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Darwin's finches

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continent. There are twenty-six land birds, and twenty-five of these are ranked by Mr. Gould as distinct species, supposed to have been created here; yet the close affinity of most of these birds to American species in every character, in their habits, gestures, and tones of voice, was manifest. ... The naturalist, looking at the inhabitants of these volcanic islands in the Pacific, distant several hundred miles from the continent, yet feels that he is standing on American land. Why should this be so? Why should the species which are supposed to have been created in the Galapagos Archipelago, and nowhere else, bear so plain a stamp of affinity to those created in America? There is nothing in the conditions of life, in the geological nature of the islands, in their height or climate, or in the proportions in which the several classes are associated together, which resembles closely the conditions of the South American coast: In fact there is a considerable dissimilarity in all these respects. On the other hand, there is a considerable degree of resemblance in the volcanic nature of the soil, in climate, height, and size of the islands, between the Galapagos and Cape de Verde Archipelagos: But what an entire and absolute difference in their inhabitants! The inhabitants of the Cape de Verde Islands are related to those of Africa, like those of the Galapagos to America. I believe this grand fact can receive no sort of explanation on the ordinary view of independent creation; whereas on the view here maintained, it is obvious that the Galapagos Islands would be likely to receive colonists, whether by occasional means of transport or by formerly continuous land, from America; and the Cape de Verde Islands from Africa; and that such colonists would be liable to modification — the principle of inheritance still betraying their original birthplace.
859:) has produced a new "big bird" population that can exploit previously unexploited food due to its larger size. They do not breed with the other species on the island, as the females do not recognize the songs of the new males. Genetic evidence shows that currently, after several generations (a time scale that suggests shorter speciation events could have occurred previously), it lives in a complete reproductive isolation from the native species. According to professor Leif Andersson of Uppsala University, a taxonomist not aware of its history would consider it a distinct species. 42: 558:, the two species may be often seen climbing about the flowers of the great cactus-trees; but all the other species of this group of finches, mingled together in flocks, feed on the dry and sterile ground of the lower districts. The males of all, or certainly of the greater number, are jet black; and the females (with perhaps one or two exceptions) are brown. The most curious fact is the perfect gradation in the size of the beaks in the different species of 74: 395: 4309: 4333: 723: 4345: 4321: 1019:(BMP4), and its differential expression during development, resulted in variation of beak size and shape among finches. BMP4 acts in the developing embryo to lay down skeletal features, including making the beak stronger. The same group showed that the development of the different beak shapes in Darwin's finches are also influenced by slightly 285:. Lack based his analysis on the large collection of museum specimens collected by the 1905–06 Galápagos expedition of the California Academy of Sciences, to whom Lack dedicated his 1947 book. The birds vary in size from 10 to 20 cm (4 to 8 in) and weigh between 8 and 38 grams (0.3 and 1.3 oz). The smallest are the 2691: 583:, is slightly parrot-shaped. Seeing this gradation and diversity of structure in one small, intimately related group of birds, one might really fancy that from an original paucity of birds in this archipelago, one species had been taken and modified for different ends. In a like manner it might be fancied that 549:
The remaining land-birds form a most singular group of finches, related to each other in the structure of their beaks, short tails, form of body and plumage: There are thirteen species, which Mr. Gould has divided into four subgroups. All these species are peculiar to this archipelago; and so is
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The most striking and important fact for us in regard to the inhabitants of islands, is their affinity to those of the nearest mainland, without being actually the same species. the Galapagos Archipelago ... almost every product of the land and water bears the unmistakable stamp of the American
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acts in a similar way to BMP4, affecting some of the features of beak growth like making them long and pointy. The authors suggest that changes in the temporal and spatial expression of these two factors are possible developmental controls of beak morphology. In a recent study genome sequencing
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pulp, which surrounds the seeds, whereas those with shorter beaks tear apart the cactus base and eat the pulp and any insect larvae and pupae (both groups eat flowers and buds). This dimorphism clearly maximises their feeding opportunities during the non-breeding season when food is scarce.
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that Darwin had labelled by island were separate species rather than just varieties. Gould found more species than Darwin had expected, and concluded that 25 of the 26 land birds were new and distinct forms, found nowhere else in the world but closely allied to those found on the
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continent. Darwin now saw that, if the finch species were confined to individual islands, like the mockingbirds, this would help to account for the number of species on the islands, and he sought information from others on the expedition. Specimens had also been collected by Captain
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Lamichhaney, Sangeet; Berglund, Jonas; Almén, Markus Sällman; Maqbool, Khurram; Grabherr, Manfred; Martinez-Barrio, Alvaro; Promerová, Marta; Rubin, Carl-Johan; Wang, Chao; Zamani, Neda; Grant, B. Rosemary; Grant, Peter R.; Webster, Matthew T.; Andersson, Leif (February 2015).
508:), he described Gould's findings on the number of birds, noting that "Although the species are thus peculiar to the archipelago, yet nearly all in their general structure, habits, colour of feathers, and even tone of voice, are strictly American". In the first edition of 402:) into three other species of finches found on the Galapagos Islands. Due to the absence of other species of birds, the finches adapted to new niches. The finches' beaks and bodies changed allowing them to eat certain types of foods such as nuts, fruits, and insects. 570:
is shown in Fig. 1, and the smallest in Fig. 3; but instead of there being only one intermediate species, with a beak of the size shown in Fig. 2, there are no less than six species with insensibly graduated beaks. The beak of the sub-group
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Narrative of the surveying voyages of His Majesty's Ships Adventure and Beagle between the years 1826 and 1836, describing their examination of the southern shores of South America, and the Beagle's circumnavigation of the globe. Journal and remarks.
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Females are dimorphic in song type: songs A and B are quite distinct. Also, males with song A have shorter bills than B males, another clear difference. With these beaks, males are able to feed differently on their favourite cactus, the prickly pear
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Seeing this gradation and diversity of structure in one small, intimately related group of birds, one might really fancy that from an original paucity of birds in this archipelago, one species had been taken and modified for different
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It is very remarkable that a nearly perfect gradation of structure in this one group can be traced in the form of the beak, from one exceeding in dimensions that of the largest gros-beak, to another differing but little from that of a
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Another interesting dimorphism is for the bills of young finches, which are either 'pink' or 'yellow'. All species of Darwin's finches exhibit this morphism, which lasts for two months. No interpretation of this phenomenon is known.
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Andersson, Leif; Lamichhaney, Sangeet; Berglund, Jonas; AlmĂ©n, Markus Sällman; Maqbool, Khurram; Grabherr, Manfred; et al. (11 February 2015), "Evolution of Darwin's finches and their beaks revealed by genome sequencing",
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gene that encodes a transcription factor affecting craniofacial development is strongly associated with beak shape diversity. Moreover, these changes in the beak size have also altered vocalizations in Darwin's finches.
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Abzhanov, Arhat; Kuo, Winston P.; Hartmann, Christine; Grant, B. Rosemary; Grant, Peter R.; Tabin, Clifford J. (3 August 2006). "The calmodulin pathway and evolution of elongated beak morphology in Darwin's finches".
487:, who had labelled them by island. From these, Darwin tried to reconstruct the locations from where he had collected his own specimens. The conclusions supported his idea of the transmutation of species. 2723:
project, a collaborative initiative for porting open source software to the Darwin platform to enable its use and evolution in the Apple Mac OS X environment. "Fink" is the German name for "finch."
