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Crotalus willardi

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measuring one to two feet (30–60 cm) in length. Color patterns are generally a dark brown base with pale or white horizontal striping, but vary slightly among subspecies. It has the odd distinction of being named last as a new species, as there is no mistaking the clear cut ridge that follows
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lasts about four to five months. Females give birth to two to 9 (average five) young in late July or August. Both sexes appear to reach reproductive maturity around 400 mm (16 in) in body (snout to vent) length. Although captive snakes have reproduced annually, wild females probably
405:(v3.1, 2001). Species are listed as such due to their wide distribution, presumed large population, or because they are unlikely to be declining fast enough to qualify for listing in a more threatened category. The population trend was stable when assessed in 2007. 385:
is rarely found outside habitats at high elevation. Wooded mountain ranges, primarily in the southwest, are where this reclusive species is found. Each subspecies’ range is limited to select mountain ranges, making human encounters rare events.
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venom is not as life-threatening as that of other rattlesnakes. No documented deaths have been caused by ridge-nosed rattlesnakes, but pain and discomfort can still result from a rare
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Rattlesnakes are primarily ambush hunters; they coil and lie waiting for prey to approach within striking distance. The diet of
1310: 1482: 914: 302:" region. The IUCN reports this snake's conservation status as being of Least Concern. It is the official state reptile of 425: 1298: 1593: 575: 1521: 1320: 603: 87: 1202:(1905). "An annotated list of a collection of reptiles from southern California and northern Lower California". 551: 873:
Beltz, Ellin (2006). Biographies of People Honored in the Names of the Reptiles and Amphibians of North America
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is the cause of declining numbers, but critical habitat designations (recovery measures) have been proposed.
1057: 581: 315: 253: 1238: 609: 1352: 1216: 322:, is in honor of its discoverer, "Professor" Frank Cottle Willard, a businessman from Tombstone, Arizona. 1390: 1163: 1033: 1124: 1116: 417: 192: 751: 1005: 1425: 1399: 1220: 1029: 747: 497: 807:. Ithaca and London: Comstock Publishing Associates. 1,105 pp. (in 2 volumes). (7th printing, 1985, 1275:. The Peterson Field Guide Series ®. Boston and New York: Houghton Mifflin Company. xiii + 533 pp. 1090: 796: 743: 437: 52: 1508: 1330: 1141: 613: 496:
animals, the young still develop within an egg inside the female snake until their time of birth.
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Brennan, Thomas (2008). "RIDGE-NOSED RATTLESNAKE Crotalus willardi". Retrieved 16 May 2019.
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the countour of its snout, a ridge formed by turning up the scales on the end of its nose.
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is the most recent rattlesnake species to be discovered in the United States. Five
1417: 1503: 1451: 1384: 1127:(2002). "Foraging ecology of the threatened New Mexico Ridge-nosed Rattlesnake ( 1375: 627: 493: 469: 465: 429: 421: 409: 370: 335: 299: 284: 1182:
A Guide to the Rattlesnakes and other Venomous Serpents of the United States
1166:(2001). "Reproduction in northern populations of the ridgenose rattlesnake, 595: 522: 518: 501: 99: 756:
Snake Species of the World: A Taxonomic and Geographic Reference, Volume 1
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The Audubon Society Field Guide to North American Reptiles and Amphibians
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spp.) and lizards, whereas adults feed primarily on mammals and birds.
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Rattlesnakes: Their Habitats, Life Histories, and Influence on Mankind
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is commonly known as the Arizona ridge-nosed rattlesnake, and is the
295: 109: 1346: 758:. Washington, District of Columbia: Herpetologists' League. 511 pp. 630:, Mexico in extreme northwestern Chihuahua and extreme northeastern 461: 159: 149: 492:, meaning it gives birth and does not lay eggs. Contrasting with 270:
ridge-nosed rattlesnake, Willard's rattlesnake, Willard's rattler
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A Field Guide to Western Reptiles and Amphibians, Third Edition
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Hammerson, G.A.; Vazquez DĂ­az, J.; Quintero DĂ­az, G.E. (2007).
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http://www.reptilesofaz.org/Snakes-Subpages/h-c-willardi.html
887:. Baltimore: Johns Hopkins UNiversity Press. xiii + 296 pp. 1152:. Eagle Mountain, Utah: Eagle Mountain Publishing. 596 pp. 532: 1324: 1243:
Reptiles of North America: A Guide to Field Identification
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Beolens, Bo; Watkins, Michael; Grayson, Michael (2011).
