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Dudleya

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37: 1998: 66: 1977: 48: 2086: 2040: 2118: 1914:, are adapted towards pollination via hummingbirds, while short, spreading, yellow flowers favor pollination by insects. Despite the evolution of long flowers towards hummingbirds, the long flowers also show no performance disadvantage in pollination environments exclusively composed of insects. While the long flowers may provide an evolutionary advantage for more effective pollination, more energy is required to produce the larger amounts of floral tissue. 2065: 85: 2530: 2208: 2019: 55: 2174: 235: 629: 4001: 2319: 678:, have a clear, sticky layer on their foliage. This adhesive layer ultimately helps the plant protect its leaves from the sun, much like ones with "white" wax. It accomplishes this solar protection by allowing dust to adhere to the sticky layer on the leaves, which coats the foliage with dust and debris, blocking solar radiation. 2286:
requires is dependent on the location. Plants growing in coastal regions may require little shade at all, whilst plants growing in the deserts, inland valleys and mountains will require shade. The majority of plants in the genus will appreciate north-facing sites and shade during the heat of the day.
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When the underground portion of the plant is lost to root rot, but the rosette and part of the caudex survive, the parts above the decayed roots can be removed cleanly, and planted provided they send out roots. Conversely, should the top of the rosette die but the lower stem still live, the deceased
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During the late fall to early spring period of growing, watering should be done frequently in order to promote good growth and flower development. Plants grown with exposure to rainfall, provided they are in a Mediterranean climate, they may obtain optimum moisture. In contrast, plants grown indoors
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have noticeably smaller seeds; this is likely a result of an evolutionary tradeoff. The investment in growing large allows plants to retain moisture in periods of drought, at the cost of energy placed into reproduction. Other reasons for smaller seeds may also be due to allocation of resources into
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species are widespread and diverse in their range, but are typically found in rock outcroppings, cliff faces, or road cuts, where their leaves help them store water in a setting too dry for most types of plants. Most are small and inconspicuous when not in bloom. The two predominant habitats where
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plants of this genera may be found are by the coast or mountains. Dudleya favor moderate temperatures, summer dormancy, winter precipitation, and rocky habitats, which means they may be found in diverse, disjunct locales from oceanic bluffs on the California coast to sky islands in Arizona.
1945:, the petals begin to fuse at the base, with the flowers becoming tubular and red, due to an association with hummingbird pollinators. This shift in pollinators corresponds to larger nectar volumes and higher energy contents in accordance with the demands of hummingbird pollinators. 2577:, to facilitate the poaching and shipping of the plants. The species was seriously threatened after Mexican soldiers discovered poachers taking nearly 5,000 rosettes in a tractor-trailer. It was suspected the poachers rappelled onto the succulent's location via helicopter, as 3874:
In 2020, posts on social media pointed to shipments of Dudleya candelabrum (candleholder liveforever) harvested from California and destined for sale in Korea. Candleholder liveforever is restricted to San Miguel, Santa Rosa, and Santa Cruz Islands off the coast of Ventura
659:. The wax on the giant chalk dudleya is a surface with some of the highest ultraviolet reflectivity ever discovered in plants. When exposed to water, the wax may coat drops on leaves, preventing their evaporation. The farina is mostly composed of pentacyclic 743:). The differing generic placements of these early descriptions suggest that taxonomic disagreements over the genus had an early start, with some of these generic placements persisting even after the proper description of the genus by Britton and Rose. 3562:
In a recent greenhouse study, conducted in Santa Cruz, California, by Stephen McCabe and Rachel Ormes, D. parva was rooted from the leaves of the plant, making it the only known species in the Dudleya subgenus to exhibit this behavior (McCabe in litt.
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During the dormant period, from late spring to fall, watering should be limited to bi-weekly intervals, but not completely cut out. In nature, plants may completely desiccate themselves during the dormant period. Deciduous members of the genus in the
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In 2020, for example, posts on social media suggested that shipments of candleholder dudleya (Dudleya candelabrum), a species restricted to San Miguel, Santa Rosa, and Santa Cruz Islands off the coast of Ventura County, were destined for sale
967:. In the book, Thompson made numerous changes and adjustments to species, and described several new species. The book included a large number of photographs, extensive descriptions, and horticultural practices for the care of 2339:
seeds are crescent shaped and near-microscopic, and can be collected by taking the dried flower buds and crushing the ovules inside, which can contain hundreds of seeds. Seeds can be sown into a soilless medium, such as
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are very susceptible to mold and fungi. How quickly the medium drains should be determined by the amount of shade in the location, which affects the rate of evaporation that the soil will go through. Thus, well-shaded
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Margulies, J. D. (2020). Korean ‘Housewives’ and ‘Hipsters’ Are Not Driving a New Illicit Plant Trade: Complicating Consumer Motivations Behind an Emergent Wildlife Trade in Dudleya farinosa.
2611:) was also targeted by poachers in numerous large-scale operations. Although not particularly rare, the size of the poaching operation pose a serious ecological threat. Starting in 2017, the 3156:
The green "form" of this species should likely be recognized as a separate species but the description by P.H. Thomson apparently lacks a designated type and may be invalidly published.
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are broad, white to yellow, and spreading, with bumblebees and bees as primary pollinators, and hummingbirds as occasional pollinators. As these species grade towards the subgenus
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Growing mediums should attempt to imitate the preferred soil of each species in nature. The most essential element for the medium is good drainage. Good drainage is important, as
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Rosettes emerge out of a long, horizontal root. This is commonplace when the plant has been decapitated on a cliff face, leaving only the long roots, which send out rosettes.
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Clay is preferable to plastic pots when growing the plants due to the advantages with water drainage. Plastic pots may also kill the plants by overheating the roots.
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that evolved recently with the formation of the summer-dry climate in California. This was evidenced by the similarities of seed-surface ornamentation between
2377:, when the plant has its main rosettes burned off, new ones will emerge from either the stem or the roots, forming a new plant within a single growing season. 2085: 2616: 1966:-type members produce the largest seeds in the genus. These larger seeds are not distributed very far, which contributes to the restricted ranges of the 3667: 1902:, although this has been inferred in some species only from flower morphology. The genus exhibits a wide array of diverse flower varieties, even within 3538: 3290:
Dudleyas are also known as "liveforevers" due to their individual longevity and hardiness, a somewhat deceptive name given the rarity of many species.
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At the edge of California and the edge of existence: How siemprevive de Isla de Cedros (Dudleya pachyphytum ) was poached ever closer to extinction.
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from state or private lands without a permit. The California Native Plant Society and conservationists have also initiated propagation programs to
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also the type species for this subgenus). The flowers have the petals spreading at the middle, and are not arranged connately like in the subgenus
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should be planted at an angle. This allows accumulated water to drain from the nestlike center of the plant, thus preventing microbial decay.
47: 4578: 2480:, in their early stages, were used as food, with a sweet flavor and juicy texture. The roots were pounded up and soaked in water, used as an 530:, the plants of this group usually have narrow leaves, often elliptic to round in cross-section, resembling fingers; hence common names like 2403:
portion may be removed, a depression will remain in the caudex where the decayed rosette was removed, and at the edges new buds will sprout.
515:). When the flowers go to fruit, they retain the tight and tubular character. The most recognizable plants of this first group would be the 3663: 3765: 3397: 2204:
should not be watered from directly above, as this may damage their chalky coating known as farina, which is present on numerous species.
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is a relatively obscure genus, in comparison to other, more widely-cultivated succulents; converging interests, by succulent collectors,
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Henricot, B. (2008). Occurrence of powdery mildew (Erysiphe sp.) on Echeveria spp., Crassula spp., Cotyledon and Dudleya in the UK.
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was characterized by a different vegetative structure and more niche adaptations, primarily corm-like stems and deciduous leaves.
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Spectral Properties of Heavily Glaucous and Non-Glaucous Leaves of a Succulent Rosette-Plant, Thomas W. Mulroy, Oecologia, 1979,
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Carlson, Cheri (April 18, 2019). "First a wildfire, then poachers. How a rare Ventura County plant has been nearly wiped out".
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The genus is traditionally divided into three subgenera, two of which were formerly their own genera. The subgenera consist of
3945:"Governor Newsom signs CNPS-sponsored AB 223 into law, providing important new protections for California's imperiled dudleya" 3918: 2770:"Linnaeus's folly -phylogeny, evolution and classification of Sedum (Crassulaceae) and Crassulaceae subfamily Sempervivoideae" 1121:, although it may not be the true sister taxon, as larger samplings are needed. Berger's hypothesis of a relationship between 4211: 2991: 1673: 1910:, although some individuals may have sterile anthers. Plants with long, tubular, reddish flowers, mostly in the subgenus 4568: 2662: 1685: 1455: 1407: 996: 1431: 1383: 4063: 2680: 2291:
is grown during a freeze or snow, it should be shaded as not to damage the plant, as a quick thaw may be detrimental.
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containing 60 species, of which 41 were newly described by Britton and Rose. The two also defined the related genera
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Amoroso, D. M., & Wilson, P. (2018). Ten Cases of Divergence in the Seedling Ecology of Dudleya (Crassulaceae).