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to different food sources. The birds are all dull-coloured. They are thought to have evolved from a single finch species that came to the islands more than a million years ago.
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Journal of researches into the natural history and geology of the countries visited during the voyage of H.M.S. Beagle round the world, under the Command of Capt. Fitz Roy, R.N
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Abzhanov, Arhat; Protas, Meredith; Grant, B. Rosemary; Grant, Peter R.; Tabin, Clifford J. (3 September 2004). "Bmp4 and Morphological Variation of Beaks in Darwin's Finches".
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Soons, Joris; Herrel, Anthony; Genbrugge, Annelies; Aerts, Peter; Podos, Jeffrey; Adriaens, Dominique; Witte, Yoni de; Jacobs, Patric; Dirckx, Joris (12 April 2010).
5106: 4985: 4812: 4639: 2324: 1657: 1606: 1956: 1412: 5132: 5011: 4838: 4665: 4518: 359:'s teaching, he was interested in the geographical distribution of species, particularly links between species on oceanic islands and on nearby continents. On 706:
checklist, places the Cocos finch in the Emberizidae, but with an asterisk indicating that the placement is probably wrong (AOU 1998–2006); in its tentative
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by expanding its feeding opportunity. The genetics of this situation cannot be clarified in the absence of a detailed breeding program, but two loci with
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he carefully noted where mockingbirds had been caught. In contrast, he paid little attention to the finches. When examining his specimens on the way to
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at that time. In early March, he met Gould again and for the first time to get a full report on the findings, including the point that his Galápagos "
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were actually "a series of ground Finches which are so peculiar an entirely new group, containing 12 species." This story made the newspapers.
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on 4 January 1837, along with other mammal and bird specimens that he had collected. The bird specimens, including the finches, were given to
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and Chatham Islands of a third. As they sailed home about nine months later, this, together with other facts, including what he had heard about
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Darwin discussed the divergence of species of birds in the Galápagos more explicitly in his chapter on geographical distribution in
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Lamichhaney, Sangeet; Han, Fan; Webster, Matthew T.; Andersson, Leif; Grant, B. Rosemary; Grant, Peter R. (12 January 2018).
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and their colleagues have made research trips to the Galápagos for about 30 years, particularly studying Darwin's finches.
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and by this stage of the voyage concentrated mainly on geology. In Galápagos he mostly left bird shooting to his servant
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On the Origin of Species by Means of Natural Selection, or the Preservation of Favoured Races in the Struggle for Life
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birds. They are well known for their remarkable diversity in beak form and function. They are often classified as the
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that shows how modulation of a certain gene during development can account for the differences seen in beak shape.
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and afterward, Darwin thought in terms of "centres of creation" and rejected ideas concerning the
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A long-term study carried out for more than 40 years by the Princeton University researchers
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exhibits a balanced genetic polymorphism and not, as originally supposed, a case of nascent
290: 232: 57: 4578: 2528:"Charles Darwin's bird collection and ornithological knowledge during the voyage of H.M.S. 1168:"Mechanical stress, fracture risk and beak evolution in Darwin's ground finches (Geospiza)" 4203: 4031: 3501: 3023: 3008: 2946: 2845: 2773: 2720: 1271: 1254: 1060: 740: 734: 372: 317: 222: 4911: 2364: 2126: 2058: 1990: 1906: 1865: 1796: 1754: 1108: 4405: 4313: 4243: 4195: 4129: 4068: 4023: 4010: 3884: 3847: 3698: 3472: 3309: 3063: 3058: 3053: 3033: 3018: 2913: 2908: 2888: 2883: 2865: 2813: 2746: 2564: 2450: 2417: 2287: 2254: 2238: 2205: 1192: 1167: 915: 641:. Those with long beaks are able to punch holes in the cactus fruit and eat the fleshy 584: 484: 480: 394: 329: 259: 2512: 1583:
Evolutionary dynamics of a natural population: the large cactus finch of the Galápagos
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Reissued in 1961 by Harper, New York. Reissued in 1983 by Cambridge University Press,
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Monroe and Sibley consider the tanagers to be a tribe (Thraupini) of a big family
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It is not clear whether this placement was made by Burns and Skutch or by Perrins.
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The life and letters of Charles Darwin, including an autobiographical chapter
2142: 2074: 2006: 1812: 1781:"Evolution of Darwin's finches and their beaks revealed by genome sequencing" 4219: 4174: 3963: 3756: 3673: 3657: 3646: 3556: 3492: 3451: 3150: 2951: 2938: 2918: 2818: 1998: 1874: 1849: 929: 462: 337: 239:. The closest known relative of the Galápagos finches is the South American 215: 211: 181: 125: 85: 2158: 2150: 2090: 2082: 2022: 2014: 1883: 1820: 1280: 1201: 1183: 1126: 2397: 5050: 4890: 4704: 4557: 4384: 4187: 3917: 3892: 3838: 3749: 3721: 3611: 3575: 3567: 3549: 3444: 3429: 3413: 3160: 3123: 2956: 756: 650: 451: 430:
Following his return from the voyage Darwin presented the finches to the
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Sato A, Tichy H, O'hUigin C, Grant PR, Grant BR, Klein J (March 2001).
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along with the New World sparrows and Old World buntings. However, the
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Charles Darwin's zoology notes & specimen lists from H.M.S. Beagle
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Philosophical Transactions of the Royal Society B: Biological Sciences
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is somewhat like that of a starling, and that of the fourth subgroup,
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Darwin's field notes on the Galapagos: 'A little world within itself'
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Phylogeny of Darwin's finches as revealed by mtDNA sequences in PNAS
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Strange Galapagos "Big Bird" reveals outcome of super-fast evolution
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Leah Burrows, Harvard University: News and Events. November 12, 2021
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For some decades, taxonomists have placed these birds in the family
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the whole group, with the exception of one species of the sub-group
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Intelligent Thought: Science versus the Intelligent Design Movement
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How and Why Species Multiply: The Radiation of Darwin's Finches
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has documented evolutionary changes in beak size affected by
554:, lately brought from Bow Island, in the Low Archipelago. Of 1222:"Beaks, Adaptation, and Vocal Evolution in Darwin's Finches" 1239:
10.1641/0006-3568(2004)054[0501:baavei]2.0.co;2
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Whereas Darwin spent just five weeks in the Galápagos, and
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and had been keen on shooting, but he had no expertise in
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Galapagos Online. List of birds of the Galapagos Islands.