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Field Book of Snakes of the United States and Canada
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are secure, the New Mexico ridge-nosed rattlesnake (
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Taxonomy of the New Mexican Ridge-nosed Rattlesnake
805:Handbook of Snakes of the United States and Canada 787:". The Reptile Database. www.reptile-database.org. 428:. Remaining populations are scattered throughout 1331:US Fish & Wildlife Service Species Profile: 1038:The Venomous Reptiles of the Western Hemisphere 1321:International Biopark: Ridge-nosed Rattelsnake 8: 972:2001 Categories & Criteria (version 3.1) 943:Campbell, Sheldon; Shaw, Charles E. (1974). 723:10.2305/IUCN.UK.2007.RLTS.T62253A12584209.en 1213:, new species, pp. 18–19 + Plate III). 500:occurs from late summer to early fall, and 1347: 1184:. Tempe, Arizona: Tricolor Books. 129 pp. 673:Southeastern Arizona, and northern Sonora 217: 61: 42: 31: 1227:. New York: G.P. Putnam's Sons. 365 pp. ( 1012:. New York: Sterling Publishers. 480 pp. 721: 1204:Field Columbian Museum Zoölogical Series 338:are currently recognized, including the 1180:Hubbs, Brian; O'Connor Brendan (2012). 923:Integrated Taxonomic Information System 850:Arizona Rattlesnakes: Rattlesnake Facts 685: 369:is a rather small rattlesnake with all 909: 907: 905: 505:reproduce every second or third year. 398:is classified as Least Concern on the 1097:. New York: Alfred A. Knopf. 743 pp. 468:. The young feed primarily on large 298:. This snake is found mainly in the " 7: 1604:Fauna of the Sierra Madre Occidental 1287:, p. 418 + Plate 52 + Map 188). 1584:IUCN Red List least concern species 1010:Living Snakes of the World in Color 709:IUCN Red List of Threatened Species 648:Chihuahuan ridge-nosed rattlesnake 619:New Mexico ridge-nosed rattlesnake 513:Due to the generally small size of 1245:. New York: Golden Press. 240 pp. 25: 885:The Eponym Dictionary of Reptiles 587:Southern ridge-nosed rattlesnake 563:Del Nido ridge-nosed rattlesnake 436:and the northern part of Mexico. 1338:U.S. Fish & Wildlife Service 1109:, pp. 695–696 + Plate 643). 855:Arizona Game and Fish Department 670:Arizona ridge-nosed rattlesnake 86: 996:Holycross & Goldberg, 2001. 622:The US in extreme southeastern 544: 541: 538: 535: 651:Western Chihuahua and eastern 403:Red List of Threatened Species 330:Originally described in 1905, 1: 1305:Reptarium.cz Reptile Database 978:. Accessed 13 September 2007. 1340:. Accessed 12 December 2007. 1327:. Accessed 12 December 2007. 1317:. Accessed 12 December 2007. 1307:. Accessed 12 December 2007. 426:US Fish and Wildlife Service 1231:, p. 311, Figure 103). 1209:(1): 1-19 + Plates I-III. ( 945:Snakes of The American West 1620: 1599:Reptiles described in 1905 1129:Crotalus willardi obscurus 408:Although four of the five 344:Crotalus willardi willardi 290:found in the southwestern 248:Crotalus willardi willardi 18:Crotalus willardi willardi 1170:(Serpentes: Viperidae)". 626:and extreme southwestern 484:Like other rattlesnakes, 232: 225: 216: 198: 191: 83:Scientific classification 81: 59: 50: 41: 34: 566:Mexico in north-central 987:Holycross et al., 2002. 