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vegetative reproduction via pup rosettes, instead of sexual reproduction. In contrast, the small, inconspicuous,
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research is still at an early stage in the genus, and is complicated by the fact that many species are becoming
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Uhl, Charles H. (1994). "INTERGENERIC HYBRIDS IN THE MEXICAN CRASSULACEAE: II. DUDLEYA (AND PLATE TECTONICS)".
1133:-type taxa could be basal to the genus. However, no truly conclusive evidence of significant divergence within 935:
far enough that intergeneric reproductive success was unlikely. Despite Uhl's research, he still believed that
716: 647:, or farina. This waxy coating is usually white, chalky, and mealy, and is the namesake in many epithets, like 4382: 4263: 2399:) sprout roots out of stoloniferous plantlets, yet this can be difficult or impossible to induce in captivity. 852:, a botanist with a longstanding interest in the genera, offered a new revision of the taxa. Moran recognized 3655: 4128: 2942: 2356: 1907: 960: 3526: 971:
derived from a long interest growing, visiting and collecting the plants. However, he failed to follow the
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in Oregon and California, has been shown in multiple phylogenetic reconstructions to be a sister taxon to
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before flowering, and widely spreading flowers and fruit. The spreading flowers, which resemble those of
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revealed that two fishermen were killed after a dispute emerged over the trafficking of the rare plant.
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Manheim Jr, B. S., & Mulroy, T. W. (1978). Triterpenoids in epicuticular waxes of Dudleya species.
308:. Yet, despite their dramatic variations in appearance, most species readily hybridize. The flowers of 4462: 4343: 3739: 3197:
Yost, J. M.; Bontrager, M.; McCabe, S. W.; Burton, D.; Simpson, M. G.; Kay, K. M.; Ritter, M. (2013).
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As Moran and Uhl conducted more cytological and taxonomic research on the genus, it became clear that
4159: 4089: 3560:. Ventura Fish and Wildlife Office, Ventura, California: U.S. Fish and Wildlife Service. March 2015. 3554:
Dudleya abramsii ssp. parva ( = Dudleya parva )(Conejo Dudleya) 5-Year Review: Summary and Evaluation
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Thiede, J. (2004). The genus Dudleya Britton & Rose (Crassulaceae): its systematics and biology.
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The rosette of the plant divides into two, eventually branching off to develop two separate rosettes.
1808: 1622: 1109: 729: 701: 648: 516: 341:, in the California region, five million years ago. Early botanists classified the larger species as 2426:
methods are also available, and becoming more advanced. Currently, plant tissue culture is used for
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as a means to deter poachers. On September 28, 2021, governor Gavin Newsom signed AB-223 into law.
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occupies a nearly inaccessible habitat. In 2019, the deaths and injuries of several fishermen from
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is characterized by broad, flattened leaves, and tight petals on the flower that form a tube (
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have failed. Uhl came to the conclusion that there are probably no intergeneric hybrids with
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formed two distinct lineages within Echeverioideae. Joachim Thiede instead proposed that
3811:"Plant smugglers take 'massive toll' on California's Dudleya farinosa succulent species" 3580:"In-vitro cultivation of liveforevers (genus Dudleya) to reduce poaching and extinction" 3357: 2590: 2501: 2249:
Fertilizers may be used to maintain good color and growth, but they should be diluted.
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enthusiasts and gardeners alike, have led to the wider cultivation of many species as
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During the summer months, 50% shade may be beneficial for plants. If a cold-tolerant
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to "tighten the gums." The roots were also boiled whole as a decoction for asthma.
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radiating northward from Mexico, evidenced by the similar tubular corollas, while
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Uhl, C. H., & Moran, R. (1953). The cytotaxonomy of Dudleya and Hasseanthus.
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will not reproduce from leaves, with the leaves simply dying after being severed.
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upended many of the early taxonomic classifications, with the newly-minted genus
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cannot be propagated through leaf cuttings. Propagation is mostly achieved via
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or in greenhouses should be watered to ensure that the plant maximizes growth.
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freely hybridize within themselves and each other, all attempts at hybridizing
3098:"Growing and Appreciating California Succulents in the Garden and in the Wild" 3097: 2481: 2370:
of old leaves around the primary rosettes, forming as many as 10 new rosettes.
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forms and, when fruiting, are filled with tiny, ovoid-crescent-shaped seeds.
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were between the leaf shape, floral structure and petal orientation, whilst
635:, which has the highest ultraviolet reflectivity ever discovered in a plant. 582: 512: 412: 343: 313: 305: 297: 4527: 4395: 4276: 1293:– Short-leaved liveforever, short-leaved dudleya. Formerly a subspecies of 959:
Around 1993, one of the only literary treatments of the genus was written,
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feature. The seeds are among the largest in the genus. The type species is
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CNPS 2018 Conservation Conference: Horticulture as a Part of Conservation.
3766:"Reconoce Marina extracción y tráfico de "siempre viva" en Isla de Cedros" 2573:. In 2016, Korean nationals began moving to Bahia Tortugas, a locality in 4522: 4429: 4390: 4310: 4271: 4033: 3837:"A poaching conspiracy is playing out on Northern California's coastline" 2647: 2509: 2445: 2391:
may send out stoloniferous rosettes, but they do not root. Only does the
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is their waxy coating. Numerous species have their foliage covered in an
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may still prove true, as in addition to morphological similarities, the
4449: 4330: 4146: 3141:"Annotated checklist of the vascular plants of Baja California, Mexico" 3102: 3053: 2758:
Britton, N. L. & Rose, J. N.: Dudleya Britton & Rose, gen. nov.
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may refer to the longevity and hardiness of some plants in the wild.
1008: 548:, and some have campanulate flowers that appear intermediate between 419:, where hybridisation and cultivation of succulents is very popular. 4475: 4010: 3579: 3244:(Crassulaceae), a new species from northern Baja California, Mexico" 3199:"Phylogenetic relationships and evolution in Dudleya (Crassulaceae)" 3591: 4356: 4185: 4107: 3712: 2705: 2631: 2528: 2317: 2259: 2206: 2172: 2116: 1137:
exists, and it remains unclear which groups are basal or derived.
808: 627: 587: 329: 273: 96: 2126:, which grows on Miocene volcanic substrates on Santa Rosa Island 876:
as a separate genus, but realized that it was closely related to
2992:"Meet the Endangered Plant Named after Rock Legend Jimi Hendrix" 2406:
Propagation via leaves. Only certain species, like those of the
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species complex: can generalization promote floral divergence?"
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According to nursery owners responsible for legally exporting
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Research in the 21st century has supported the hypothesis of
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Orange County Chapter of the California Native Plant Society
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Messerschmid, Thibaud; Klein, Johannes T. (September 2020).
581:-like stems, with small, inconspicuous leaves that usually 435:
poaching through propagation programs and protection laws.
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populations of plants. Poached plants are often shipped to
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Said to have been published in the December 2016 issue of
1360:– Knife-leaved liveforever, maritime succulent liveforever 312:
have parts numbered in fives, with the petals arranged in
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material, including references to the lost continents of
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wildfires. However, one of the most critical threats to
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this grouping is typically distinguished by underground
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grouping should not be watered at all during dormancy.
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Bornstein, Carol; Fross, David; O'Brien, Bart (2005).
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nationals have both been arrested in the smuggling of
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who were on Cedros Island was allegedly the result of
3338:"Variation in pollination performance gradients in a 3324:
An Ethnobotany of Baja California's Kumeyaay Indians.
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Thiede, J., & Eggli, U. (2007). Crassulaceae. In
1490:– Bluff lettuce, powdery liveforever, powdery dudleya 832:
was believed to occupy a transitory position between
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propagation in commercial and conservation settings.
