5522: 5516: 5513: 5507: 5505: 5501: 5495: 5492: 5486: 5484: 5480: 5474: 5471: 5465: 5462: 5456: 5453: 5447: 5444: 5438: 5435: 5429: 5426: 5420: 5417: 5411: 5408: 5402: 5399: 5393: 5391: 5387: 5381: 5378: 5372: 5370: 5366: 5360: 5357: 5351: 5348: 5342: 5340: 5338: 5334: 5328: 5325: 5319: 5316: 5310: 5307: 5301: 5298: 5292: 5289: 5283: 5280: 5274: 5271: 5265: 5262: 5256: 5253: 5247: 5244: 5238: 5235: 5229: 5226: 5220: 5217: 5211: 5208: 5202: 5199: 5193: 5190: 5184: 5181: 5175: 5173: 5169: 5163: 5160: 5154: 5151: 5145: 5142: 5136: 5133: 5127: 5124: 5118: 5115: 5109: 5106: 5100: 5097: 5091: 5088: 5082: 5079: 5073: 5070: 5064: 5061: 5055: 5052: 5046: 5043: 5037: 5034: 5028: 5025: 5019: 5017: 5013: 5007: 5004: 4998: 4996: 4994: 4992: 4990: 4988: 4984: 4978: 4975: 4969: 4967: 4965: 4961: 4955: 4952: 4946: 4944: 4940: 4934: 4931: 4925: 4922: 4916: 4913: 4907: 4905: 4901: 4895: 4892: 4886: 4883: 4877: 4874: 4868: 4865: 4859: 4856: 4850: 4847: 4841: 4838: 4832: 4830: 4826: 4820: 4817: 4811: 4809: 4807: 4805: 4803: 4799: 4793: 4790: 4784: 4781: 4775: 4773: 4769: 4763: 4761: 4757: 4751: 4748: 4742: 4739: 4733: 4730: 4724: 4721: 4715: 4713: 4709: 4703: 4700: 4694: 4691: 4685: 4682: 4676: 4673: 4667: 4664: 4658: 4656: 4652: 4646: 4643: 4637: 4634: 4628: 4625: 4619: 4617: 4613: 4607: 4604: 4598: 4595: 4589: 4586: 4580: 4577: 4571: 4569: 4565: 4559: 4556: 4550: 4547: 4541: 4538: 4532: 4529: 4523: 4520: 4514: 4511: 4505: 4502: 4496: 4493: 4487: 4484: 4478: 4475: 4469: 4466: 4460: 4457: 4451: 4448: 4442: 4439: 4433: 4430: 4424: 4421: 4415: 4412: 4406: 4403: 4397: 4394: 4388: 4385: 4379: 4376: 4370: 4367: 4361: 4359: 4355: 4349: 4346: 4340: 4337: 4331: 4328: 4322: 4320: 4318: 4316: 4314: 4310: 4304: 4301: 4295: 4293: 4289: 4283: 4281: 4277: 4271: 4268: 4262: 4260: 4258: 4254: 4248: 4246: 4242: 4236: 4234: 4230: 4224: 4222: 4218: 4212: 4209: 4203: 4200: 4194: 4192: 4188: 4182: 4180: 4176: 4170: 4168: 4164: 4158: 4155: 4149: 4146: 4140: 4137: 4131: 4129: 4125: 4119: 4116: 4110: 4107: 4101: 4099: 4095: 4089: 4086: 4080: 4077: 4071: 4069: 4067: 4063: 4057: 4054: 4048: 4045: 4039: 4037: 4035: 4031: 4025: 4022: 4016: 4014: 4010: 4004: 4001: 3995: 3992: 3986: 3984: 3980: 3974: 3972: 3968: 3962: 3959: 3953: 3950: 3944: 3941: 3935: 3932: 3926: 3923: 3917: 3914: 3908: 3905: 3899: 3896: 3890: 3887: 3884: 3878: 3875: 3869: 3866: 3860: 3857: 3851: 3848: 3842: 3839: 3833: 3831: 3827: 3821: 3818: 3812: 3810: 3808: 3806: 3802: 3796: 3794: 3792: 3790: 3786: 3780: 3777: 3771: 3768: 3762: 3760: 3758: 3756: 3754: 3752: 3750: 3748: 3744: 3738: 3735: 3729: 3727: 3725: 3721: 3715: 3705: 3702: 3696: 3693: 3687: 3684: 3678: 3675: 3671: 3667: 3663: 3657: 3654: 3650: 3649: 3642: 3639: 3632: 3630: 3627: 3623: 3619: 3615: 3614:Least Concern 3611: 3607: 3599: 3597: 3595: 3591: 3587: 3583: 3579: 3575: 3571: 3567: 3563: 3559: 3558: 3554: 3550: 3549: 3544: 3543: 3538: 3537: 3532: 3531: 