1131:)". pp. 243–251. 1150:Biology of the Vipers 716:: e.T62253A12584209. 418:endangered subspecies 1265:, pp. 206–207). 590:Mexico in southern 438:Habitat destruction 390:Conservation status 340:nominate subspecies 53:Conservation status 1594:Symbols of Arizona 1196:, pp. 58–61). 1148:(editors) (2002). 917:Crotalus willardi 835:2016-08-11 at the 576:C. w. meridionalis 346:, described here. 1571: 1570: 1543:Open Tree of Life 1405:Crotalus_willardi 1391:Crotalus willardi 1361:Crotalus willardi 1353:Taxon identifiers 1333:Crotalus willardi 1300:Crotalus willardi 1285:Crotalus willardi 1281:978-0-395-98272-3 1263:Crotalus willardi 1229:Crotalus willardi 1211:Crotalus willardi 1194:Crotalus willardi 1190:978-0-9754641-3-7 1168:Crotalus willardi 1107:Crotalus willardi 958:978-0-394-48882-0 897:Crotalus willardi 893:978-1-4214-0135-5 817:Crotalus willardi 785:Crotalus willardi 702:Crotalus willardi 677: 676: 594:and southwestern 545:Geographic range 444:Behavior and diet 367:Crotalus willardi 332:Crotalus willardi 277:Crotalus willardi 263: 262: 236:Crotalus willardi 202:Crotalus willardi 76: 36:Crotalus willardi 16:(Redirected from 1611: 1564: 1563: 1551: 1550: 1538: 1537: 1525: 1524: 1512: 1511: 1499: 1498: 1486: 1485: 1473: 1472: 1460: 1459: 1447: 1446: 1434: 1433: 1421: 1420: 1408: 1407: 1395: 1394: 1393: 1380: 1379: 1378: 1348: 1073: 1055: 1049: 1027: 1021: 1003: 997: 994: 988: 985: 979: 969: 963: 962: 940: 934: 933: 931: 929: 911: 900: 881: 875: 870: 864: 863: 862: 861: 845: 839: 826: 820: 794: 788: 781: 775: 741: 735: 734: 732: 730: 725: 695: 533: 257: 244: 221: 204: 184:C. willardi 91: 90: 70: 65: 64: 46: 32: 27:Species of snake 21: 1619: 1618: 1614: 1613: 1612: 1610: 1609: 1608: 1574: 1573: 1572: 1567: 1559: 1554: 1546: 1541: 1533: 1530:Observation.org 1528: 1520: 1515: 1507: 1502: 1494: 1489: 1481: 1476: 1468: 1463: 1455: 1450: 1442: 1437: 1429: 1424: 1416: 1411: 1403: 1398: 1389: 1388: 1383: 1374: 1373: 1368: 1355: 1344: 1295: 1290: 1082: 1080:Further reading 1077: 1076: 1056: 1052: 1028: 1024: 1004: 1000: 995: 991: 986: 982: 970: 966: 959: 949:Alfred A. 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( 1232: 1214: 1197: 1178: 1162:Holycross AT, 1160: 1158:978-0972015400 1144:, Douglas ME, 1110: 1083: 1081: 1078: 1075: 1074: 1050: 1022: 998: 989: 980: 964: 957: 935: 901: 876: 865: 840: 821: 789: 776: 736: 684: 683: 681: 678: 675: 674: 671: 668: 662: 660:C. w. willardi 656: 655: 649: 646: 645:Klauber, 1949 643: 635: 634: 620: 617: 607: 604:C. w. obscurus 599: 598: 588: 585: 579: 571: 570: 564: 561: 555: 552:C. w. amabilis 547: 546: 543: 540: 537: 530: 527: 510: 507: 481: 478: 445: 442: 420:and listed as 414:C. w. obscurus 391: 388: 379: 376: 363: 360: 348:C. w. willardi 327: 324: 311: 308: 273: 272: 261: 260: 259: 258: 245: 230: 229: 223: 222: 214: 213: 207: 196: 195: 189: 188: 181: 179: 175: 174: 167: 163: 162: 157: 153: 152: 147: 143: 142: 137: 133: 132: 127: 123: 122: 117: 113: 112: 107: 103: 102: 97: 93: 92: 79: 78: 60: 57: 56: 51: 48: 47: 39: 38: 26: 24: 14: 13: 10: 9: 6: 4: 3: 2: 1616: 1605: 1602: 1600: 1597: 1595: 1592: 1590: 1587: 1585: 1582: 1581: 1579: 1562: 1557: 1553: 1549: 1544: 