1907:"Galapagos finches caught in act of becoming new species" 1755:"Galapagos finches caught in act of becoming new species" 498:
At the time that he rewrote his diary for publication as
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revealed a 240 kilobase haplotype encompassing the
2243:"Chapter 1, The Foundations of the 'Origin of Species'" 483:, FitzRoy's steward Harry Fuller, and Darwin's servant 1953:"Galapagos Finch Evolution – Dan Lewitt – HHMI (2013)" 1333:
For Darwin's finches, beak shape goes beyond evolution
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Four of Darwin's finches, clockwise (from top left):
270:, the others are found only on the Galápagos Islands. 27:
Group of related bird species in the Galápagos Islands
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A classification of the bird species of South America
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For the 1845 second edition of 2711:Darwin's Finches Evolve Before Scientists' Eyes 2538:, vol. 145, no. 4, pp. 300–320, 1343: 1341: 1142:"Evolution of Darwin's finches and their beaks" 611: 547: 2573:collections of Darwin's finches (Geospizinae)" 2353:(1940), "Evolution of the Galapagos Finches", 2754: 2616:Darwin in Galapagos: Footsteps to a New World 1850:"Rapid hybrid speciation in Darwin's finches" 1350:Darwin in Galapagos: Footsteps to a New World 8: 2719:Darwin's finches inspired the naming of the 2332:Grant, Peter R.; Grant, B. Rosemary (2008), 1581:Grant, B. Rosemary; Grant, Peter R. (1989). 1468: 1436: 1352:. Princeton, NJ: Princeton University Press. 1348:Grant, K. Thalia; Estes, Gregory B. (2009). 398:Seen here is adaptive radiation of finch A ( 2635:, vol. 15, no. 1, pp. 1–53, 2476:, New Haven, Conn.: Yale University Press, 1362: 1317: 714:, of uncertain place (Remsen et al. 2007). 4362: 3922: 3843: 3834: 3825: 3563: 3409: 3396: 3357: 3353: 3326: 3265: 3261: 3250: 3246: 2761: 2747: 2739: 2323:: CS1 maint: location missing publisher ( 1656:: CS1 maint: location missing publisher ( 1605:: CS1 maint: location missing publisher ( 1093:"Darwin's iconic finches join genome club" 1078: 1015:Developmental research in 2004 found that 40: 31: 2640: 2311:Ecology and evolution of Darwin's finches 2265:, London: Michael Joseph, Penguin Group, 1873: 1723: 1712:"Learning about birds from their genomes" 1270: 1237: 1191: 1116: 2713:: new developments reported 13 July 2006 2683:Genetics and the Origin of Birds Species 2592:"Why Darwin Rejected Intelligent Design" 1700:Peter Hess, Inverse. November 27th, 2017 1682: 1632:John Maynard Smith (1998). "Chapter 5". 1480: 1453: 1425: 1385: 1220:Podos, Jeffrey; Nowiki, Stephen (2004). 2619:, Princeton: Princeton University Press 1215: 1213: 1211: 1071: 575:, is shown in Fig. 4. The beak of 371:, and after finding a different one on 5192:Endemic birds of the Galápagos Islands 2316: 2298:, no. Spring 2006, pp. 32–53 2213:, vol. III, London: Henry Colburn 1649: 1598: 1564: 1548: 1537: 1522: 1506: 1491: 1449: 1447: 1445: 1377: 710:check-list, the Galápagos species are 2601:, New York: Vintage, pp. 107–126 1670: 1619: 1272:10.1093/oxfordjournals.molbev.a003806 7: 5164:E9CFF3AD-4E18-4236-8887-146E75D96F14 4320: 2222:(2nd. ed.), London: John Murray 1928:Starr, Michelle (24 November 2017). 367:was similar to those he had seen in 277:in 1936, and popularised in 1947 by 4344: 2701:Galápagos Online. Darwin's Finches. 2613:Grant, K. T.; Estes, G. B. (2009), 2497:rather than a family of their own ( 2233:(1st ed.), London: John Murray 1905:Galloway, Rory (23 November 2017). 1753:Galloway, Rory (23 November 2017). 1255:"On the Origin of Darwin's Finches" 1140:Koffmar, Linda (12 February 2015). 1027:of a gene called calmodulin (CaM). 235:and are not closely related to the 2624:Sulloway, Frank J. (Spring 1982), 2389:Check-list of North American Birds 2249:, vol. 2, London: John Murray 247:). They were first collected when 25: 3222:Late Quaternary prehistoric birds 2678:Different bills and song melodies 2633:Journal of the History of Biology 2428:The Firefly Encyclopedia of Birds 1091:Marsh, Geoff (11 February 2015). 1011:Molecular basis of beak evolution 4343: 4331: 4319: 4308: 4307: 4198:(New World vultures and condors) 1940:from the original on 2017-12-09. 1917:from the original on 2017-11-30. 1710:Holmes, Bob (10 February 2022). 619:Polymorphism in Darwin's finches 72: 5212:History of evolutionary biology 2179:Kaplan, Sarah (22 April 2016). 1959:from the original on 2017-11-07 1415:from the original on 2011-08-21 1148:from the original on 2018-04-24 694:puts Darwin's finches with the 2418:"Tanagers and Tanager-Finches" 2394:American Ornithologists' Union 2336:, Princeton University Press, 1585:. Chicago. p. 241 first para. 1144:. Uppsala University, Sweden. 700:American Ornithologists' Union 525:development of Darwin's theory 1: 2296:The Virginia Quarterly Review 1298:. Cambridge University Press. 