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were related closer to each other than they were to
4419: 4300: 4023: 3919:"Plant poachers threaten California's biodiversity" 3174:(pp. 83-118). Springer, Berlin, Heidelberg. Chicago 3018:"Two species of Dudleya were named in recent years" 1921:also corresponds with the aforementioned groups of 828:, primarily flowers with broadly-spreading petals. 711:The same species would later be described again as 3902:: 30 – via California Native Plant Society. 1273:– Arizona chalk dudleya. Formerly a subspecies of 844:Reid Moran, Charles H. Uhl and early phylogenetics 769:, with 4 species. The primary differences between 1792:to the pencil-shaped leaves of the type species, 1626:– Chalk lettuce, chalk dudleya, chalk liveforever 296:. The species come in many forms, some large and 2315:by seed, or in nurseries, plant tissue culture. 955:Modern classifications and phylogenetic research 696:The earliest species of the genus described was 353:, while the geophytic species were placed under 3870:California State Senate Public Safety Committee 1811:, a surgeon and collector of California flora. 605:. Perhaps the most widely known members of the 2550:), which was nearly wiped out during the 2013 288:, consisting of about 68 taxa in southwestern 3527:"Defeat Dudleya Poaching Through Propagation" 3411:Wilson, Bert; Wilson, Celeste (9 July 2013). 3275:"Proposed Bill Would Stop Succulent Poaching" 3185:Evolution and systematics of the Crassulaceae 1773:, the first head of the botany department at 1482:– Fingertips, lady fingers, San Diego dudleya 8: 3139:Rebman, J. P.; Gibson, J.; Rich, K. (2016). 1285:– Blochman's liveforever, Blochman's dudleya 1265:– Orcutt's liveforever, tapertip liveforever 1103:is much closer to North American members of 973:International Code of Botanical Nomenclature 3917:Jensen, Nick; Foy, Patrick (23 July 2021). 570:, but still remain tight towards the base. 4011: 3390:"Native Succulents for Water-Wise Gardens" 2617:California Department of Fish and Wildlife 1313:– Coast dudleya, sea lettuce, sand lettuce 1034:Later research challenged the notion that 233: 20: 3484:. Los Olivos, California: Cachuma Press. 3365: 3238:Dodero, M. W. and M. G. Simpson. (2012). 3123:Cactus & Succulent Journal of America 2785: 1080:, and the reproductive isolation between 327:, from ancestors in the stonecrop genus, 2182:, found on the Baja California peninsula 1305:– Britton's dudleya, giant chalk dudleya 812:. This was in part due to a belief that 639:One of the most famous features of some 566:are more spreading than in the subgenus 469: 333:. The ancestors radiated southward from 4574:Flora of the Southwestern United States 3713:"Militares decomisan plantas endemicas" 3482:California Native Plants for the Garden 3446:from the original on 30 September 2021. 3440:Henry Shaw Cactus and Succulent Society 3400:from the original on 27 September 2015. 2751: 2687:, which would make it illegal to poach 2496:are threatened by urban development in 1972: 939:had evolved northward from Mexico with 704:in 1803, which would later be known as 3134: 3132: 3110:from the original on 13 December 2021. 2565:) is a rare and extremely specialized 1175:– San Bernardino Mountains liveforever 1046:evolved completely independently from 3835:Robertson, Michelle (20 March 2019). 3693:from the original on 10 December 2019 3681:Vargas, Elizabeth (9 December 2019). 3573: 3571: 3520: 3518: 3469:from the original on 25 October 2007. 3383: 3381: 3379: 3377: 3318: 3316: 3314: 3183:Hart, H. T., & Eggli, U. (1995). 3166: 3164: 3016:McCabe, Stephen W. (9 January 2017). 2937: 2935: 2933: 2931: 2929: 2927: 2270:has also been recorded in cultivated 1953:Seed morphology also differs between 1736:– Alabaster plant, Island liveforever 1050:, evolving from relatives within the 7: 3740:"CJNG en desaparición de pescadores" 3664:University of California, Santa Cruz 3541:from the original on 17 August 2021. 3148:San Diego Society of Natural History 3034:from the original on 28 January 2021 2968: 2966: 2925: 2923: 2921: 2919: 2917: 2915: 2913: 2911: 2909: 2907: 2879: 2877: 2875: 2873: 2871: 2869: 2867: 2853: 2851: 2849: 2847: 2845: 2843: 2841: 2839: 2821: 2819: 2817: 2815: 2813: 2811: 2809: 2807: 2805: 2522:paying lucrative prices for certain 2512:, partially caused by a demand from 1538:– Baja liveforever, rock liveforever 868:(which was changed to just subgenus 824:possessed aesthetic similarities to 609:group are the critically endangered 448:This genus is the only taxon of the 3943:Bryant, David (28 September 2021). 3423:from the original on 16 March 2015. 3279:the Northcoast Environmental Center 3273:Griffith, Caroline (28 June 2021). 3086:. Desert Plant Life. 14: 191. 1942. 2464:leaves were chewed on to alleviate 2452:region utilized the genus for both 1474:– San Gabriel Mountains liveforever 4515:urn:lsid:ipni.org:names:30012235-2 3623:Frontiers in Ecology and Evolution 3457:Hodgkiss, R.J. (5 February 2021). 3025:International Crassulaceae Network 2990:Platt, John R. (12 January 2017). 2626:being shipped out of the country. 2613:U.S. Customs and Border Protection 2538:, which was threatened by poachers 1712:– Santa Barbara Island liveforever 1107:than to Echeverioideae. The plant 765:, which contained 12 species, and 753:'s revision of the North American 373:'s investigations into the genus. 14: 3729:McCabe, S.W. (September 24, 2019) 3670:from the original on 22 May 2019. 3654:Townsend, Peggy (25 April 2019). 3394:San Joaquin UCCE Master Gardeners 2827:Cactus and Succulent Journal (US) 2589:trafficking, a conflict with the 2282:The amount of shade a species of 1886:Pollinators and flower morphology 1634:– Sierra de la Laguna liveforever 526:Formerly segregated as the genus 4383:urn:lsid:ipni.org:names:298178-2 4264:urn:lsid:ipni.org:names:297170-2 3999: 3994:Treatment from the Jepson Manual 3974:Dudleya and Hassenthaus Handbook 3367:10.1111/j.1365-2435.2005.01009.x 3302:Britton, N. L. & Rose, J. N. 2947:Dudleya and Hasseanthus Handbook 2084: 2063: 2038: 2017: 1996: 1975: 1440:– Santa Monica Mountains dudleya 1027:and the scientifically accepted 965:Dudleya and Hasseanthus Handbook 796:into a sectional ranking within 357:. Taxonomic efforts, started by 284:plants in the stonecrop family, 83: 64: 53: 46: 35: 3949:California Native Plant Society 3872:– via California Senate. 3809:Callahan, Mary (9 March 2019). 3798:California Native Plant Society 3535:California Native Plant Society 2685:California Native Plant Society 2668:In response to the poaching of 1586:– Santa Cruz Island liveforever 1562:– San Benito Island liveforever 1464:– Transverse ranges liveforever 562:. The flowers and fruit of the 395:. In the wild, many species of 3976:. Bonsall Publications. 