3526: 3525: 3520: 3519: 3515: 3514:sucking louse 3511: 3507: 3506: 3501: 3500: 3496: 3492: 3491: 3487:of the genus 3486: 3482: 3481: 3476: 3475: 3470: 3466: 3462: 3458: 3457: 3452: 3448: 3444: 3440: 3439: 3434: 3429: 3427: 3423: 3418: 3414: 3409: 3405: 3402: 3398: 3394: 3390: 3389: 3384: 3380: 3376: 3372: 3368: 3365: 3364:coastal plain 3361: 3357: 3356: 3351: 3347: 3343: 3339: 3335: 3327: 3325: 3323: 3319: 3315: 3311: 3307: 3303: 3299: 3295: 3291: 3283: 3281: 3279: 3275: 3271: 3267: 3263: 3259: 3254: 3252: 3248: 3244: 3239: 3237: 3233: 3229: 3225: 3221: 3217: 3213: 3209: 3205: 3189: 3181: 3179: 3177: 3173: 3169: 3165: 3161: 3156: 3154: 3150: 3146: 3142: 3138: 3134: 3130: 3126: 3122: 3118: 3114: 3109: 3107: 3103: 3099: 3095: 3091: 3087: 3083: 3079: 3075: 3066:Alston, 1877) 3065: 3061: 3057: 3047: 3038: 3026: 3024: 3022: 3021: 3016: 3012: 3008: 3004: 3003: 2998: 2994: 2990: 2989:cartilaginous 2986: 2982: 2977: 2975: 2971: 2967: 2963: 2959: 2955: 2951: 2947: 2943: 2939: 2935: 2931: 2928:is absent, a 2927: 2923: 2919: 2915: 2911: 2906: 2904: 2900: 2896: 2892: 2888: 2883: 2871: 2867: 2864: 2861: 2858: 2855: 2851: 2850: 2846: 2843: 2840: 2837: 2834: 2830: 2829: 2824: 2820: 2817: 2814: 2811: 2808: 2807: 2803: 2801:137 (124–151) 2800: 2797: 2794: 2791: 2787: 2786: 2783:28.8 (27–31) 2782: 2779: 2776: 2773: 2770: 2766: 2765: 2762:29.1 (28–32) 2761: 2759:138 (130–145) 2758: 2755: 2752: 2749: 2745: 2744: 2740: 2738:172 (163–177) 2737: 2735:306 (285–327) 2734: 2731: 2728: 2724: 2723: 2720:27 (24.6–29) 2719: 2717:122 (107–146) 2716: 2713: 2710: 2707: 2703: 2702: 2699:30.7 (30–31) 2698: 2696:130 (127–135) 2695: 2693:252 (242–265) 2692: 2689: 2686: 2682: 2681: 2677: 2675:148 (139–174) 2674: 2672:263 (248–294) 2671: 2668: 2665: 2661: 2660: 2656: 2654:157 (143–167) 2653: 2651:281 (265–294) 2650: 2647: 2644: 2640: 2639: 2636:34.5 (32–38) 2635: 2633:161 (138–180) 2632: 2630:297 (283–310) 2629: 2626: 2623: 2619: 2618: 2613: 2609: 2606: 2603: 2600: 2597: 2593: 2592: 2588: 2586:168 (155–180) 2585: 2583:308 (285–320) 2582: 2579: 2576: 2572: 2571: 2567: 2564: 2561: 2558: 2555: 2552:Mexico City ( 2551: 2550: 2546: 2544:161 (154–170) 2543: 2541:307 (297–318) 2540: 2537: 2534: 2530: 2529: 2524: 2520: 2517: 2514: 2511: 2508: 2507: 2503: 2500: 2497: 2494: 2491: 2487: 2486: 2483:30.5 (27–33) 2482: 2479: 2476: 2473: 2470: 2466: 2465: 2462:30.2 (26–34) 2461: 2458: 2455: 2452: 2449: 2445: 2444: 2441:28.9 (27–35) 2440: 2437: 2434: 2431: 2428: 2424: 2423: 2419: 2416: 2413: 2410: 2407: 2403: 2402: 2397: 2393: 2390: 2387: 2385: 2382: 2379: 2378: 2375: 2366: 2364: 2362: 2357: 2353: 2345: 2343: 2341: 2337: 2333: 2329: 2325: 2321: 2317: 2313: 2311: 2307: 2303: 2299: 2295: 2291: 2287: 2283: 2279: 2275: 2271: 2267: 2263: 2259: 2255: 2251: 2247: 2243: 2239: 2235: 2231: 2227: 2223: 2219: 2215: 2211: 2207: 2203: 2199: 2190: 2186: 2182: 2172: 2163: 2154: 2152: 2148: 2144: 2140: 