446:that Darwin had thought were 2459:, Cambridge University Press 2292:"Confessions of a Darwinist" 2245:, in Darwin, Francis (ed.), 1017:bone morphogenetic protein 4 432:Zoological Society of London 334:inception of Darwin's theory 4262:(woodpeckers and relatives) 2597:, in Brockman, John (ed.), 2590:Sulloway, Frank J. (2006), 2526:Steinheimer, F. D. (2004), 2505:Zimmer, J. (5 April 2007), 2470:Sibley, Charles G. (1993), 2396:, 1998–2006, archived from 909:) – sometimes separated in 585:a bird originally a buzzard 383:of another, and those from 5238: 4124:(cormorants and relatives) 3217:Extinct species since 1500 2473:A World Checklist of Birds 2431:, Firefly Books, pp.  591:of the American continent. 206:) are a group of about 18 4303: 4254:(kingfishers and rollers) 4108:(albatrosses and petrels) 3992:(swifts and hummingbirds) 3974:(nightjars and relatives) 3824: 3798:(pheasants and relatives) 3356: 3325: 3264: 3260: 3249: 3245: 2781: 2544:10.1007/s10336-004-0043-8 1725:10.1146/knowable-021022-1 783:Sharp-beaked ground finch 249:the second voyage of the 156: 69:Scientific classification 48: 39: 4153:(seriemas and relatives) 4132:(pelicans and relatives) 2309:Grant, Peter R. (1999), 2227:Darwin, Charles (1859), 2218:Darwin, Charles (1845), 1636:(2nd ed.). Oxford. 1469:Desmond & Moore 1991 1437:Desmond & Moore 1991 1001:Peter and Rosemary Grant 986:Platyspiza crassirostris 797:Geospiza septentrionalis 692:Sibley–Ahlquist taxonomy 629:Peter and Rosemary Grant 606:On the Origin of Species 598:On the Origin of Species 511:The Voyage of the Beagle 505:The Voyage of the Beagle 493:The Voyage of the Beagle 353:transmutation of species 4246:(hornbills and hoopoes) 4161:(falcons and relatives) 2735:Kimball's Biology Pages 2727:Aug 2006 Nature Article 1999:10.1126/science.1098095 1875:10.1126/science.aao4593 1007:cycles in the Pacific. 920:Camarhynchus heliobates 877:Camarhynchus psittacula 618: 322:University of Edinburgh 289:and the largest is the 241:dull-coloured grassquit 5217:Polymorphism (biology) 4230:(trogons and quetzals) 4042:(cranes and relatives) 3312:(emus and cassowaries) 3202:Glossary of bird terms 3014:Confuciusornithiformes 2536:Journal of Ornithology 1622:, p. 428 in Afterword. 1184:10.1098/rstb.2009.0280 1079:Grant & Grant 2008 750: 667:linkage disequilibrium 616: 593: 546: 521: 427: 300: 5025:Paleobiology Database 4865:Paleobiology Database 4532:Paleobiology Database 4071:(kagu and sunbittern) 4034:(gulls and relatives) 3039:Songlingornithiformes 3004:Omnivoropterygiformes 2685:, Grant and Grant in 1634:Evolutionary genetics 968:Pinaroloxias inornata 907:Camarhynchus pallidus 897:Camarhynchus parvulus 837:Geospiza magnirostris 813:Genovesa cactus finch 773:Española cactus finch 767:Geospiza acutirostris 763:Genovesa ground finch 747:Camarhynchus parvulus 729:Geospiza magnirostris 725: 649:If the population is 541: 537:Journal of Researches 516: 438:, the famous English 407:Geospiza magnirostris 400:Geospiza magnirostris 397: 363:, he recorded that a 228:. They belong to the 18:Darwin's Finches 3893:Phoenicopteriformes 2413:Skutch, Alexander F. 1397:Chancellor, Gordon; 1294:Lack, David (1947). 777:Geospiza conirostris 659:sympatric speciation 655:Geospiza conirostris 627:spent three months, 3864:Mesitornithiformes 3858:(doves and pigeons) 3192:Families and orders 3054:Ichthyornithiformes 3029:Patagopterygiformes 2423:Christopher Perrins 2365:1940Natur.146..324L 2185:Speaking of Science 2135:10.1038/nature14181 2127:2015Natur.518..371L 2067:10.1038/nature04843 2059:2006Natur.442..563A 1991:2004Sci...305.1462A 1985:(5689): 1462–1465. 1866:2018Sci...359..224L 1805:10.1038/nature14181 1797:2015Natur.518..371L 1178:(1543): 1093–1098. 1109:2015Natur.518..147M 1025:spatial expressions 936:Green warbler-finch 887:Camarhynchus pauper 843:Common cactus finch 827:Geospiza fuliginosa 803:Medium ground finch 787:Geospiza difficilis 500:Journal and Remarks 461:Darwin had been in 389:Galápagos tortoises 202:(also known as the 62:Green warbler-finch 54:Medium ground finch 5197:Evolution of birds 4206:(eagles and hawks) 4106:Procellariiformes 4011:Opisthocomiformes 3980:Steatornithiformes 3207:List by population 3044:Hongshanornithidae 2969:Evolution of birds 2651:10.1007/BF00132004 2565:Sulloway, Frank J. 1439:, pp. 208–209 1052:Adaptive radiation 946:Grey warbler-finch 940:Certhidea olivacea 833:Large ground finch 823:Small ground finch 817:Geospiza propinqua 751: 669:is a possibility. 514:, Darwin said that 428: 417:Certhidea olivacea 245:Asemospiza obscura 50:Large ground finch 5222:Bird common names 5174: 5173: 5146:Open Tree of Life 4852:Open Tree of Life 4679:Open Tree of Life 4368:Taxon identifiers 4359: 4358: 4299: 4298: 4295: 4294: 4291: 4290: 4287: 4286: 4283: 4282: 4279: 4278: 4275: 4274: 4271: 4270: 4092:(loons or divers) 4061:Phaethontiformes 4053:Phaethontimorphae 4003:Opisthocomiformes 3972:Caprimulgiformes 3912: 3911: 3901:Podicipediformes 3820: 3819: 3816: 3815: 3812: 3811: 3538: 3537: 3321: 3320: 3278:Struthioniformes 3146:Waterfowl hunting 3079:Gastornithiformes 3074:Aepyornithiformes 3064:Lithornithiformes 2483:978-0-300-07083-5 2442:978-1-55297-777-4 2411:Burns, Kevin J.; 2343:978-0-691-13360-7 2272:978-0-7181-3430-3 2053:(7102): 563–567. 1860:(6372): 224–228. 1791:(7539): 371–375. 