1993. 3889:"New Protections for Dudleya?" 3578:Alison, Kevin (25 June 2020). 3525:Chesnut, John (October 2018). 2619:, discovered large amounts of 2444:The indigenous peoples of the 2246:must have excellent drainage. 2158:. The diversity of species of 979:new taxa, forgoing designated 381:and over-harvested (poached). 16:North American succulent genus 1: 3388:Zane, Nadia (9 August 2015). 3096:Heiple, Paul (9 March 2021). 444:Characteristics and subgenera 4579:Flora of Northwestern Mexico 3887:Jensen, Nick (Spring 2021). 2554:, was targeted by poachers. 2359:may occur in multiple ways: 2299:Unlike their related genera 1223:– Santa Clara Valley dudleya 573:Formerly known as the genus 407:often threaten particularly 2890:Jepson Flora Project (eds.) 2886:"Dudleya, in Jepson eFlora" 2366:Plantlets will form in the 1650:– Red flowering liveforever 1404:– Pierpoint Springs dudleya 1392:– San Gabriel River dudleya 337:during the creation of the 4600: 3322:Wilken, Michael A. (2012) 3172:Flowering Plants· Eudicots 2859:American Journal of Botany 2741:, who described the genus. 2661:), native to the northern 1803:The name of the subgenera 1784:The name of the subgenera 1704:– Laguna Beach liveforever 1329:– Candleholder liveforever 1249:– Todos Santos liveforever 1241:– White-flower liveforever 872:). Moran still recognized 864:as a subgenera along with 3861:Bradford, Steven (2021). 3326:Retrieved 13 October 2021 2798:– via ResearchGate. 2673:California State Assembly 2559:Cedros Island liveforever 2410:subgenus and others like 1917:The flower morphology of 1906:. The flowers are mostly 1821:is also named after him. 1769:The genus is named after 1670:– Gila County liveforever 1642:– Playa Maria liveforever 1321:– Punta Banda liveforever 599:, which is a synonym for 474: 241: 232: 227: 220: 210: 203: 80:Scientific classification 78: 28: 23: 3660:UC Santa Cruz Newscenter 3606:Santa Ysabel Ethnobotany 3463:The Succulent Plant Page 3459:"Dudleya - Crassulaceae" 3218:10.1600/036364413X674760 2949:. Bonsall Publications. 2593:, or both. In 2020, the 2393:Laguna Beach Liveforever 2106:Distribution and habitat 1720:– Variegated liveforever 1618:– Few-flower liveforever 1554:– Lanceleaf liveforever. 903:was also a subgenera of 717:Joseph Franz von Jacquin 3610:San Diego Museum of Man 2693:oversaturate the market 2357:Vegetative reproduction 1658:– Panamint liveforever 1594:– Cedros Island dudleya 1546:– Hendrix's liveforever 1530:– Guadalupe liveforever 1498:– La Mision liveforever 1337:– Coronados liveforever 1257:– Anthony's liveforever 1113:, which co-exists with 856:as being distinct from 666:Other members, such as 617:, famous for its name. 3863:"AB 223 Bill Analysis" 3413:"Dudleya pulverulenta" 3336:Aigner, P. A. (2005). 2884:McCabe, Stephen Ward. 2735:Nathaniel Lord Britton 2650:and weathered leaves. 2646:quality of their long 2539: 2460:purposes. The tender, 2328: 2217: 2183: 2127: 1728:– Verity's liveforever 1694:– Panamint liveforever 1602:– Palmer's liveforever 1570:– Many-stemmed dudleya 1452:– Diablo range dudleya 1380:– Agoura Hills dudleya 1233:– Vizcaino liveforever 788:revised the status of 747:Nathaniel Lord Britton 725:Echeveria pulverulenta 636: 597:Hasseanthus variegatus 452:tribe to have evolved 363:Nathaniel Lord Britton 4497:Paleobiology Database 4008:at Wikimedia Commons 2721:, a genus similar to 2709:, a genus related to 2532: 2502:anthropogenic-induced 2321: 2210: 2176: 2134:can be found include 2120: 1848:iempreviva, mezcalito 1771:William Russel Dudley 1744:– Colonet liveforever 1527:Dudleya guadalupensis 1416:– Coast Range dudleya 1368:– Canyon liveforever 1163:– Abrams' liveforever 1007:directly induced the 719:in 1811, and in 1840 631: 461:Dudleya, Stylophyllum 323:The genus evolved as 3770:Investigaciones Zeta 3744:Investigaciones Zeta 3434:Kasten, Roy (2004). 2655:candleholder dudleya 2544:Verity's liveforever 2424:Plant tissue culture 2385:stoloniferous growth 2383:Plants multiply via 2352:Asexual reproduction 1881:Reproductive biology 1837:Dudleya, liveforever 1807:was in honor of Dr. 1682:– Desert liveforever 1623:Dudleya pulverulenta 1506:– Gates' liveforever 1428:– Marcescent dudleya 1352:– Thick-leaf dudleya 1275:Dudleya pulverulenta 1211:– Mouse-gray dudleya 1110:Sedum spathulifolium 802:, while merging the 735:Echeveria lanceolata 730:Dudleya pulverulenta 702:Adrian Hardy Haworth 698:Cotyledon caespitosa 611:short-leaved dudleya 369:, proper, following 260:, commonly known as 4569:Crassulaceae genera 3641:Ventura County Star 3604:Hedges, K. (1986). 3594:– via Zenodo. 3417:Las Pilitas Nursery 3358:2005FuEco..19..681A 3242:Dudleya crassifolia 2996:SCIENTIFIC AMERICAN 2683:, sponsored by the 2659:Dudleya candelabrum 2599:Second Naval Region 2575:Baja California Sur 2563:Dudleya pachyphytum 2492:Several species of 2468:, or used to treat 2440:Kumeyaay and Paipai 2397:Dudleya stolonifera 2332:Sexual reproduction 2164:Southern California 2078:Dudleya blochmaniae 1890:The pollinators of 1775:Stanford University 1701:Dudleya stolonifera 1639:Dudleya rigidiflora 1591:Dudleya pachyphytum 1567:Dudleya multicaulis 1349:Dudleya crassifolia 1342:Dudleya cedrosensis 1326:Dudleya candelabrum 1318:Dudleya campanulata 1282:Dudleya blochmaniae 1199:– limestone dudleya 1023:, and criticism of 1005:divine intervention 657:giant chalk dudleya 653:powdery liveforever 559:Dudleya campanulata 521:giant chalk dudleya 471: 454:sympodial branching 399:are vulnerable, as 316:, star-shaped, and 3815:The Press Democrat 3790:Uhler, M. (2018). 3656:"Succulent savior" 3644:. August 16, 2021. 3346:Functional Ecology 3306:Britton & Rose 2892:. Jepson Herbarium 2739:Joseph Nelson Rose 2542:The population of 2540: 2498:Coastal California 2416:, will root. Most 2329: 2262:are main pests of 2218: 2184: 2128: 1990:Dudleya caespitosa 1794:Stylophyllum edule 1741:Dudleya viridicata 1615:Dudleya pauciflora 1578:– Cape liveforever 1551:Dudleya lanceolata 1522:– Greene's dudleya 1514:– Munchkin dudleya 1471:Dudleya densiflora 1310:Dudleya caespitosa 1290:Dudleya brevifolia 1076:, which is unlike 751:Joseph Nelson Rose 740:Dudleya lanceolata 707:Dudleya caespitosa 637: 470: 359:Joseph Nelson Rose 339:dry summer climate 213:Dudleya lanceolata 4551: 4550: 4233:Open Tree of Life 4017:Taxon identifiers 4004:Media related to 3772:. 20 January 2021 3746:. 