2136: 2132: 2128: 2124: 2112: 2108: 2105: 2100: 2097: 2096: 2095: 2094: 2091: 2083: 2081: 2079: 2075: 2071: 2067: 2063: 2059: 2055: 2051: 2047: 2046: 2041: 2037: 2033: 2029: 2025: 2020: 2018: 2014: 2010: 2006: 2002: 1997: 1993: 1989: 1986:and mainland 1985: 1981: 1977: 1973: 1969: 1965: 1961: 1951: 1947: 1943: 1933: 1924: 1915: 1912: 1908: 1904: 1900: 1896: 1892: 1888: 1885:in Guatemala— 1884: 1883:type locality 1880: 1876: 1871: 1869: 1865: 1861: 1857: 1853: 1849: 1845: 1841: 1837: 1832: 1830: 1826: 1822: 1818: 1814: 1813:hybridization 1810: 1806: 1802: 1798: 1794: 1790: 1786: 1782: 1778: 1774: 1770: 1760: 1756: 1752: 1742: 1733: 1724: 1722: 1718: 1714: 1710: 1706: 1702: 1698: 1694: 1690: 1678: 1674: 1671: 1667: 1663: 1660: 1656: 1652: 1649: 1645: 1641: 1638: 1633: 1630: 1626: 1622: 1619: 1615: 1611: 1608: 1604: 1600: 1597: 1593: 1589: 1586: 1582: 1578: 1575: 1571: 1567: 1564: 1559: 1556: 1555: 1554: 1553: 1550: 1542: 1540: 1538: 1534: 1530: 1526: 1522: 1517: 1515: 1511: 1507: 1503: 1499: 1495: 1491: 1487: 1483: 1479: 1475: 1471: 1470: 1465: 1461: 1457: 1453: 1449: 1445: 1441: 1437: 1433: 1429: 1425: 1424: 1419: 1409: 1406:, Michoacán ( 1405: 1401: 1391: 1382: 1373: 1371: 1367: 1363: 1359: 1355: 1351: 1347: 1343: 1339: 1335: 1331: 1327: 1323: 1319: 1315: 1311: 1298: 1294: 1291: 1287: 1283: 1280: 1276: 1272: 1269: 1265: 1261: 1258: 1257: 1256: 1255: 1252: 1244: 1242: 1240: 1236: 1232: 1228: 1224: 1220: 1216: 1212: 1208: 1204: 1202: 1198: 1194: 1191: 1187: 1183: 1179: 1175: 1171: 1167: 1163: 1159: 1155: 1151: 1147: 1143: 1139: 1135: 1131: 1127: 1123: 1121: 1117: 1113: 1109: 1105: 1101: 1100:biogeographic 1097: 1096: 1091: 1090: 1085: 1084: 1079: 1078: 1073: 1069: 1065: 1061: 1057: 1053: 1048: 1047:in the east. 1046: 1042: 1038: 1034: 1030: 1026: 1022: 1018: 1014: 1010: 1006: 1002: 998: 994: 990: 977: 973: 970: 966: 962: 959: 955: 951: 948: 943: 940: 936: 932: 929: 928: 927: 926: 923: 915: 913: 911: 907: 903: 899: 895: 891: 887: 883: 879: 875: 871: 867: 863: 858: 854: 850: 849: 844: 843: 838: 834: 830: 826: 822: 818: 814: 810: 806: 802: 798: 794: 790: 786: 785: 779: 778:mitochondrial 774: 772: 768: 764: 760: 759: 754: 750: 749: 744: 740: 736: 732: 731: 726: 722: 721: 716: 712: 708: 704: 696: 695: 690: 689: 684: 683: 678: 674: 670: 666: 662: 661: 656: 652: 648: 644: 640: 636: 632: 628: 623: 619: 617: 613: 609: 605: 601: 597: 593: 589: 585: 581: 577: 573: 569: 565: 563: 559: 555: 551: 547: 543: 539: 538: 533: 529: 528:Elliott Coues 525: 521: 520:specific name 517: 516: 511: 508:(then called 507: 503: 502: 497: 493: 489: 488:Edward Alston 482: 480: 478: 474: 473:Sigmodontinae 470: 465: 461: 457: 453: 449: 445: 444: 439: 435: 428: 426: 424: 421:. Generally, 420: 416: 412: 408: 404: 400: 396: 395: 390: 389: 384: 380: 376: 372: 368: 364: 360: 355: 353: 349: 345: 340: 336: 332: 328: 324: 320: 316: 312: 309: 305: 301: 300: 291: 284: 279: 