1540: 1536: 1531: 1527: 1523: 1518: 1514: 1510: 1505: 1501: 1497: 1492: 1488: 1484: 1479: 1475: 1471: 1466: 1462: 1458: 1453: 1449: 1445: 1440: 1436: 1432: 1427: 1423: 1419: 1414: 1410: 1406: 1401: 1397: 1392: 1386: 1382: 1377: 1371: 1367: 1366: 1364: 1362: 1358: 1354: 1349: 1345: 1339: 1335: 1334: 1329: 1326: 1322: 1319: 1316: 1315:Virginia Tech 1312: 1309: 1306: 1302: 1301: 1297: 1296: 1292: 1286: 1282: 1278: 1274: 1270: 1267: 1264: 1260: 1259:0-307-47009-1 1256: 1253:(paperback), 1252: 1251:0-307-13666-3 1248: 1244: 1240: 1236: 1233: 1230: 1226: 1222: 1218: 1215: 1212: 1208: 1205: 1201: 1198: 1195: 1191: 1187: 1183: 1179: 1176: 1173: 1169: 1165: 1161: 1159: 1155: 1151: 1147: 1143: 1139: 1136: 1135: 1130: 1126: 1122: 1118: 1114: 1111: 1108: 1104: 1103:0-394-50824-6 1100: 1096: 1092: 1088: 1085: 1084: 1079: 1071: 1070:0-520-21056-5 1067: 1063: 1059: 1054: 1051: 1047: 1046:0-8014-4141-2 1043: 1039: 1035: 1031: 1026: 1023: 1019: 1018:0-8069-6460-X 1015: 1011: 1007: 1002: 999: 993: 990: 984: 981: 977: 976:IUCN Red List 973: 968: 965: 960: 954: 950: 946: 939: 936: 924: 920: 918: 910: 908: 906: 902: 898: 894: 890: 886: 880: 877: 874: 869: 866: 856: 852: 851: 844: 841: 838: 834: 831: 825: 822: 818: 814: 813:0-8014-0463-0 810: 806: 802: 798: 793: 790: 786: 780: 777: 773: 772:1-893777-01-4 769: 765: 764:1-893777-00-6 761: 757: 753: 749: 745: 740: 737: 724: 719: 715: 711: 710: 705: 703: 694: 692: 690: 686: 679: 672: 669: 666: 663: 661: 658: 657: 654: 650: 647: 644: 642: 641: 637: 636: 633: 629: 625: 621: 618: 615: 611: 608: 606: 605: 601: 600: 597: 593: 589: 586: 583: 580: 578: 577: 573: 572: 569: 565: 562: 559: 556: 554: 553: 549: 548: 539:Taxon author 534: 528: 526: 524: 520: 516: 508: 506: 503: 499: 495: 491: 490:ovoviviparous 487: 479: 477: 475: 471: 467: 463: 459: 455: 451: 443: 441: 439: 435: 431: 427: 423: 419: 415: 411: 406: 404: 401: 397: 389: 387: 384: 377: 375: 372: 368: 361: 359: 357: 353: 352:state reptile 349: 345: 341: 337: 333: 325: 323: 321: 317: 316:specific name 309: 307: 305: 301: 297: 293: 292:United States 289: 286: 283: 279: 278: 271: 269: 268:Common names: 265: 264: 255: 249: 246: 242: 237: 234: 233: 231: 228: 224: 220: 215: 210: 205: 203: 197: 194: 193:Binomial name 190: 186: 185: 180: 177: 176: 173: 172: 168: 165: 164: 161: 158: 155: 154: 151: 148: 145: 144: 141: 138: 135: 134: 131: 128: 125: 124: 121: 118: 115: 114: 111: 108: 105: 104: 101: 98: 95: 94: 89: 84: 80: 74: 69: 68:Least Concern 58: 54: 49: 45: 40: 37: 33: 30: 19: 1360: 1343: 1332: 1299: 1284: 1272: 1262: 1242: 1239:Brodie ED Jr 1228: 1224: 1210: 1206: 1203: 1193: 1181: 1174: 1171: 1167: 1149: 1133: 1132: 1128: 1113:Holycross AT 1106: 1094: 1061: 1053: 1037: 1025: 1009: 1001: 992: 983: 967: 947:. New York: 944: 938: 926:. Retrieved 916: 896: 884: 879: 868: 858:, retrieved 849: 843: 824: 816: 804: 792: 784: 779: 755: 744:McDiarmid RW 739: 727:. Retrieved 713: 707: 701: 659: 638: 602: 574: 550: 542:Common name 514: 512: 485: 483: 480:Reproduction 473: 464:, and large 449: 447: 413: 407: 395: 394:The species 393: 382: 381: 366: 365: 347: 343: 331: 329: 319: 313: 276: 275: 274: 267: 266: 247: 235: 201: 199: 183: 182: 170: 35: 29: 1504:NatureServe 1452:iNaturalist 1385:Wikispecies 1269:Stebbins RC 1164:Goldberg SR 1030:Campbell JA 1006:Mehrtens JM 748:Campbell JA 729:19 November 640:C. w. silus 536:Subspecies 515:C. willardi 486:C. willardi 474:Scolopendra 450:C. willardi 396:C. willardi 383:C. willardi 362:Description 1578:Categories 1217:Schmidt KP 1177:: 473-481. 