9 December 2019 3206:Systematic Botany 2974:Systematic Botany 2787:10.1002/tax.12316 2615:, along with the 2324:Dudleya variegata 2196:In horticulture, 2179:Dudleya albiflora 2099:Dudleya variegata 2097:- The flowers of 2076:— The flowers of 2051:— The flowers of 2030:— The flowers of 2009:— The flowers of 1988:— The flowers of 1933:. The flowers in 1904:species complexes 1717:Dudleya variegata 1583:Dudleya nesiotica 1543:Dudleya hendrixii 1302:Dudleya brittonii 1270:Dudleya arizonica 1262:Dudleya attenuata 1238:Dudleya albiflora 1230:Dudleya acuminata 1187:– Betty's dudleya 1009:chromosome counts 927:, believing that 816:had evolved from 687:Taxonomic history 633:Dudleya brittonii 621: 620: 615:Hendrix's dudleya 602:Dudleya variegata 393:ornamental plants 253: 252: 199: 4591: 4584:Succulent plants 4544: 4543: 4531: 4530: 4518: 4517: 4505: 4504: 4492: 4491: 4479: 4478: 4466: 4465: 4453: 4452: 4440: 4439: 4438: 4412: 4411: 4399: 4398: 4386: 4385: 4373: 4372: 4360: 4359: 4347: 4346: 4334: 4333: 4321: 4320: 4319: 4293: 4292: 4280: 4279: 4267: 4266: 4254: 4253: 4241: 4240: 4228: 4227: 4215: 4214: 4202: 4201: 4189: 4188: 4176: 4175: 4163: 4162: 4150: 4149: 4137: 4136: 4124: 4123: 4111: 4110: 4098: 4097: 4085: 4084: 4072: 4071: 4059: 4058: 4057: 4044: 4043: 4042: 4012: 4003: 3960: 3959: 3957: 3955: 3940: 3934: 3933: 3931: 3929: 3914: 3908: 3907: 3893: 3884: 3878: 3877: 3867: 3858: 3852: 3851: 3849: 3847: 3832: 3826: 3825: 3823: 3821: 3806: 3800: 3788: 3782: 3781: 3779: 3777: 3762: 3756: 3755: 3753: 3751: 3736: 3730: 3727: 3721: 3720: 3717:MAS Noticias BCS 3709: 3703: 3702: 3700: 3698: 3678: 3672: 3671: 3651: 3645: 3636: 3630: 3619: 3613: 3602: 3596: 3595: 3575: 3566: 3565: 3559: 3549: 3543: 3542: 3522: 3513: 3502: 3496: 3495: 3477: 3471: 3470: 3454: 3448: 3447: 3431: 3425: 3424: 3408: 3402: 3401: 3385: 3372: 3371: 3369: 3333: 3327: 3320: 3309: 3299: 3293: 3292: 3287: 3285: 3270: 3264: 3257: 3251: 3236: 3230: 3229: 3212:(4): 1096–1104. 3203: 3194: 3188: 3181: 3175: 3168: 3159: 3158: 3145: 3136: 3127: 3126: 3118: 3112: 3111: 3093: 3087: 3084:(Davidson) Moran 3078: 3072: 3071:(10), 1799-1800. 3061: 3055: 3050: 3044: 3043: 3041: 3039: 3033: 3022: 3013: 3007: 3006: 3004: 3002: 2987: 2981: 2970: 2961: 2960: 2943:Thomson, Paul H. 2939: 2902: 2901: 2899: 2897: 2881: 2862: 2855: 2834: 2823: 2800: 2799: 2789: 2765: 2759: 2756: 2609:Dudleya farinosa 2535:Dudleya farinosa 2500:and Mexico, and 2095: 2088: 2074: 2067: 2049: 2042: 2028: 2021: 2011:Dudleya anthonyi 2007: 2000: 1986: 1979: 1809:Hermann E. Hasse 1759:– Sticky dudleya 1709:Dudleya traskiae 1610:– Conejo dudleya 1575:Dudleya nubigena 1559:Dudleya linearis 1487:Dudleya farinosa 1357:Dudleya cultrata 1254:Dudleya anthonyi 1216:Dudleya abramsii 1204:Dudleya abramsii 1192:Dudleya abramsii 1180:Dudleya abramsii 1168:Dudleya abramsii 1156:Dudleya abramsii 1149:Dudleya abramsii 1141:Selected species 1029:age of the Earth 999:, a belief that 985:pseudoscientific 645:epicuticular wax 624:Epicuticular wax 545:Dudleya traskiae 472: 421:Conservationists 401:land development 294:Guadalupe Island 242:Distribution of 237: 191: 88: 87: 68: 57: 50: 39: 21: 4599: 4598: 4594: 4593: 4592: 4590: 4589: 4588: 4554: 4553: 4552: 4547: 4539: 4534: 4526: 4521: 4513: 4508: 4500: 4495: 4487: 4482: 4474: 4469: 4461: 4456: 4448: 4443: 4434: 4433: 4428: 4415: 4407: 4402: 4394: 4389: 4381: 4376: 4368: 4363: 4355: 4350: 4342: 4337: 4329: 4324: 4315: 4314: 4309: 4296: 4288: 4283: 4275: 4270: 4262: 4257: 4249: 4244: 4236: 4231: 4223: 4218: 4210: 4205: 4197: 4192: 4184: 4179: 4171: 4166: 4158: 4153: 4145: 4140: 4132: 4127: 4119: 4114: 4106: 4101: 4093: 4088: 4080: 4075: 4067: 4062: 4053: 4052: 4047: 4038: 4037: 4032: 4019: 3969: 3967:Further reading 3964: 3963: 3953: 3951: 3942: 3941: 3937: 3927: 3925: 3916: 3915: 3911: 3891: 3886: 3885: 3881: 3865: 3860: 3859: 3855: 3845: 3843: 3834: 3833: 3829: 3819: 3817: 3808: 3807: 3803: 3789: 3785: 3775: 3773: 3764: 3763: 3759: 3749: 3747: 3738: 3737: 3733: 3728: 3724: 3711: 3710: 3706: 3696: 3694: 3680: 3679: 3675: 3653: 3652: 3648: 3637: 3633: 3620: 3616: 3603: 3599: 3577: 3576: 3569: 3557: 3551: 3550: 3546: 3524: 3523: 3516: 3506:Plant Pathology 3503: 3499: 3492: 3479: 3478: 3474: 3456: 3455: 3451: 3433: 3432: 3428: 3410: 3409: 3405: 3387: 3386: 3375: 3335: 3334: 3330: 3321: 3312: 3304:: Stylophyllum 3300: 3296: 3283: 3281: 3272: 3271: 3267: 3258: 3254: 3237: 3233: 3201: 3196: 3195: 3191: 3182: 3178: 3169: 3162: 3143: 3138: 3137: 3130: 3120: 3119: 3115: 3095: 3094: 3090: 3081:Dudleya anomala 3079: 3075: 3062: 3058: 3051: 3047: 3037: 3035: 3031: 3020: 3015: 3014: 3010: 3000: 2998: 2989: 2988: 2984: 2971: 2964: 2957: 2941: 2940: 2905: 2895: 2893: 2883: 2882: 2865: 2856: 2837: 2824: 2803: 2780:(45): 892–926. 2767: 2766: 2762: 2757: 2753: 2748: 2701: 2663:Channel Islands 2548:Dudleya verityi 2490: 2442: 2437: 2354: 2334: 2297: 2280: 2235: 2213:Dudleya candida 2194: 2189: 2168:Baja California 2162:is centered in 2150:, and southern 2136:The Californias 2108: 2101: 2093: 2089: 2080: 2072: 2068: 2059: 2047: 2043: 2034: 2026: 2022: 2013: 2005: 2001: 1992: 1984: 1980: 1951: 1888: 1883: 1827: 1796:, now known as 1767: 1762: 1756:Dudleya viscida 1749:Dudleya viridis 1725:Dudleya verityi 1599:Dudleya palmeri 1519:Dudleya greenei 1503:Dudleya gatesii 1495:Dudleya formosa 1433:Dudleya cymosa. 1334:Dudleya candida 1246:Dudleya anomala 1143: 1095:'s relation to 1025:paleontologists 975:guidelines for 961:Paul H. Thomson 957: 931:had split from 846: 723:described both 713:Sedum cotyledon 694: 689: 684: 626: 536:Dudleya edulis, 446: 441: 216: 190: 166:Sempervivoideae 82: 74: 73: 72: 71: 70: 69: 60: 59: 58: 51: 42: 41: 40: 17: 12: 11: 5: 4597: 4595: 4587: 4586: 4581: 4576: 4571: 4566: 4556: 4555: 4549: 4548: 4546: 4545: 4541:wfo-4000037012 4532: 4519: 4506: 4493: 4480: 4467: 4454: 4441: 4425: 4423: 4417: 4416: 4414: 4413: 4409:wfo-4000017003 4400: 4387: 4374: 4361: 4348: 4335: 4322: 4306: 4304: 4298: 4297: 4295: 4294: 4290:wfo-4000012706 4281: 4268: 4255: 4242: 4229: 4216: 4203: 4190: 4177: 4164: 4151: 4138: 4125: 4112: 4099: 4086: 4073: 4060: 4045: 4029: 4027: 4021: 4020: 4015: 3997: 3996: 3991: 3968: 3965: 3962: 3961: 3935: 3909: 3879: 3853: 3827: 3801: 3783: 3757: 3731: 3722: 3719:. 28 May 2017. 3704: 3673: 3646: 3631: 3614: 3597: 3592:10.7280/D1NH4W 3567: 3544: 3514: 3497: 3490: 3472: 3449: 3426: 3403: 3373: 3352:(4): 681–689. 3328: 3310: 3294: 3265: 3252: 3250:59(4) 223–229. 