276: 272: 267: 264: 259: 256: 255: 253: 250: 246: 242: 238: 233: 229: 224: 219: 213: 211: 205: 202: 201:Binomial name 198: 194: 193: 188: 185: 184: 181: 180: 176: 173: 172: 169: 168:Sigmodontinae 166: 163: 162: 159: 156: 153: 152: 149: 146: 143: 142: 139: 136: 133: 132: 129: 126: 123: 122: 119: 116: 113: 112: 109: 106: 103: 102: 97: 92: 88: 82: 77: 76:Least Concern 66: 62: 57: 53: 48: 43: 37: 33: 30: 19: 8556: 8330: 8301: 8280: 8273: 8265:Zygodontomys 8263: 8251: 8246:T. bolivaris 8244: 8234: 8222: 8212: 8202:S. aphrastus 8200: 8193: 8183: 8171: 8164: 8154: 8142: 8131: 8118: 8108: 8096: 8085: 8072: 8066: 8058: 8050: 8043: 8037: 8036: 8029: 8021: 8011: 8001:O. balneator 7999: 7989: 7977: 7969: 7962: 7957:O. rupestris 7955: 7948: 7941: 7934: 7927: 7920: 7915:O. griseolus 7913: 7906: 7899: 7892: 7885: 7878: 7871: 7864: 7857: 7849:Oligoryzomys 7847: 7835: 7828: 7821: 7816:O. speciosus 7814: 7807: 7800: 7793: 7786: 7779: 7772: 7767:O. flavicans 7765: 7758: 7751: 7744: 7737: 7730: 7720: 7710:N. vespuccii 7708: 7697: 7684: 7677: 7670: 7663: 7655: 7644: 7634:N. pirrensis 7632: 7625: 7618: 7611: 7604: 7597: 7590: 7583: 7576: 7569: 7564:N. caracolus 7562: 7555: 7548: 7538: 7528:N. squamipes 7526: 7519: 7512: 7505: 7498: 7488: 7476: 7469: 7462: 7455: 7448: 7441: 7434: 7427: 7417: 7405: 7395: 7383: 7376: 7368:Microryzomys 7366: 7354: 7344: 7332: 7325: 7318: 7308: 7296: 7290:M. georginae 7288: 7280: 7272: 7264: 7253: 7240: 7230: 7218: 7211: 7204: 7197: 7190: 7183: 7176: 7166: 7154: 7149:H. chacarius 7147: 7140: 7130: 7118: 7113:H. rostratus 7111: 7104: 7099:H. melanotis 7097: 7090: 7083: 7076: 7069: 7059: 7047: 7040: 7033: 7026: 7019: 7012: 7004:Euryoryzomys 7002: 6990: 6982:Eremoryzomys 6980: 6968: 6958: 6946: 6941:C. subflavus 6939: 6932: 6925: 6918: 6913:C. langguthi 6911: 6904: 6894: 6882: 6873:Carletonomys 6871: 6858: 6848: 6836: 6828: 6819:Agathaeromys 6817: 6804: 6797: 6787: 6693: 6679: 6659: 6653: 6641: 6635: 6626: 6615: 6601: 6595: 6583: 6569: 6563: 6554: 6549: 6537: 6531: 6512:65:205–214. 6509: 6503: 6487: 6473: 6466: 6455: 6440: 6432: 6421: 6415: 6406: 6400: 6391: 6385: 6376: 6358: 6347:. Retrieved 6342: 6336: 6330: 6317: 6299: 6289: 6271: 6264: 6250: 6245: 6236: 6225: 6219: 6210: 6189: 6184: 6180: 6169: 6163: 6154: 6143: 6137: 6131:56(36):1–11. 6128: 6117: 6112: 6103:78:857–865. 6100: 6089: 6068: 6054: 6040: 6029: 6018: 6007: 5989: 5979: 5968: 5962: 5953: 5939: 5928: 5914: 5903: 5896: 5890: 5878: 5867: 5856: 5852: 5845: 5835: 5829: 5817: 5811: 5807: 5803: 5787: 5778: 5773: 5764: 5753: 5742: 5731: 5710: 5701: 5692: 5683: 5674: 5665: 5656: 5647: 5638: 5629: 5620: 5611: 5602: 5593: 5584: 5551: 5542: 5533: 5524: 5515: 5494: 5473: 5464: 5455: 5446: 5437: 5428: 5419: 5410: 5401: 5380: 5359: 5350: 5327: 5318: 5309: 5300: 5291: 5282: 5273: 5264: 5255: 5246: 5237: 5228: 5219: 5210: 5201: 5192: 5183: 5162: 5153: 5144: 5135: 5126: 5117: 5108: 5099: 