1716:Knowable Magazine 883:Medium tree finch 847:Geospiza scandens 529:natural selection 444:Galápagos Islands 349:Galápagos Islands 342:natural selection 307:the survey voyage 256:Galápagos Islands 204:Galápagos finches 197: 196: 35:Darwin's finches 16:(Redirected from 5229: 5167: 5166: 5154: 5153: 5141: 5140: 5128: 5127: 5115: 5114: 5102: 5101: 5089: 5088: 5076: 5075: 5074: 5061: 5060: 5059: 5033: 5032: 5020: 5019: 5007: 5006: 4994: 4993: 4981: 4980: 4968: 4967: 4955: 4954: 4942: 4941: 4929: 4928: 4926:DDACC3F87B82FF4C 4916: 4915: 4914: 4901: 4900: 4899: 4873: 4872: 4860: 4859: 4847: 4846: 4834: 4833: 4821: 4820: 4808: 4807: 4795: 4794: 4782: 4781: 4769: 4768: 4756: 4755: 4753:5F7D29F76143F763 4743: 4742: 4730: 4729: 4728: 4715: 4714: 4713: 4687: 4686: 4674: 4673: 4661: 4660: 4648: 4647: 4635: 4634: 4622: 4621: 4609: 4608: 4596: 4595: 4583: 4582: 4581: 4568: 4567: 4566: 4540: 4539: 4527: 4526: 4514: 4513: 4501: 4500: 4488: 4487: 4475: 4474: 4462: 4461: 4449: 4448: 4436: 4435: 4433:358FEF970ECCD004 4423: 4422: 4410: 4409: 4408: 4395: 4394: 4393: 4363: 4347: 4346: 4335: 4323: 4322: 4311: 4310: 4263: 4255: 4247: 4239: 4236:Leptosomiformes 4231: 4223: 4215: 4207: 4204:Accipitriformes 4199: 4178: 4177:(perching birds) 4170: 4162: 4154: 4133: 4125: 4117: 4109: 4101: 4098:Sphenisciformes 4093: 4072: 4064: 4043: 4035: 4032:Charadriiformes 4014: 3993: 3975: 3954: 3946: 3943:Musophagiformes 3938: 3923: 3904: 3896: 3875: 3867: 3859: 3844: 3835: 3826: 3799: 3786: 3564: 3560: 3553: 3466: 3459: 3423: 3410: 3406: 3397: 3393: 3358: 3354: 3327: 3313: 3305: 3297: 3289: 3281: 3266: 3262: 3251: 3247: 3114:Bird collections 3069:Dinornithiformes 2974:Darwin's finches 2964:Origin of flight 2904:Seabird breeding 2894:Sexual selection 2763: 2756: 2749: 2740: 2667: 2666: 2665: 2644: 2630: 2620: 2602: 2596: 2586: 2585: 2584: 2560: 2559: 2558: 2522: 2521: 2520: 2492: 2491: 2490: 2466: 2465: 2464: 2445: 2420: 2407: 2406: 2405: 2383: 2373:10.1038/146324a0 2346: 2328: 2322: 2314: 2305: 2304: 2303: 2283: 2250: 2234: 2223: 2214: 2193: 2192: 2187:. Archived from 2176: 2170: 2169: 2109: 2103: 2102: 2041: 2035: 2034: 1974: 1968: 1967: 1965: 1964: 1948: 1942: 1941: 1925: 1919: 1918: 1902: 1896: 1895: 1877: 1845: 1839: 1838: 1836: 1835: 1775: 1769: 1768: 1766: 1765: 1750: 1744: 1743: 1741: 1740: 1727: 1707: 1701: 1695: 1689: 1680: 1674: 1668: 1662: 1661: 1655: 1647: 1629: 1623: 1617: 1611: 1610: 1604: 1596: 1578: 1572: 1562: 1556: 1546: 1540: 1535: 1529: 1520: 1514: 1504: 1498: 1489: 1483: 1478: 1472: 1466: 1460: 1451: 1440: 1434: 1428: 1423: 1417: 1416: 1401:(October 2006), 1394: 1388: 1375: 1369: 1363:Steinheimer 2004 1360: 1354: 1353: 1345: 1336: 1330: 1324: 1318:Steinheimer 2004 1315: 1309: 1299: 1296:Darwin's Finches 1291: 1285: 1284: 1274: 1250: 1244: 1243: 1241: 1217: 1206: 1205: 1195: 1163: 1157: 1156: 1154: 1153: 1137: 1131: 1130: 1120: 1088: 1082: 1076: 1057:Island gigantism 1021:different timing 982:Vegetarian finch 903:Woodpecker finch 893:Small tree finch 873:Large tree finch 412:Geospiza parvula 291:vegetarian finch 283:Darwin's Finches 266:, which is from 200:Darwin's finches 152:Groups included 77: 76: 58:Small tree finch 44: 32: 21: 5237: 5236: 5232: 5231: 5230: 5228: 5227: 5226: 5177: 5176: 5175: 5170: 5162: 5157: 5149: 5144: 5136: 5131: 5123: 5118: 5110: 5105: 5097: 5092: 5084: 5079: 5070: 5069: 5064: 5055: 5054: 5049: 5036: 5028: 5023: 5015: 5010: 5002: 4997: 4989: 4984: 4976: 4971: 4963: 4958: 4950: 4945: 4937: 4932: 4924: 4919: 4910: 4909: 4904: 4895: 4894: 4889: 4876: 4868: 4863: 4855: 4850: 4842: 4837: 4829: 4824: 4816: 4811: 4803: 4798: 4790: 4785: 4777: 4772: 4764: 4759: 4751: 4746: 4738: 4733: 4724: 4723: 4718: 4709: 4708: 4703: 4690: 4682: 4677: 4669: 4664: 4656: 4651: 4643: 4638: 4630: 4625: 4617: 4612: 4604: 4599: 4591: 4586: 4577: 4576: 4571: 4562: 4561: 4556: 4543: 4535: 4530: 4522: 4517: 4509: 4504: 4496: 4491: 4483: 4478: 4470: 4465: 4457: 4452: 4444: 4439: 4431: 4426: 4418: 4413: 4404: 4403: 4398: 4389: 4388: 4383: 4370: 4360: 4355: 4267: 4261: 4253: 4245: 4244:Bucerotiformes 4238:(cuckoo-roller) 4237: 4229: 4221: 4213: 4205: 4197: 4196:Cathartiformes 4182: 4176: 4168: 4167:Psittaciformes 4160: 4152: 4137: 4131: 4130:Pelecaniformes 4123: 4115: 4107: 4099: 4091: 4076: 4070: 4069:Eurypygiformes 4062: 4047: 4041: 4033: 4018: 4012: 3997: 3991: 3973: 3958: 3952: 3944: 3936: 3908: 3902: 3894: 3879: 3873: 3872:Pterocliformes 3865: 3857: 3808: 3797: 3784: 3769: 3693: 3652: 3590: 3555: 3554: 3547: 3546: 3534: 3514: 3487: 3464: 3457: 3417: 3416: 3404: 3403: 3387: 3386: 3383: 3381: 3379: 3377: 3375: 3373: 3371: 3369: 3367: 3365: 3363: 3348: 3346: 3344: 3342: 3340: 3338: 3336: 3334: 3332: 3317: 3311: 3310:Casuariiformes 3303: 3302:Apterygiformes 3295: 3287: 3279: 3256: 3241: 3212:Lists by region 3180: 3090: 3083: 3024:Chaoyangiformes 3009:Jeholornithidae 2983: 2947:Origin of birds 2933: 2914:Brood parasites 2860: 2790: 2777: 2767: 2674: 2663: 2661: 2642:10.1.1.458.3975 2628: 2623: 2612: 2609: 2607:Further reading 2594: 2589: 2582: 2580: 2563: 2556: 2554: 2525: 2518: 2516: 2504: 2488: 2486: 2484: 2469: 2462: 2460: 2451:Keynes, Richard 2449: 2443: 2410: 2403: 2401: 2386: 2349: 2344: 2331: 2315: 2313:, Princeton, NJ 2308: 2301: 2299: 2288:Eldredge, Niles 2286: 2273: 2255:Desmond, Adrian 2253: 2239:Darwin, Francis 2237: 2226: 2217: 2206:Darwin, Charles 2204: 2201: 2196: 2178: 2177: 2173: 2121:(7539): 371–5, 2111: 2110: 2106: 2043: 2042: 2038: 1976: 1975: 1971: 1962: 1960: 1950: 1949: 1945: 