3231: 3189: 3176: 3160: 3128: 3113: 3088: 3073: 3065:Phytochemistry 3056: 3045: 3008: 2982: 2962: 2956:978-0960206650 2955: 2903: 2863: 2835: 2801: 2760: 2750: 2749: 2747: 2744: 2743: 2742: 2732: 2726: 2714: 2700: 2697: 2679:proposed Bill 2591:Sinaloa Cartel 2583:Bahía Tortugas 2579:D. pachyphytum 2489: 2486: 2478:inflorescences 2476:. The budding 2441: 2438: 2436: 2433: 2432: 2431: 2421: 2404: 2400: 2381: 2378: 2371: 2364: 2353: 2350: 2333: 2330: 2296: 2293: 2279: 2276: 2268:Powdery mildew 2234: 2233:Growing medium 2231: 2216:in cultivation 2193: 2190: 2188: 2185: 2130:Regions where 2107: 2104: 2103: 2102: 2090: 2083: 2081: 2069: 2062: 2060: 2053:Dudleya virens 2044: 2037: 2035: 2032:Dudleya edulis 2023: 2016: 2014: 2002: 1995: 1993: 1981: 1974: 1950: 1947: 1908:hermaphroditic 1887: 1884: 1882: 1879: 1878: 1877: 1866: 1850: 1839: 1826: 1823: 1814:Dudleya virens 1798:Dudleya edulis 1766: 1763: 1761: 1760: 1752: 1745: 1737: 1733:Dudleya virens 1729: 1721: 1713: 1705: 1697: 1696: 1695: 1687:Dudleya saxosa 1683: 1675:Dudleya saxosa 1671: 1663:Dudleya saxosa 1655:Dudleya saxosa 1651: 1647:Dudleya rubens 1643: 1635: 1631:Dudleya rigida 1627: 1619: 1611: 1603: 1595: 1587: 1579: 1571: 1563: 1555: 1547: 1539: 1535:Dudleya ingens 1531: 1523: 1515: 1507: 1499: 1491: 1483: 1479:Dudleya edulis 1475: 1467: 1466: 1465: 1457:Dudleya cymosa 1453: 1445:Dudleya cymosa 1441: 1429: 1421:Dudleya cymosa 1417: 1409:Dudleya cymosa 1405: 1397:Dudleya cymosa 1393: 1385:Dudleya cymosa 1381: 1373:Dudleya cymosa 1365:Dudleya cymosa 1361: 1353: 1345: 1338: 1330: 1322: 1314: 1306: 1298: 1295:D. blochmaniae 1286: 1278: 1266: 1258: 1250: 1242: 1234: 1226: 1225: 1224: 1212: 1200: 1188: 1176: 1164: 1144: 1142: 1139: 1060:. This places 981:type specimens 956: 953: 845: 842: 784:In the 1930s, 721:Thomas Nuttall 693: 690: 688: 685: 683: 680: 625: 622: 619: 618: 571: 524: 500: 499: 492: 485: 477: 476: 445: 442: 440: 437: 251: 250: 239: 238: 230: 229: 228:About 68 taxa 225: 224: 218: 217: 208: 207: 201: 200: 183: 179: 178: 173: 169: 168: 163: 159: 158: 153: 149: 148: 143: 139: 138: 133: 126: 125: 120: 113: 112: 107: 100: 99: 94: 90: 89: 76: 75: 63: 62: 61: 52: 45: 44: 43: 34: 33: 32: 31: 30: 29: 26: 25: 15: 13: 10: 9: 6: 4: 3: 2: 4596: 4585: 4582: 4580: 4577: 4575: 4572: 4570: 4567: 4565: 4562: 4561: 4559: 4542: 4537: 4533: 4529: 4524: 4520: 4516: 4511: 4507: 4503: 4498: 4494: 4490: 4485: 4481: 4477: 4472: 4468: 4464: 4459: 4455: 4451: 4446: 4442: 4437: 4431: 4427: 4426: 4424: 4422: 4418: 4410: 4405: 4401: 4397: 4392: 4388: 4384: 4379: 4375: 4371: 4366: 4362: 4358: 4353: 4349: 4345: 4340: 4336: 4332: 4327: 4323: 4318: 4312: 4308: 4307: 4305: 4303: 4299: 4291: 4286: 4282: 4278: 4273: 4269: 4265: 4260: 4256: 4252: 4247: 4243: 4239: 4234: 4230: 4226: 4221: 4217: 4213: 4208: 4204: 4200: 4195: 4191: 4187: 4182: 4178: 4174: 4169: 4165: 4161: 4156: 4152: 4148: 4143: 4139: 4135: 4130: 4126: 4122: 4117: 4113: 4109: 4104: 4100: 4096: 4091: 4087: 4083: 4078: 4074: 4070: 4065: 4061: 4056: 4050: 4046: 4041: 4035: 4031: 4030: 4028: 4026: 4022: 4018: 4013: 4009: 4007: 4002: 3995: 3992: 3990: 3989:9780960206650 3986: 3983: 3982:0-9602066-5-5 3979: 3975: 3972:Thompson, P. 3971: 3970: 3966: 3950: 3946: 3939: 3936: 3924: 3920: 3913: 3910: 3906: 3901: 3897: 3890: 3883: 3880: 3876: 3871: 3864: 3857: 3854: 3842: 3838: 3831: 3828: 3816: 3812: 3805: 3802: 3799: 3796: 3793: 3787: 3784: 3771: 3767: 3761: 3758: 3745: 3741: 3735: 3732: 3726: 3723: 3718: 3714: 3708: 3705: 3692: 3688: 3684: 3677: 3674: 3669: 3665: 3661: 3657: 3650: 3647: 3643: 3642: 3635: 3632: 3628: 3624: 3618: 3615: 3611: 3607: 3601: 3598: 3593: 3589: 3585: 3581: 3574: 3572: 3568: 3564: 3556: 3555: 3548: 3545: 3540: 3536: 3532: 3528: 3521: 3519: 3515: 3511: 3507: 3501: 3498: 3493: 3491:0-9628505-8-6 3487: 3483: 3476: 3473: 3468: 3464: 3460: 3453: 3450: 3445: 3441: 3437: 3430: 3427: 3422: 3418: 3414: 3407: 3404: 3399: 3395: 3391: 3384: 3382: 3380: 3378: 3374: 3368: 3363: 3359: 3355: 3351: 3347: 3343: 3341: 3332: 3329: 3325: 3319: 3317: 3315: 3311: 3307: 3303: 3298: 3295: 3291: 3280: 3276: 3269: 3266: 3262: 3256: 3253: 3249: 3245: 3243: 3235: 3232: 3227: 3223: 3219: 3215: 3211: 3207: 3200: 3193: 3190: 3186: 3180: 3177: 3173: 3167: 3165: 3161: 3157: 3153: 3149: 3142: 3135: 3133: 3129: 3124: 3117: 3114: 3109: 3105: 3104: 3099: 3092: 3089: 3085: 3082: 3077: 3074: 3070: 3066: 3060: 3057: 3054: 3049: 3046: 3030: 3026: 3019: 3012: 3009: 2997: 2993: 2986: 2983: 2980:(4), 889-900. 2979: 2975: 2969: 2967: 2963: 2958: 2952: 2948: 2944: 2938: 2936: 2934: 2932: 2930: 2928: 2926: 2924: 2922: 2920: 2918: 2916: 2914: 2912: 2910: 2908: 2904: 2891: 2887: 2880: 2878: 2876: 2874: 2872: 2870: 2868: 2864: 2860: 2854: 2852: 2850: 2848: 2846: 2844: 2842: 2840: 2836: 2832: 2828: 2822: 2820: 2818: 2816: 2814: 2812: 2810: 2808: 2806: 2802: 2797: 2793: 2788: 2783: 2779: 2775: 2771: 2764: 2761: 2755: 2752: 2745: 2740: 2736: 2733: 2730: 2727: 2724: 2720: 2719: 2715: 2712: 2708: 2707: 2703: 2702: 2698: 2696: 2694: 2690: 2686: 2682: 2678: 2674: 2671: 2666: 2664: 2660: 2656: 2651: 2649: 2645: 2641: 2637: 2633: 2629: 2625: 2622: 2618: 2614: 2610: 2606: 2605:Bluff lettuce 2602: 2600: 2596: 2592: 2588: 2584: 2580: 2576: 2572: 2571:Cedros Island 2568: 2564: 2560: 2555: 2553: 2549: 2545: 2537: 2536: 2531: 2527: 2525: 2521: 2518: 2515: 2511: 2507: 2503: 2499: 2495: 2487: 2485: 2483: 2479: 2475: 2471: 2467: 2463: 2459: 2455: 2451: 2447: 2439: 2434: 2429: 2425: 2422: 2419: 2415: 2414: 2413:Dudleya parva 2409: 2405: 2401: 2398: 2394: 2390: 2386: 2382: 2379: 2376: 2372: 2369: 2368:axillary buds 2365: 2362: 2361: 2360: 2358: 2351: 2349: 2347: 2343: 2338: 2331: 2326: 2325: 2320: 2316: 2314: 2310: 2306: 2302: 2294: 2292: 2290: 2285: 2277: 2275: 2273: 2269: 2265: 2261: 2257: 2253: 2250: 2247: 2245: 2240: 2232: 2230: 2228: 2222: 2215: 2214: 2209: 2205: 2203: 2199: 2191: 2186: 2181: 2180: 2175: 2171: 2169: 2166:and northern 2165: 2161: 2157: 2153: 2149: 2145: 2141: 2137: 2133: 2125: 2124: 2123:Dudleya gnoma 2119: 2115: 2112: 2105: 2100: 2096: 2087: 2082: 2079: 2075: 2066: 2061: 2058: 2054: 2050: 2041: 2036: 2033: 2029: 2020: 2015: 2012: 2008: 1999: 1994: 1991: 1987: 1978: 1973: 1971: 1969: 1965: 1960: 1956: 1948: 1946: 1944: 1940: 1936: 1932: 1928: 1924: 1920: 1915: 1913: 1909: 1905: 1901: 1897: 1893: 1885: 1880: 1876: 1872: 1871: 1867: 1865:(narrow-leaf) 1864: 1861:(wide-leaf); 1860: 1856: 1855: 1851: 1849: 1845: 1844: 1840: 1838: 1834: 1833: 1829: 1828: 1824: 1822: 1820: 1819: 1815: 1810: 1806: 1801: 1799: 1795: 1791: 1787: 1782: 1780: 1776: 1772: 1764: 1758: 1757: 1753: 1751: 1750: 1746: 1743: 1742: 1738: 1735: 1734: 1730: 1727: 1726: 1722: 1719: 1718: 1714: 1711: 1710: 1706: 1703: 1702: 1698: 1693: 1692: 1688: 1684: 1681: 1680: 1676: 1672: 1669: 1668: 1664: 1660: 1659: 1657: 1656: 1652: 1649: 1648: 1644: 1641: 1640: 1636: 1633: 1632: 1628: 1625: 1624: 1620: 1617: 1616: 1612: 1609: 1608: 1607:Dudleya parva 1604: 1601: 1600: 1596: 1593: 1592: 1588: 1585: 1584: 1580: 1577: 1576: 1572: 1569: 1568: 1564: 1561: 1560: 1556: 1553: 1552: 1548: 1545: 1544: 1540: 1537: 1536: 1532: 1529: 1528: 1524: 1521: 1520: 1516: 1513: 1512: 1511:Dudleya gnoma 1508: 1505: 1504: 1500: 1497: 1496: 1492: 1489: 1488: 1484: 1481: 1480: 1476: 1473: 1472: 1468: 1463: 1462: 1458: 1454: 1451: 1450: 1446: 1442: 1439: 1438: 1434: 1430: 1427: 1426: 1422: 1418: 1415: 1414: 1410: 1406: 1403: 1402: 1398: 1394: 1391: 1390: 1386: 1382: 1379: 1378: 1374: 1370: 1369: 1367: 1366: 1362: 1359: 1358: 1354: 1351: 