5090: 5081: 5072: 5063: 5054: 5045: 5036: 5027: 5006: 4977: 4954: 4933: 4924: 4915: 4894: 4885: 4876: 4867: 4858: 4849: 4840: 4819: 4792: 4783: 4750: 4741: 4732: 4723: 4702: 4693: 4684: 4675: 4666: 4645: 4636: 4627: 4606: 4597: 4588: 4579: 4558: 4549: 4540: 4531: 4522: 4513: 4504: 4495: 4486: 4477: 4468: 4459: 4450: 4441: 4432: 4423: 4414: 4405: 4396: 4387: 4378: 4369: 4348: 4339: 4330: 4303: 4270: 4211: 4202: 4157: 4148: 4139: 4118: 4109: 4088: 4079: 4056: 4047: 4024: 4003: 3994: 3961: 3952: 3943: 3934: 3925: 3916: 3907: 3898: 3889: 3882: 3877: 3868: 3859: 3850: 3841: 3820: 3779: 3770: 3737: 3704: 3695: 3686: 3677: 3669: 3665: 3661: 3656: 3646: 3641: 3609: 3603: 3581: 3577: 3562:hantaviruses 3555: 3553:apicomplexan 3546: 3540: 3534: 3528: 3522: 3516: 3509: 3503: 3497: 3488: 3478: 3472: 3464: 3460: 3454: 3450: 3449:recorded on 3445:on Cozumel. 3442: 3436: 3430: 3421: 3416: 3413:cloud forest 3406: 3400: 3392: 3386: 3378: 3370: 3369: 3353: 3333: 3331: 3314:hemal arches 3287: 3266:anterolophid 3255: 3240: 3231: 3185: 3178:, is large. 3157: 3153:mastoid bone 3110: 3074:premaxillary 3071: 3063: 3062:, Honduras ( 3055: 3018: 3002:Oligoryzomys 3000: 2996: 2980: 2978: 2973: 2958:X chromosome 2930:synapomorphy 2926:gall bladder 2907: 2891:ungual tufts 2881: 2880: 2853: 2832: 2789: 2768: 2747: 2726: 2714:108 (96–128) 2705: 2684: 2663: 2642: 2621: 2595: 2574: 2553: 2532: 2489: 2468: 2447: 2426: 2405: 2388:Total length 2383: 2373: 2360: 2355: 2352:common names 2349: 2346:Common names 2335: 2315: 2314: 2309: 2305: 2301: 2297: 2293: 2289: 2285: 2281: 2277: 2273: 2269: 2265: 2261: 2257: 2253: 2249: 2245: 2237: 2233: 2217: 2205: 2201: 2197: 2196: 2188: 2180: 2150: 2146: 2142: 2138: 2134: 2130: 2126: 2122: 2121: 2106: 2098: 2077: 2053: 2049: 2043: 2039: 2035: 2031: 2027: 2023: 2021: 2008: 2004: 2000: 1995: 1991: 1987: 1983: 1979: 1971: 1967: 1959: 1957: 1949: 1941: 1910: 1906: 1902: 1894: 1890: 1886: 1878: 1872: 1867: 1863: 1859: 1835: 1833: 1828: 1824: 1820: 1816: 1804: 1784: 1780: 1776: 1768: 1766: 1761:Allen, 1891) 1758: 1750: 1720: 1716: 1712: 1708: 1704: 1700: 1696: 1692: 1688: 1687: 1672: 1661: 1650: 1639: 1631: 1620: 1609: 1598: 1587: 1576: 1565: 1557: 1536: 1532: 1528: 1524: 1518: 1513: 1509: 1505: 1501: 1497: 1489: 1485: 1481: 1477: 1467: 1463: 1459: 1455: 1451: 1443: 1439: 1435: 1432:morphometric 1427: 1421: 1417: 1415: 1407: 1399: 1369: 1365: 1361: 1357: 1349: 1341: 1333: 1329: 1325: 1321: 1317: 1313: 1309: 1307: 1292: 1281: 1270: 1259: 1238: 1234: 1230: 1226: 1222: 1218: 1214: 1210: 1206: 1205: 1200: 1196: 1192: 1185: 1177: 1173: 1165: 1161: 1157: 1153: 1149: 1145: 1141: 1136:from nearby 1133: 1129: 1124: 1119: 1115: 1111: 1107: 1103: 1093: 1087: 1081: 1075: 1051: 1049: 1044: 1040: 1028: 1024: 1004: 1000: 996: 988: 986: 971: 960: 949: 941: 930: 909: 905: 901: 896:. Using the 893: 889: 885: 881: 877: 873: 869: 865: 861: 852: 846: 840: 