1138:Schuett GW 1125:Douglas ME 1117:Painter CW 1058:Klauber LM 899:, p. 286). 860:2019-05-16 766:(series). 680:References 628:New Mexico 529:Subspecies 498:Copulation 494:viviparous 470:centipedes 466:centipedes 430:New Mexico 422:threatened 410:subspecies 371:subspecies 336:subspecies 300:sky island 146:Suborder: 1146:Greene HW 1142:Höggren M 1121:Barker DG 1087:Behler JL 801:Wright AA 797:Wright AH 774:(volume). 596:Zacatecas 568:Chihuahua 519:hemotoxic 502:gestation 310:Etymology 285:pit viper 178:Species: 160:Viperidae 150:Serpentes 106:Kingdom: 100:Eukaryota 1589:Crotalus 1561:willardi 1509:2.102294 1470:10803183 1376:Q1517877 1370:Wikidata 1271:(2003). 1241:(1982). 1235:Smith HM 1223:(1941). 1221:Davis DD 1093:(1979). 1060:(1997). 1036:(2004). 1034:Lamar WW 1008:(1987). 833:Archived 803:(1957). 754:(1999). 558:Anderson 416:) is an 326:Taxonomy 320:willardi 282:venomous 227:Synonyms 171:Crotalus 156:Family: 140:Squamata 130:Reptilia 120:Chordata 116:Phylum: 110:Animalia 96:Domain: 73:IUCN 3.1 1444:2444415 1325:Biopark 1303:at the 1200:Meek SE 1091:King FW 974:at the 752:TourĂ© T 667:, 1905 624:Arizona 616:, 1974 614:Simmons 592:Durango 584:, 1949 582:Klauber 560:, 1962 458:lizards 454:mammals 434:Arizona 424:by the 378:Habitat 356:Arizona 304:Arizona 288:species 254:Klauber 166:Genus: 136:Order: 126:Class: 71: ( 1548:899047 1522:125876 1483:174320 1431:455016 1279:  1257:  1249:  1188:  1172:Copeia 1156:  1101:  1068:  1044:  1016:  955:  928:16 May 891:  811:  770:  762:  653:Sonora 632:Sonora 612:& 610:Harris 296:Mexico 256:, 1949 243:, 1905 211:, 1905 1535:98430 1496:62253 1465:IRMNG 1457:30705 509:Venom 462:birds 280:is a 1517:NCBI 1491:IUCN 1478:ITIS 1439:GBIF 1418:ZPDR 1277:ISBN 1255:ISBN 1247:ISBN 1186:ISBN 1175:2001 1154:ISBN 1099:ISBN 1066:ISBN 1042:ISBN 1014:ISBN 953:ISBN 930:2007 889:ISBN 815:). ( 809:ISBN 768:ISBN 760:ISBN 731:2021 714:2007 665:Meek 523:bite 400:IUCN 314:The 294:and 241:Meek 209:Meek 1426:EoL 1413:CoL 1400:ADW 1336:at 1323:at 1313:at 1283:. ( 1192:. ( 1134:In: 1105:. ( 895:. ( 718:doi 488:is 354:of 1580:: 1558:: 1556:RD 1545:: 1532:: 1519:: 1506:: 1493:: 1480:: 1467:: 1454:: 1441:: 1428:: 1415:: 1402:: 1387:: 1372:: 1237:, 1219:, 1140:, 1123:, 1119:, 1115:, 1089:, 1072:). 1032:, 951:. 921:. 904:^ 853:, 799:, 750:, 746:, 712:. 706:. 688:^ 525:. 460:, 456:, 432:, 358:. 342:, 318:, 306:. 252:— 1207:7 1048:. 1020:. 961:. 932:. 919:" 915:" 783:" 733:. 720:: 704:" 700:" 472:( 75:) 20:)

Index

Crotalus willardi willardi

Conservation status
Least Concern
IUCN 3.1
Scientific classification
Edit this classification
Eukaryota
Animalia
Chordata
Reptilia
Squamata
Serpentes
Viperidae
Crotalus
Binomial name
Meek

Synonyms
Meek
Klauber
venomous
pit viper
species
United States
Mexico
sky island
Arizona
specific name
subspecies

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