1927: 1926: 1922: 1904: 1903: 1899: 1847: 1846: 1842: 1833: 1831: 1777: 1776: 1772: 1763: 1761: 1752: 1751: 1747: 1738: 1736: 1709: 1708: 1704: 1696: 1692: 1681: 1677: 1669: 1665: 1648: 1644: 1631: 1630: 1626: 1618: 1614: 1597: 1593: 1580: 1579: 1575: 1563: 1559: 1547: 1543: 1536: 1532: 1521: 1517: 1505: 1501: 1490: 1486: 1479: 1475: 1467: 1463: 1452: 1443: 1435: 1431: 1424: 1420: 1396: 1395: 1391: 1376: 1372: 1361: 1357: 1347: 1346: 1339: 1331: 1327: 1316: 1312: 1293: 1292: 1288: 1259:Mol. Biol. Evol 1252: 1251: 1247: 1219: 1218: 1209: 1165: 1164: 1160: 1151: 1149: 1139: 1138: 1134: 1118:10.1038/518147a 1090: 1089: 1085: 1077: 1073: 1069: 1061:island dwarfism 1043: 1013: 1005:El Niño/La Niña 997: 995:Modern research 950:Certhidea fusca 807:Geospiza fortis 741:Certhidea fusca 735:Geospiza fortis 720: 684: 679: 621: 601: 496: 426: 422:Geospiza fortis 318:John Edmonstone 303: 301:Darwin's theory 287:warbler-finches 71: 56: 28: 23: 22: 15: 12: 11: 5: 5235: 5233: 5225: 5224: 5219: 5214: 5209: 5204: 5199: 5194: 5189: 5187:Charles Darwin 5179: 5178: 5172: 5171: 5169: 5168: 5155: 5142: 5129: 5116: 5103: 5090: 5077: 5062: 5046: 5044: 5038: 5037: 5035: 5034: 5021: 5008: 4995: 4982: 4969: 4956: 4943: 4930: 4917: 4902: 4886: 4884: 4878: 4877: 4875: 4874: 4861: 4848: 4835: 4822: 4809: 4796: 4783: 4770: 4757: 4744: 4731: 4716: 4700: 4698: 4692: 4691: 4689: 4688: 4675: 4662: 4649: 4636: 4623: 4610: 4597: 4584: 4569: 4553: 4551: 4545: 4544: 4542: 4541: 4528: 4515: 4502: 4489: 4476: 4463: 4450: 4437: 4424: 4411: 4396: 4380: 4378: 4372: 4371: 4366: 4357: 4356: 4354: 4353: 4341: 4329: 4317: 4304: 4301: 4300: 4297: 4296: 4293: 4292: 4289: 4288: 4285: 4284: 4281: 4280: 4277: 4276: 4273: 4272: 4269: 4268: 4266: 4265: 4257: 4252:Coraciiformes 4249: 4241: 4233: 4228:Trogoniformes 4225: 4217: 4209: 4201: 4192: 4190: 4184: 4183: 4181: 4180: 4175:Passeriformes 4172: 4164: 4159:Falconiformes 4156: 4151:Cariamiformes 4147: 4145: 4139: 4138: 4136: 4135: 4127: 4119: 4114:Ciconiiformes 4111: 4103: 4095: 4086: 4084: 4078: 4077: 4075: 4074: 4066: 4057: 4055: 4049: 4048: 4046: 4045: 4037: 4028: 4026: 4024:Cursorimorphae 4020: 4019: 4017: 4016: 4007: 4005: 3999: 3998: 3996: 3995: 3987: 3982: 3977: 3968: 3966: 3960: 3959: 3957: 3956: 3948: 3940: 3931: 3929: 3920: 3914: 3913: 3910: 3909: 3907: 3906: 3898: 3889: 3887: 3885:Mirandornithes 3881: 3880: 3878: 3877: 3869: 3861: 3856:Columbiformes 3852: 3850: 3848:Columbimorphae 3841: 3832: 3822: 3821: 3818: 3817: 3814: 3813: 3810: 3809: 3807: 3806: 3801: 3793: 3788: 3783:Meleagridinae 3779: 3777: 3771: 3770: 3768: 3767: 3760: 3753: 3746: 3739: 3732: 3725: 3718: 3711: 3703: 3701: 3699:Odontophoridae 3695: 3694: 3692: 3691: 3684: 3677: 3670: 3662: 3660: 3654: 3653: 3651: 3650: 3643: 3636: 3629: 3622: 3615: 3608: 3600: 3598: 3592: 3591: 3589: 3588: 3583: 3578: 3572: 3570: 3561: 3540: 3539: 3536: 3535: 3533: 3532: 3524: 3522: 3516: 3515: 3513: 3512: 3505: 3497: 3495: 3489: 3488: 3486: 3485: 3480: 3478:Stictonettinae 3475: 3473:Dendrocygninae 3470: 3469: 3468: 3461: 3449: 3448: 3447: 3442: 3437: 3426: 3424: 3407: 3394: 3351: 3323: 3322: 3319: 3318: 3316: 3315: 3307: 3299: 3291: 3283: 3274: 3272: 3258: 3257: 3254: 3243: 3242: 3240: 3239: 3238: 3237: 3232: 3226:Notable birds 3224: 3219: 3214: 3209: 3204: 3199: 3194: 3188: 3186: 3182: 3181: 3179: 3178: 3176:Egg collecting 3173: 3168: 3163: 3158: 3153: 3148: 3143: 3138: 3133: 3128: 3127: 3126: 3116: 3111: 3106: 3101: 3095: 3093: 3085: 3084: 3082: 3081: 3076: 3071: 3066: 3061: 3059:Hesperornithes 3056: 3051: 3046: 3041: 3036: 3034:Ambiortiformes 3031: 3026: 3021: 3019:Enantiornithes 3016: 3011: 3006: 3001: 2993: 2991: 2985: 2984: 2982: 2981: 2976: 2971: 2966: 2961: 2960: 2959: 2954: 2943: 2941: 2935: 2934: 2932: 2931: 2926: 2921: 2916: 2911: 2906: 2901: 2896: 2891: 2886: 2881: 2876: 2870: 2868: 2862: 2861: 2859: 2858: 2853: 2848: 2843: 2838: 2833: 2828: 2823: 2822: 2821: 2811: 2806: 2800: 2798: 2792: 2791: 2789: 2788: 2782: 2779: 2778: 2768: 2766: 2765: 2758: 2751: 2743: 2737: 2736: 2730: 2724: 2714: 2708: 2703: 2698: 2689: 2680: 2673: 2672:External links 2670: 2669: 2668: 2621: 2608: 2605: 2604: 2603: 2587: 2561: 2523: 2502: 2482: 2467: 2447: 2441: 2408: 2384: 2347: 2342: 2329: 2306: 2284: 2271: 2251: 2235: 2224: 2215: 2200: 2197: 2195: 2194: 2191:on 2016-05-31. 2171: 2104: 2036: 1969: 1943: 1920: 1897: 1840: 1770: 1745: 1702: 1690: 1675: 1663: 1643:978-0198502319 1642: 1624: 1612: 1592:978-0226305905 1591: 1573: 1557: 1541: 1530: 1515: 1499: 1484: 1473: 1461: 1441: 1429: 1418: 1399:Keynes, Randal 1389: 1370: 1355: 1337: 1325: 1310: 1286: 1265:(3): 299–311. 1245: 1232:(6): 501–510. 