828: 824: 820: 816: 804: 788: 783: 775: 770: 765:included 22 762: 756: 746: 742: 738: 728: 718: 714: 710: 706: 702: 700: 692: 686: 680: 676: 672: 668: 664: 658: 654: 650: 646: 642: 638: 634: 630: 626: 615: 611: 607: 604:Raymond Hall 599: 595: 591: 587: 575: 571: 566: 561: 553: 549: 541: 535: 523: 513: 509: 499: 495: 491: 486: 463: 459: 451: 441: 437: 433: 432: 422: 410: 407:DNA sequence 402: 392: 386: 382: 378: 369:united this 367:Raymond Hall 365:and in 1960 362: 356: 352:hantaviruses 338: 331:ungual tufts 322: 303: 298: 297: 296: 289: 277: 265: 261:Alston, 1877 257: 240: 236: 209: 207: 191: 190: 178: 35: 29: 8500:NatureServe 8422:iNaturalist 8355:Wikispecies 8282:Z. brunneus 8166:S. melanops 7991:Oreoryzomys 7950:O. nigripes 7936:O. microtis 7866:O. arenalis 7823:O. superans 7788:O. phaeotis 7781:O. paricola 7760:O. concolor 7646:Nesoryzomys 7620:N. nimbosus 7514:N. palmipes 7500:N. apicalis 7478:N. tenuipes 7471:N. spinosus 7407:M. hammondi 7266:M. audreyae 7220:H. yunganus 7185:H. laticeps 7156:H. sciureus 7106:H. rhabdops 7092:H. intectus 7085:H. fuscatus 7078:H. chapmani 7061:Handleyomys 7049:E. russatus 7014:E. emmonsae 6960:Drymoreomys 6927:C. marinhus 6830:A. donovani 6776:Subfamily: 6735:(rice rats) 6733:Oryzomyini 6349:24 December 6146:27:127–130. 6120:52:833–834. 5899:331:94–127. 5859:47:107–108. 5806:sp. n. and 3699:With claws. 3586:arboviruses 3574:Bayou virus 3490:Raillietina 3375:semiaquatic 3360:oxbow lakes 3346:Pleistocene 3302:Anapophyses 3290:articulates 3258:anteroconid 3212:opisthodont 2962:acrocentric 2950:acrocentric 2938:chromosomes 2657:34 (33–35) 2589:35 (34–36) 2573:Michoacán ( 2547:34 (33–35) 2509:El Salvador 2490:zygomaticus 2367:Description 2230:nasal bones 1856:Pleistocene 1755:Brownsville 1635:Murie, 1932 1561:Eliot, 1904 1231:zygomaticus 1066:, southern 1062:, southern 1045:zygomaticus 1015:, stronger 782:cytochrome 578:, with ten 350:and by two 335:chromosomes 308:semiaquatic 164:Subfamily: 42:Pleistocene 8563:Categories 8224:S. angouya 8214:Sooretamys 8120:P. simplex 8098:P. nivalis 8087:Pennatomys 8060:O. nelsoni 8052:O. gorgasi 7971:O. vegetus 7943:O. moojeni 7901:O. fornesi 7873:O. brendae 7859:O. andinus 7809:O. rutilus 7802:O. roberti 7774:O. mamorae 7753:O. cleberi 7739:O. bicolor 7699:Noronhomys 7686:N. swarthi 7657:N. darwini 7592:N. levipes 7540:Nephelomys 7507:N. grandis 7457:N. paracou 7450:N. musseri 7443:N. minutus 7436:N. guianae 7429:N. dubosti 7385:M. minutus 7334:M. zunigae 7242:L. molitor 7199:H. oniscus 7178:H. acritus 7132:Holochilus 7071:H. alfaroi 7042:E. nitidus 7028:E. legatus 6906:C. goytaca 6896:Cerradomys 6860:A. savamis 6789:Aegialomys 6772:Cricetidae 6629:3537:1–29. 6618:296:1–149. 6469:178:1–705. 6394:3:273–295. 6379:14:99–104. 6021:3:263–270. 5848:7:140–196. 5832:) in Texas 5774:Hesperomys 5767:9:197–208. 5756:28:87–115. 5734:3:263–310. 3716:References 3433:introduced 3428:is short. 