1350: 1346: 1344: 1343: 1339: 1336: 1335: 1331: 1328: 1327: 1323: 1320: 1319: 1315: 1312: 1311: 1307: 1304: 1303: 1299: 1296: 1292: 1291: 1287: 1284: 1283: 1279: 1276: 1272: 1271: 1267: 1264: 1263: 1259: 1256: 1255: 1251: 1248: 1247: 1243: 1240: 1239: 1235: 1232: 1231: 1227: 1222: 1221: 1217: 1213: 1210: 1209: 1205: 1201: 1198: 1197: 1193: 1189: 1186: 1185: 1181: 1177: 1174: 1173: 1169: 1165: 1162: 1161: 1157: 1153: 1152: 1151: 1150: 1146: 1145: 1140: 1138: 1136: 1132: 1128: 1124: 1120: 1116: 1112: 1111: 1106: 1102: 1098: 1094: 1089: 1087: 1083: 1079: 1075: 1071: 1067: 1063: 1059: 1058: 1057:Sedum moranii 1053: 1049: 1045: 1041: 1037: 1032: 1030: 1026: 1022: 1018: 1014: 1010: 1006: 1002: 998: 994: 990: 986: 982: 978: 974: 970: 966: 962: 954: 952: 950: 946: 942: 938: 934: 930: 926: 922: 918: 914: 910: 906: 902: 897: 895: 891: 887: 883: 879: 875: 871: 867: 863: 860:, and merged 859: 855: 851: 843: 841: 839: 835: 831: 827: 823: 819: 815: 811: 810: 805: 801: 800: 795: 791: 787: 782: 780: 776: 772: 768: 764: 760: 756: 752: 748: 744: 742: 741: 736: 732: 731: 726: 722: 718: 714: 710: 708: 703: 699: 692:Early history 691: 686: 681: 679: 677: 676: 671: 670: 664: 662: 661:triterpenoids 658: 654: 650: 649:chalk dudleya 646: 642: 634: 630: 623: 616: 612: 608: 604: 603: 598: 594: 593:plesiomorphic 590: 589: 584: 580: 576: 572: 569: 565: 561: 560: 555: 554:Stylophyllum, 551: 547: 546: 541: 537: 533: 529: 525: 522: 518: 517:chalk dudleya 514: 510: 506: 503:The subgenus 502: 501: 498: 497: 493: 491: 490: 486: 484: 483: 479: 478: 473: 468: 466: 462: 457: 455: 451: 443: 438: 436: 434: 430: 426: 422: 418: 415:, especially 414: 410: 406: 402: 398: 394: 390: 386: 382: 380: 376: 372: 368: 364: 360: 356: 352: 351: 346: 345: 340: 336: 332: 331: 326: 321: 319: 315: 311: 307: 303: 299: 295: 291: 290:North America 287: 283: 279: 275: 271: 267: 263: 259: 258: 249: 248:North America 245: 240: 236: 231: 226: 223: 219: 215: 214: 209: 206: 202: 198: 194: 189: 188: 184: 181: 180: 177: 174: 171: 170: 167: 164: 161: 160: 157: 154: 151: 150: 147: 144: 141: 140: 137: 134: 131: 128: 127: 124: 121: 118: 115: 114: 111: 110:Tracheophytes 108: 105: 102: 101: 98: 95: 92: 91: 86: 81: 77: 67: 56: 49: 38: 27: 24:Liveforevers 22: 19: 4421:Stylophyllum 4420: 4301: 4024: 3998: 3973: 3952:. Retrieved 3948: 3938: 3926:. Retrieved 3922: 3912: 3903: 3899: 3895: 3882: 3873: 3869: 3856: 3844:. Retrieved 3840: 3830: 3818:. Retrieved 3814: 3804: 3794: 3786: 3774:. Retrieved 3769: 3760: 3748:. Retrieved 3743: 3734: 3725: 3716: 3707: 3695:. Retrieved 3687:Ensenada.net 3686: 3676: 3659: 3649: 3639: 3634: 3626: 3622: 3617: 3605: 3600: 3583: 3561: 3553: 3547: 3530: 3509: 3505: 3500: 3481: 3475: 3462: 3452: 3439: 3429: 3416: 3406: 3393: 3349: 3345: 3339: 3331: 3323: 3305: 3301: 3297: 3289: 3282:. Retrieved 3278: 3268: 3260: 3255: 3247: 3241: 3234: 3209: 3205: 3192: 3184: 3179: 3171: 3155: 3151: 3147: 3122: 3116: 3101: 3091: 3083: 3080: 3076: 3068: 3064: 3059: 3048: 3036:. Retrieved 3024: 3011: 2999:. Retrieved 2995: 2985: 2977: 2973: 2946: 2894:. Retrieved 2889: 2858: 2830: 2826: 2777: 2773: 2763: 2754: 2722: 2716: 2710: 2704: 2688: 2669: 2667: 2658: 2652: 2639: 2636:D. farinosa. 2635: 2628:South Korean 2623: 2620: 2608: 2603: 2595:Mexican Navy 2586: 2578: 2566: 2562: 2556: 2552:Springs Fire 2547: 2541: 2533: 2523: 2505: 2493: 2491: 2488:Conservation 2458:agricultural 2443: 2427: 2417: 2411: 2407: 2396: 2388: 2355: 2336: 2335: 2322: 2304: 2300: 2298: 2288: 2283: 2281: 2271: 2263: 2254: 2251: 2248: 2243: 2238: 2236: 2226: 2223: 2219: 2211: 2201: 2197: 2195: 2187:Horticulture 2177: 2159: 2131: 2129: 2121: 2110: 2109: 2098: 2092: 2077: 2071: 2056: 2052: 2048:Stylophyllum 2046: 2031: 2027:Stylophyllum 2025: 2010: 2004: 1989: 1983: 1967: 1963: 1958: 1954: 1952: 1942: 1939:Stylophyllum 1938: 1934: 1930: 1927:Stylophyllum 1926: 1922: 1918: 1916: 1911: 1896:hummingbirds 1891: 1889: 1874: 1869: 1863:milh kajmila 1862: 1858: 1853: 1847: 1841: 1836: 1830: 1825:Nomenclature 1817: 1813: 1804: 1802: 1797: 1793: 1786:Stylophyllum 1785: 1783: 1778: 1768: 1754: 1747: 1739: 1731: 1723: 1715: 1707: 1699: 1690: 1686: 1678: 1674: 1666: 1662: 1653: 1645: 1637: 1629: 1621: 1613: 1605: 1597: 1589: 1581: 1573: 1565: 1557: 1549: 1541: 1533: 1525: 1517: 1509: 1501: 1493: 1485: 1477: 1469: 1460: 1456: 1448: 1444: 1436: 1432: 1424: 1420: 1412: 1408: 1401:costatifolia 1400: 1396: 1388: 1384: 1376: 1372: 1363: 1355: 1347: 1340: 1332: 1324: 1316: 1308: 1300: 1294: 1288: 1280: 1274: 1268: 1260: 1252: 1244: 1236: 1228: 1219: 1215: 1207: 1203: 1195: 1191: 1183: 1179: 1171: 1167: 1159: 1155: 1147: 1134: 1130: 1126: 1122: 1118: 1114: 1108: 1100: 1096: 1092: 1090: 1085: 1081: 1077: 1073: 1069: 1061: 1055: 1051: 1047: 1043: 1039: 1035: 1033: 1020: 1016: 968: 964: 958: 949:paleoendemic 944: 940: 936: 932: 928: 924: 920: 916: 912: 908: 904: 900: 898: 893: 889: 885: 881: 877: 873: 869: 865: 862:Stylophyllum 861: 857: 853: 847: 837: 833: 830:Stylophyllum 829: 825: 821: 817: 813: 807: 806:genera into 803: 797: 794:Stylophyllum 793: 789: 786:Alwin Berger 783: 778: 775:Stylophyllum 774: 770: 766: 763:Stylophyllum 762: 758: 755:Crassulaceae 745: 738: 734: 728: 724: 712: 705: 697: 695: 673: 667: 665: 640: 638: 632: 606: 600: 596: 586: 575:Hasseanthus, 574: 567: 564:Stylophyllum 563: 557: 553: 549: 543: 539: 535: 528:Stylophyllum 527: 508: 504: 495: 494: 489:Stylophyllum 488: 487: 481: 480: 464: 460: 458: 447: 432: 396: 389:native plant 384: 383: 375:Phylogenetic 366: 354: 348: 342: 334: 328: 322: 309: 286:Crassulaceae 270:siemprevivas 269: 262:liveforevers 261: 256: 255: 254: 243: 211: 205:Type species 186: 185: 156:Crassulaceae 146:Saxifragales 129: 116: 103: 18: 4302:Hasseanthus 4168:iNaturalist 4049:Wikispecies 3308:, gen. nov. 3187:. Backhuys. 2569:endemic to 2508:species is 2435:Ethnobotany 2408:Hasseanthus 2346:vermiculite 2313:germination 2295:Propagation 2227:Hasseanthus 2094:Hasseanthus 2073:Hasseanthus 1968:Hasseanthus 1964:Hasseanthus 1935:Hasseanthus 1923:Hasseanthus 1894:are mostly 1805:Hasseanthus 1779:liveforever 1777:. The term 1389:crebrifolia 1131:Hasseanthus 1123:Hasseanthus 1054:clade like 1017:Hasseanthus 1001:cosmic rays 901:Hasseanthus 886:Hasseanthus 874:Hasseanthus 838:Hasseanthus 822:Hasseanthus 804:Hasseanthus 779:Hasseanthus 767:Hasseanthus 607:Hasseanthus 591:, may be a 496:Hasseanthus 465:Hasseanthus 439:Description 429:governments 417:South Korea 325:neoendemics 318:bell-shaped 162:Subfamily: 123:Angiosperms 4558:Categories 4476:30012235-2 3954:25 October 3923:CalMatters 3284:18 October 2861:, 492-502. 2833:, 224-231. 