1207: 1158: 1132: 1083: 1070: 1068: 1065: 1064: 1063: 1054: 1049: 1042: 1039: 1012: 1009: 996: 993: 992: 991: 990: 989: 973: 972: 971: 955: 954: 953: 943: 925: 924: 923: 916:Mangrove finch 913: 900: 890: 880: 862: 861: 860: 850: 840: 830: 820: 810: 800: 790: 780: 770: 719: 716: 712:incertae sedis 708:South American 704:North American 683: 680: 678: 675: 620: 617: 600: 594: 495: 489: 481:Robert FitzRoy 476:South American 425: 424: 419: 414: 409: 403: 373:Charles Island 361:Chatham Island 330:Syms Covington 302: 299: 260:Charles Darwin 195: 194: 193: 192: 185: 178: 171: 164: 154: 153: 149: 148: 143: 139: 138: 133: 129: 128: 123: 119: 118: 113: 109: 108: 103: 99: 98: 93: 89: 88: 83: 79: 78: 65: 64: 46: 45: 37: 36: 26: 24: 14: 13: 10: 9: 6: 4: 3: 2: 5234: 5223: 5220: 5218: 5215: 5213: 5210: 5208: 5205: 5203: 5200: 5198: 5195: 5193: 5190: 5188: 5185: 5184: 5182: 5165: 5160: 5156: 5152: 5147: 5143: 5139: 5134: 5130: 5126: 5121: 5117: 5113: 5108: 5104: 5100: 5095: 5091: 5087: 5082: 5078: 5073: 5067: 5063: 5058: 5052: 5048: 5047: 5045: 5043: 5039: 5031: 5026: 5022: 5018: 5013: 5009: 5005: 5000: 4996: 4992: 4987: 4983: 4979: 4974: 4970: 4966: 4961: 4957: 4953: 4948: 4944: 4940: 4935: 4931: 4927: 4922: 4918: 4913: 4907: 4903: 4898: 4892: 4888: 4887: 4885: 4883: 4879: 4871: 4866: 4862: 4858: 4853: 4849: 4845: 4840: 4836: 4832: 4827: 4823: 4819: 4814: 4810: 4806: 4801: 4797: 4793: 4788: 4784: 4780: 4775: 4771: 4767: 4762: 4758: 4754: 4749: 4745: 4741: 4736: 4732: 4727: 4721: 4717: 4712: 4706: 4702: 4701: 4699: 4697: 4693: 4685: 4680: 4676: 4672: 4667: 4663: 4659: 4654: 4650: 4646: 4641: 4637: 4633: 4628: 4624: 4620: 4615: 4611: 4607: 4602: 4598: 4594: 4589: 4585: 4580: 4574: 4570: 4565: 4559: 4555: 4554: 4552: 4550: 4546: 4538: 4533: 4529: 4525: 4520: 4516: 4512: 4507: 4503: 4499: 4494: 4490: 4486: 4481: 4477: 4473: 4468: 4464: 4460: 4455: 4451: 4447: 4442: 4438: 4434: 4429: 4425: 4421: 4416: 4412: 4407: 4401: 4397: 4392: 4386: 4382: 4381: 4379: 4377: 4373: 4369: 4364: 4352: 4351: 4342: 4340: 4339: 4334: 4330: 4328: 4327: 4318: 4316: 4315: 4306: 4305: 4302: 4264: 4258: 4256: 4250: 4248: 4242: 4240: 4234: 4232: 4226: 4224: 4218: 4216: 4212:Strigiformes 4210: 4208: 4202: 4200: 4194: 4193: 4191: 4189: 4185: 4179: 4173: 4171: 4165: 4163: 4157: 4155: 4149: 4148: 4146: 4144: 4140: 4134: 4128: 4126: 4120: 4118: 4112: 4110: 4104: 4102: 4096: 4094: 4088: 4087: 4085: 4083: 4079: 4073: 4067: 4065: 4063:(tropicbirds) 4059: 4058: 4056: 4054: 4050: 4044: 4038: 4036: 4030: 4029: 4027: 4025: 4021: 4015: 4009: 4008: 4006: 4004: 4000: 3994: 3988: 3986: 3985:Podargiformes 3983: 3981: 3978: 3976: 3970: 3969: 3967: 3965: 3961: 3955: 3949: 3947: 3941: 3939: 3935:Cuculiformes 3933: 3932: 3930: 3928: 3924: 3921: 3919: 3915: 3905: 3899: 3897: 3891: 3890: 3888: 3886: 3882: 3876: 3870: 3868: 3862: 3860: 3854: 3853: 3851: 3849: 3845: 3842: 3840: 3836: 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3308: 3306: 3300: 3298: 3294:Tinamiformes 3292: 3290: 3284: 3282: 3276: 3275: 3273: 3271: 3270:Palaeognathae 3267: 3263: 3259: 3252: 3248: 3244: 3236: 3233: 3231: 3228: 3227: 3225: 3223: 3220: 3218: 3215: 3213: 3210: 3208: 3205: 3203: 3200: 3198: 3195: 3193: 3190: 3189: 3187: 3183: 3177: 3174: 3172: 3169: 3167: 3164: 3162: 3159: 3157: 3156:Pigeon racing 3154: 3152: 3149: 3147: 3144: 3142: 3139: 3137: 3134: 3132: 3129: 3125: 3122: 3121: 3120: 3117: 3115: 3112: 3110: 3107: 3105: 3102: 3100: 3097: 3096: 3094: 3092: 3086: 3080: 3077: 3075: 3072: 3070: 3067: 3065: 3062: 3060: 3057: 3055: 3052: 3050: 3047: 3045: 3042: 3040: 3037: 3035: 3032: 3030: 3027: 3025: 3022: 3020: 3017: 3015: 3012: 3010: 3007: 3005: 3002: 3000: 2999: 2998:Archaeopteryx 2995: 2994: 2992: 2990: 2986: 2980: 2977: 2975: 2972: 2970: 2967: 2965: 2962: 2958: 2955: 2953: 2950: 2949: 2948: 2945: 2944: 2942: 2940: 2936: 2930: 2927: 2925: 2922: 2920: 2917: 2915: 2912: 2910: 2907: 2905: 2902: 2900: 2897: 2895: 2892: 2890: 2887: 2885: 2882: 2880: 2877: 2875: 2872: 2871: 2869: 2867: 2863: 2857: 2854: 2852: 2849: 2847: 2844: 2842: 2839: 2837: 2834: 2832: 2829: 2827: 2824: 2820: 2817: 2816: 2815: 2812: 2810: 2807: 2805: 2802: 2801: 2799: 2797: 2793: 2787: 2784: 2783: 2780: 2775: 2771: 2764: 2759: 2757: 2752: 2750: 2745: 2744: 2741: 2734: 2731: 2728: 2725: 2722: 2718: 2715: 2712: 2709: 2707: 2704: 2702: 2699: 2697: 2695: 2690: 2688: 2684: 2681: 2679: 2676: 2675: 2671: 2660: 2656: 2652: 2648: 2643: 2638: 2634: 2627: 2622: 2618: 2617: 2611: 2610: 2606: 2600: 2593: 2588: 2578: 2574: 2572: 2566: 2562: 2553: 2549: 2545: 2541: 2537: 2533: 2531: 2524: 2515:on 2009-03-02 2514: 2510: 2509: 2503: 2500: 2496: 2485: 2479: 2475: 2474: 2468: 2458: 2457: 2452: 2448: 2444: 2438: 2434: 2430: 2429: 2424: 2419: 2414: 2409: 2400:on 2007-04-04 2399: 2395: 2391: 2390: 2385: 2382: 2378: 2374: 2370: 2366: 2362: 2358: 2357: 2352: 2348: 2345: 2339: 2335: 2330: 2326: 2320: 2312: 2307: 2297: 2293: 2289: 2285: 2282: 2278: 2274: 2268: 2264: 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1534: 1531: 1528: 1524: 1519: 1516: 1512: 1508: 1503: 1500: 1497: 1493: 1488: 1485: 1482: 1481:Sulloway 2006 1477: 1474: 1471:, p. 248 1470: 1465: 1462: 1459: 1455: 1454:Sulloway 1982 1450: 1448: 1446: 1442: 1438: 1433: 1430: 1427: 1426:Eldredge 2006 1422: 1419: 1414: 1410: 1409:Darwin Online 1406: 1405: 1400: 1393: 1390: 1387: 1386:Eldredge 2006 1383: 1379: 1374: 1371: 1368: 1364: 1359: 1356: 1351: 1344: 1342: 1338: 1334: 1329: 1326: 1323: 1319: 1314: 1311: 1307: 1306:0-521-25243-1 1303: 1297: 1290: 1287: 1282: 1278: 1273: 1268: 1264: 1260: 1256: 1249: 1246: 1240: 1235: 1231: 1227: 1223: 1216: 1214: 1212: 1208: 1203: 1199: 1194: 1189: 1185: 1181: 1177: 1173: 1169: 1162: 1159: 1147: 1143: 1136: 1133: 1128: 1124: 1119: 1114: 1110: 1106: 1103:(7538): 147. 