3426:life cycle 3350:shrublands 3270:protoconid 3251:anteroloph 3243:anterocone 3238:(ridges). 3228:mesolophid 3220:brachydont 3088:meet. The 2960:is either 2914:unilocular 2854:gatunensis 2833:azuerensis 2380:Population 2332:ochraceous 2310:gatunensis 2306:azuerensis 2294:azuerensis 2282:azuerensis 2278:gatunensis 2266:gatunensis 2262:azuerensis 2254:azuerensis 2250:gatunensis 2246:gatunensis 2214:Canal Zone 2189:gatunensis 2187:, Panama ( 2147:dimidiatus 2102:Bole, 1937 1948:, Mexico ( 1773:Tamaulipas 1498:peninsulae 1370:albiventer 1330:albiventer 1314:albiventer 1009:hantavirus 945:Burt, 1934 791:) and two 580:subspecies 501:Hesperomys 477:Cricetidae 469:Oryzomyini 456:classified 419:Costa Rica 319:shrublands 315:Cricetidae 158:Cricetidae 8195:S. alfari 8038:O. couesi 7979:O. victus 7613:N. moerex 7585:N. keaysi 7578:N. devius 7571:N. childi 7521:N. rattus 7464:N. pictus 7310:Melanomys 7298:M. luciae 7255:Megalomys 7168:Hylaeamys 6992:E. polius 6934:C. scotti 6884:C. cailoi 6746:Kingdom: 6654:Cryptotis 6636:Cryptotis 6267:123:1–57. 6239:476:1–15. 6157:43:1–100. 5870:117:1–69. 3633:Footnotes 3594:reservoir 3447:Parasites 3441:preys on 3274:metaconid 3054:Skull of 2946:autosomal 2934:karyotype 2790:richmondi 2727:cozumelae 2725:Cozumel ( 2622:aquaticus 2594:Unknown ( 2531:Morelos ( 2469:mexicanus 2467:Nayarit ( 2448:mexicanus 2446:Jalisco ( 2427:mexicanus 2394:Hindfoot 2226:braincase 2218:richmondi 2179:Skull of 2151:richmondi 2040:richmondi 1992:cozumelae 1984:cozumelae 1950:cozumelae 1940:Skull of 1868:aquaticus 1842:basin of 1840:Río Verde 1834:The form 1825:aquaticus 1817:aquaticus 1809:gene flow 1801:sympatric 1777:aquaticus 1759:aquaticus 1757:, Texas ( 1749:Skull of 1510:mexicanus 1486:mexicanus 1460:mexicanus 1448:holotypes 1428:mexicanus 1404:Los Reyes 1398:Skull of 1354:Michoacán 1336:from the 1233:, but in 1211:mexicanus 1186:mexicanus 1174:mexicanus 1166:mexicanus 1158:mexicanus 1146:mexicanus 1060:Michoacán 1041:mexicanus 1027:" clade, 842:Melanomys 799:1 of the 795:markers, 723:from the 397:from the 348:parasites 186:Species: 114:Kingdom: 108:Eukaryota 44:to Recent 8594:Oryzomys 8505:2.104170 8492:13000793 8440:10896849 8346:Q1763359 8340:Wikidata 8156:Scolomys 8133:Reigomys 8013:Oryzomys 7490:Nectomys 7419:Neacomys 7397:Mindomys 7232:Lundomys 7213:H. tatei 7021:E. lamia 6948:C. vivoi 6770:Family: 6766:Rodentia 6760:Mammalia 6754:Chordata 6752:Phylum: 6748:Animalia 6662:51:1–30. 6564:Oryzomys 6550:Oryzomys 6510:Mammalia 6435:26:1–30. 6424:245:1–3. 6416:Oryzomys 6386:Oryzomys 6272:Oryzomys 6164:Oryzomys 6138:Oryzomys 6082:Archived 6032:6:1–761. 5853:Oryzomys 5814:(Harlan) 5788:Mammalia 5745:9:47–57. 