2746:References 2729:Reid Moran 2677:Chris Ward 2520:collectors 2514:East Asian 2482:astringent 2142:, coastal 1859:milhka’mey 1449:paniculata 1437:ovatifolia 1425:marcescens 1377:agourensis 1220:setchellii 1066:neoendemic 1052:Leucosedum 977:describing 850:Reid Moran 675:D. anomala 669:D. viscida 532:fingertips 509:Eududleya, 475:Subgenera 379:endangered 371:Reid Moran 4436:Q15733393 4317:Q87176181 3928:16 August 3905:overseas. 3846:16 August 3820:16 August 3776:21 August 3750:21 August 3697:2 January 3436:"Dudleya" 3038:5 October 3001:5 October 2796:225261669 2718:Echeveria 2644:aesthetic 2517:succulent 2462:succulent 2454:medicinal 2301:Echeveria 2256:Mealybugs 1970:subgenus. 1957:. Larger 1875:awi mielh 1765:Etymology 1667:collomiae 1196:calcicola 1105:Sedoideae 1082:Echeveria 1078:Echeveria 1048:Echeveria 1040:Echeveria 1013:evolution 941:Echeveria 933:Echeveria 921:Echeveria 909:Echeveria 890:Echeveria 866:Eududleya 858:Echeveria 848:In 1942, 818:Echeveria 799:Echeveria 583:disappear 513:connation 507:, or the 425:nurseries 413:East Asia 350:Cotyledon 344:Echeveria 306:deciduous 302:geophytic 300:, others 298:evergreen 282:succulent 280:-forming 222:Diversity 93:Kingdom: 4528:40007654 4523:Tropicos 4430:Wikidata 4396:40021598 4391:Tropicos 4357:298178-2 4311:Wikidata 4277:40036253 4272:Tropicos 4186:297170-2 4034:Wikidata 3691:Archived 3668:Archived 3539:Archived 3531:CNPS-SLO 3467:Archived 3444:Archived 3421:Archived 3398:Archived 3226:15715233 3108:Archived 3029:Archived 2945:(1993). 2699:See also 2670:Dudleya, 2648:caudices 2624:farinosa 2526:species. 2510:poaching 2470:calluses 2446:Kumeyaay 2375:wildfire 2373:After a 2192:Watering 1790:allusion 1184:bettinae 1160:abramsii 989:Atlantis 969:Dudleya, 682:Taxonomy 405:poachers 152:Family: 136:Eudicots 4564:Dudleya 4489:1301294 4450:7333965 4370:1064817 4331:7334672 4199:1039287 4147:2985872 4055:Dudleya 4040:Q138291 4025:Dudleya 4006:Dudleya 3875:County. 3563:2008a). 3354:Bibcode 3340:Dudleya 3261:Madroño 3248:Madroño 3103:YouTube 2723:Dudleya 2711:Dudleya 2689:Dudleya 2675:member 2640:Dudleya 2632:Chinese 2597:in the 2587:Dudleya 2567:Dudleya 2524:Dudleya 2506:Dudleya 2494:Dudleya 2428:Dudleya 2418:Dudleya 2389:Dudleya 2387:. Many 2337:Dudleya 2309:offsets 2305:Dudleya 2303:, many 2289:Dudleya 2284:Dudleya 2272:Dudleya 2264:Dudleya 2244:Dudleya 2239:Dudleya 2202:Dudleya 2198:Dudleya 2160:Dudleya 2140:Arizona 2132:Dudleya 2111:Dudleya 2055:subsp. 2006:Dudleya 1985:Dudleya 1959:Dudleya 1955:Dudleya 1943:Dudleya 1931:Dudleya 1919:Dudleya 1912:Dudleya 1892:Dudleya 1843:Spanish 1832:English 1816:subsp. 1689:subsp. 1679:aloides 1677:subsp. 1665:subsp. 1459:subsp. 1447:subsp. 1435:subsp. 1423:subsp. 1411:subsp. 1399:subsp. 1387:subsp. 1375:subsp. 1218:subsp. 1206:subsp. 1194:subsp. 1182:subsp. 1172:affinis 1170:subsp. 1158:subsp. 1135:Dudleya 1119:Dudleya 1115:Dudleya 1101:Dudleya 1093:Dudleya 1086:Dudleya 1070:Dudleya 1062:Dudleya 1044:Dudleya 1036:Dudleya 1021:Dudleya 993:Lemuria 945:Dudleya 937:Dudleya 929:Dudleya 925:Dudleya 917:Dudleya 905:Dudleya 882:Dudleya 878:Dudleya 870:Dudleya 854:Dudleya 834:Dudleya 814:Dudleya 790:Dudleya 771:Dudleya 759:Dudleya 641:Dudleya 568:Dudleya 550:Dudleya 540:Dudleya 505:Dudleya 482:Dudleya 433:Dudleya 431:combat 397:Dudleya 385:Dudleya 367:Dudleya 314:tubular 310:Dudleya 278:rosette 272:) is a 266:Spanish 257:Dudleya 246:within 244:Dudleya 193:Britton 187:Dudleya 182:Genus: 172:Tribe: 142:Order: 97:Plantae 4246:PLANTS 4238:437858 4212:195811 4134:111029 4121:111029 4069:423052 3987:  3980:  3841:SFGATE 3629:, 367. 3488:  3224:  2953:  2896:8 June 2794:  2681:AB-223 2466:thirst 2450:Paipai 2342:pumice 2260:aphids 2156:Nevada 2148:Oregon 2144:Sonora 2091:subg. 2070:subg. 2057:hassei 2045:subg. 2024:subg. 2003:subg. 1982:subg. 1929:, and 1870:Paipai 1854:Tiipai 1818:hassei 1788:is an 1691:saxosa 1461:pumila 1413:cymosa 1208:murina 733:) and 655:, and 613:, and 463:, and 450:Sedeae 195:& 176:Sedeae 4484:IRMNG 4463:16658 4365:IRMNG 4344:17298 4251:DUDLE 4225:23003 4194:IRMNG 4173:51891 4108:1DUDG 4095:38024 3896:Flora 3892:(PDF) 3866:(PDF) 3558:(PDF) 3222:S2CID 3202:(PDF) 3144:(PDF) 3032:(PDF) 3021:(PDF) 2792:S2CID 2774:Taxon 2706:Sedum 2474:corns 2278:Shade 1949:Seeds 1127:Sedum 1097:Sedum 1074:Sedum 1064:as a 919:with 913:Sedum 894:Sedum 826:Sedum 809:Sedum 700:, by 588:Sedum 556:like 409:niche 355:Sedum 335:Sedum 330:Sedum 274:genus 130:Clade 117:Clade 104:Clade 4510:POWO 4502:6903 4471:IPNI 4458:GRIN 4445:GBIF 4378:POWO 4352:IPNI 4339:GRIN 4326:GBIF 4259:POWO 4220:NCBI 4207:ITIS 4181:IPNI 4160:4026 4155:GRIN 4142:GBIF 4103:EPPO 4082:48MW 4064:BOLD 3985:ISBN 3978:ISBN 3956:2021 3930:2021 3848:2021 3822:2021 3778:2021 3752:2021 3699:2022 3512:(4). 3486:ISBN 3286:2021 3040:2021 3003:2021 2951:ISBN 2898:2020 2737:and 2653:The 2630:and 2557:The 2472:and 2456:and 2448:and 2258:and 2154:and 2152:Utah 2146:and 1937:and 1900:bees 1898:and 1125:and 1084:and 1072:and 1038:and 1019:and 1011:and 1003:and 995:and 911:and 884:and 836:and 792:and 773:and 749:and 672:and 579:corm 552:and 519:and 427:and 403:and 361:and 347:and 304:and 292:and 197:Rose 4536:WFO 4404:WFO 4285:WFO 4129:FoC 4116:FNA 4090:EoL 4077:CoL 3588:doi 3362:doi 3214:doi 2782:doi 2344:or 1846:: s 1015:of 963:'s 892:or 715:by 276:of 4560:: 4538:: 4525:: 4512:: 4499:: 4486:: 4473:: 4460:: 4447:: 4432:: 4406:: 4393:: 4380:: 4367:: 4354:: 4341:: 4328:: 4313:: 4287:: 4274:: 4261:: 4248:: 4235:: 4222:: 4209:: 4196:: 4183:: 4170:: 4157:: 4144:: 4131:: 4118:: 4105:: 4092:: 4079:: 4066:: 4051:: 4036:: 3947:. 3921:. 3898:. 3894:. 3868:. 3839:. 3813:. 3768:. 3742:. 3715:. 3689:. 3685:. 3666:. 3662:. 3658:. 3625:, 3608:. 3586:. 3582:. 3570:^ 3537:. 3533:. 3529:. 3517:^ 3510:57 3508:, 3465:. 3461:. 3442:. 3438:. 3419:. 3415:. 3396:. 3392:. 3376:^ 3360:. 3350:19 3348:. 3344:. 3313:^ 3288:. 3277:. 3246:. 3220:. 3210:38 3208:. 3204:. 3163:^ 3154:. 3152:45 3150:. 3146:. 3131:^ 3106:. 3100:. 3069:17 3067:, 3027:. 3023:. 2994:. 2978:43 2976:, 2965:^ 2906:^ 2888:. 2866:^ 2838:^ 2831:76 2829:, 2804:^ 2790:. 2778:69 2776:. 2772:. 2621:D. 2311:, 2274:. 2266:. 2170:. 2138:, 1925:, 1873:: 1857:: 1835:: 1800:. 1088:. 1031:. 997:Mu 991:, 951:. 947:a 896:. 663:. 651:, 523:. 467:. 456:. 423:, 268:: 132:: 119:: 106:: 3958:. 3932:. 3900:4 3850:. 3824:. 3780:. 3754:. 3701:. 3627:8 3612:. 3590:: 3494:. 3370:. 3364:: 3356:: 3263:. 3240:" 3228:. 3216:: 3125:. 3042:. 3005:. 2959:. 2900:. 2784:: 2725:. 2713:. 2657:( 2607:( 2561:( 2546:( 2395:( 1297:. 1277:. 737:( 727:( 709:. 534:( 264:(

Index

D. virens ssp. insularis
D. nesiotica
D. arizonica
D. farinosa
Scientific classification
Edit this classification
Plantae
Tracheophytes
Angiosperms
Eudicots
Saxifragales
Crassulaceae
Sempervivoideae
Sedeae
Dudleya
Britton
Rose
Type species
Dudleya lanceolata
Diversity

North America
Spanish
genus
rosette
succulent
Crassulaceae
North America
Guadalupe Island
evergreen

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