3485:cestodes 3469:nematode 3342:Taxonomy 3298:thoracic 3294:cervical 3224:mesoloph 3216:bunodont 3200:× 2 = 16 3160:mandible 3129:arteries 3078:lacrimal 3056:Oryzomys 2887:eyeshine 2706:pinicola 2643:peragrus 2575:regillus 2565:161, 148 2562:307, 280 2554:crinitus 2350:Several 2324:Montería 2234:Oryzomys 2202:Oryzomys 2198:Oryzomys 2181:Oryzomys 2133:(11.93% 2123:Oryzomys 2050:Oryzomys 2036:Oryzomys 2024:Oryzomys 1996:Oryzomys 1972:Oryzomys 1942:Oryzomys 1852:Veracruz 1836:peragrus 1751:Oryzomys 1689:Oryzomys 1537:Oryzomys 1529:crinitus 1521:holotype 1514:Oryzomys 1502:crinitus 1490:crinitus 1478:regillus 1464:crinitus 1452:regillus 1440:regillus 1436:crinitus 1418:Oryzomys 1408:regillus 1400:Oryzomys 1366:regillus 1362:crinitus 1350:regillus 1334:crinitus 1326:regillus 1318:crinitus 1190:nominate 1110:species— 1108:Oryzomys 870:Oryzomys 848:Nectomys 815:gene 1 ( 767:synonyms 739:Oryzomys 572:Oryzomys 537:Oryzomys 464:Oryzomys 443:Oryzomys 429:Taxonomy 344:omnivore 249:Synonyms 241:Oryzomys 179:Oryzomys 154:Family: 148:Rodentia 138:Mammalia 128:Chordata 124:Phylum: 118:Animalia 104:Domain: 81:IUCN 3.1 8479:1002659 8414:2438070 7722:Oecomys 6764:Order: 6758:Class: 3883:Tylomys 3355:resacas 3322:humerus 3236:cingula 3197:1.0.0.3 3194:1.0.0.3 3176:incisor 3158:In the 3133:derived 3098:foramen 3082:frontal 3011:process 2985:baculum 2910:stomach 2596:fulgens 2568:37, 35 2533:aztecus 2518:109–194 2515:190–304 2212:in the 2111:Goldman 2058:Ometepe 1976:Cozumel 1946:Cozumel 1899:Tabasco 1854:. Late 1848:Hidalgo 1797:Cameron 1793:Willacy 1677:Merriam 1666:Merriam 1655:Merriam 1625:Merriam 1603:Merriam 1592:Merriam 1482:aztecus 1456:aztecus 1444:aztecus 1358:aztecus 1346:Morelos 1342:aztecus 1322:aztecus 1297:Goldman 1275:Merriam 1264:Merriam 1182:Sinaloa 1138:Nayarit 1072:Jalisco 1064:Morelos 1037:Chiapas 993:Jalisco 976:Merriam 965:Merriam 878:Adh1-I2 866:Adh1-I2 817:Adh1-I2 793:nuclear 515:Tylomys 483:History 306:, is a 220:, 1877) 174:Genus: 144:Order: 134:Class: 79: ( 8544:431917 8531:577260 8518:160399 8453:180337 8401:328432 7795:O. rex 6522: 6494: 6480: 6448: 6414:A new 6365: 6306: 5996: 3608:lists 3545:, and 3493:. The 3471:worms 3435:snake 3338:Sonora 3306:lumbar 3204:molars 3135:. The 3084:, and 3064:couesi 3060:Yaruca 2944:of 56 2940:and a 2924:. The 2903:mammae 2769:couesi 2748:couesi 2685:couesi 2664:couesi 2521:25–33 2270:couesi 2258:couesi 2113:, 1912 2074:palate 2017:Roatán 2005:couesi 1988:couesi 1968:couesi 1964:Belize 1805:couesi 1789:Kenedy 1679:, 1901 1668:, 1901 1657:, 1901 1646:, 1910 1627:, 1901 1605:, 1901 1594:, 1901 1583:, 1877 1581:Alston 1572:, 1891 1442:, and 1340:, and 1324:, and 1310:couesi 1299:, 1915 1288:, 1893 1286:Thomas 1277:, 1901 1266:, 1901 1239:couesi 1215:couesi 1201:couesi 1197:couesi 1193:couesi 1170:jugals 1118:, and 1092:, and 1068:Puebla 1056:Oaxaca 1033:Oaxaca 1013:molars 978:, 1901 967:, 1901 956:, 1897 937:, 1897 823:. The 809:intron 807:) and 803:gene ( 691:; 15, 685:; 14, 679:; 13, 675:; 12, 671:; 11, 667:; 10, 550:couesi 524:couesi 518:. 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