1102: 1098: 1094: 1087: 1084: 1080: 1075: 1072: 1066: 1062: 1058: 1055: 1053: 1050: 1048: 1047:Species flock 1045: 1044: 1040: 1038: 1035: 1030: 1026: 1022: 1018: 1010: 1008: 1006: 1002: 994: 987: 983: 980: 979: 978: 974: 969: 965: 962: 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276: 271: 269: 265: 261: 257: 254:visited the 253: 252: 246: 242: 238: 234: 231: 227: 224: 220: 217: 213: 209: 205: 201: 191: 190: 186: 184: 183: 179: 177: 176: 172: 170: 169: 165: 163: 162: 158: 157: 155: 150: 147: 144: 141: 140: 137: 134: 132:Superfamily: 131: 130: 127: 126:Passeriformes 124: 121: 120: 117: 114: 111: 110: 107: 104: 101: 100: 97: 94: 91: 90: 87: 84: 81: 80: 75: 70: 66: 63: 59: 55: 51: 47: 43: 38: 33: 30: 19: 5072:Pinaroloxias 5042:Pinaroloxias 5041: 4881: 4740:Camarhynchus 4726:Camarhynchus 4696:Camarhynchus 4695: 4548: 4375: 4348: 4336: 4324: 4312: 4222:(mousebirds) 4220:Coliiformes 4090:Gaviiformes 4082:Aequornithes 3990:Apodiformes 3951:Otidiformes 3927:Otidimorphae 3874:(sandgrouse) 3796:Phasianinae 3762: 3755: 3748: 3743:Odontophorus 3741: 3734: 3727: 3720: 3713: 3706: 3686: 3679: 3672: 3665: 3645: 3638: 3631: 3624: 3617: 3610: 3603: 3581:Oreophasinae 3527: 3507: 3500: 3405:(waterfowls) 3401:Anseriformes 3151:Cockfighting 3136:Conservation 3131:Bird feeding 3119:Birdwatching 3109:Ornithomancy 3049:Gansuiformes 2996: 2989:Fossil birds 2973: 2879:Intelligence 2693: 2682: 2662:, retrieved 2632: 2615: 2598: 2581:, retrieved 2576: 2570: 2555:, retrieved 2535: 2532:, 1831–1836" 2529: 2517:, retrieved 2513:the original 2507: 2495:Fringillidae 2487:, retrieved 2472: 2461:, retrieved 2455: 2427: 2402:, retrieved 2398:the original 2388: 2354: 2333: 2310: 2300:, retrieved 2295: 2262: 2259:Moore, James 2246: 2229: 2219: 2209: 2189:the original 2184: 2174: 2118: 2114: 2107: 2050: 2046: 2039: 1982: 1978: 1972: 1961:. 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Retrieved 1135: 1100: 1096: 1086: 1074: 1014: 998: 985: 976: 967: 959:Pinaroloxias 958: 949: 939: 928: 919: 910: 906: 896: 886: 876: 867:Camarhynchus 865: 856: 846: 836: 826: 816: 806: 796: 786: 776: 766: 755: 745: 739: 733: 727: 711: 685: 671: 654: 648: 636: 633: 622: 612: 604: 602: 597: 581:Camarhynchus 580: 576: 572: 567: 563: 559: 555: 551: 548: 542: 536: 535:(now titled 532: 522: 517: 509: 503: 499: 497: 492: 471:mockingbirds 460: 429: 421: 416: 411: 406: 399: 346: 312: 304: 282: 281:in his book 272: 268:Cocos Island 250: 244: 237:true finches 225: 218: 203: 199: 198: 189:Pinaroloxias 187: 180: 173: 168:Camarhynchus 166: 159: 136:Emberizoidea 29: 5094:iNaturalist 5066:Wikispecies 4973:iNaturalist 4906:Wikispecies 4800:iNaturalist 4720:Wikispecies 4627:iNaturalist 4573:Wikispecies 4493:iNaturalist 4400:Wikispecies 4350:WikiProject 4260:Piciformes 4143:Australaves 4122:Suliformes 4040:Gruiformes 3895:(flamingos) 3804:Tetraoninae 3775:Phasianidae 3764:Rhynchortyx 3729:Dactylortyx 3596:Megapodidae 3586:Penelopinae 3544:Galliformes 3286:Rheiformes 3280:(ostriches) 3230:individuals 3104:Ornithology 3091:interaction 2846:Preen gland 2717:Fink F.A.Q. 2351:Lack, David 1685:, pp.  1567:, pp.  1565:Darwin 1859 1549:Darwin 1845 1538:Darwin 1887 1525:, pp.  1523:Darwin 1845 1507:Darwin 1839 1494:, pp.  1492:Darwin 1839 1456:, pp.  1378:Keynes 2000 1365:, pp.  1081:, p. 3 964:Cocos finch 688:Emberizidae 452:gross-beaks 365:mockingbird 326:ornithology 264:Cocos finch 219:Geospizinae 5207:Geospizini 5181:Categories 4579:Platyspiza 4549:Platyspiza 4100:(penguins) 3953:(bustards) 3791:Perdicinae 3736:Dendrortyx 3708:Callipepla 3640:Megapodius 3605:Aepypodius 3483:Tadorninae 3465:true geese 3296:(tinamous) 3255:Neornithes 3166:Pheasantry 3141:Aviculture 2909:Incubation 2899:Lek mating 2733:Speciation 2664:2008-12-09 2583:2008-12-08 2557:2008-12-08 2519:2007-04-09 2499:Thraupidae 2489:2013-12-06 2463:2008-12-08 2404:2007-04-09 2302:2008-11-04 2199:References 1963:2017-06-06 1834:2022-02-14 1764:2022-02-14 1739:2022-02-11 1673:, plate 7. 1671:Grant 1999 1620:Grant 1999 1551:, p.  1509:, p.  1380:, p.  1320:, p.  1226:BioScience 1152:2018-04-24 1029:Calmodulin 977:Platyspiza 911:Cactospiza 625:David Lack 596:Text from 533:The Voyage 491:Text from 448:blackbirds 436:John Gould 279:David Lack 275:Percy Lowe 226:Geospizini 175:Platyspiza 146:Thraupidae 5057:Q10807310 4912:Certhidea 4882:Certhidea 4564:Q10807482 4169:(parrots) 4013:(hoatzin) 3964:Strisores 3945:(turacos) 3937:(cuckoos) 3866:(mesites) 3785:(turkeys) 3757:Philortyx 3674:Agelastes 3667:Acryllium 3658:Numididae 3647:Talegalla 3557:gamebirds 3550:landfowls 3529:Anseranas 3493:Anhimidae 3452:Anserinae 3235:fictional 2957:dinosaurs 2952:Theropoda 2939:Evolution 2884:Migration 2866:Behaviour 2637:CiteSeerX 2281:185764721 2211:1832–1836 2143:0028-0836 2075:0028-0836 2007:0036-8075 1892:206663426 1813:1476-4687 1734:246769765 1652:cite book 1601:cite book 930:Certhidea 857:G. fortis 702:, in its 651:panmictic 577:Cactornis 573:Certhidea 564:Certhidea 556:Cactornis 552:Cactornis 519:warbler". 485:Covington 463:Cambridge 381:Albemarle 338:evolution 216:subfamily 212:passerine 182:Certhidea 92:Kingdom: 86:Eukaryota 5051:Wikidata 4891:Wikidata 4711:Q2173058 4705:Wikidata 4558:Wikidata 4420:Geospiza 4406:Geospiza 4385:Wikidata 4376:Geospiza 4314:Category 4188:Afroaves 4116:(storks) 3918:Passerea 3903:(grebes) 3839:Columbea 3750:Oreortyx 3722:Cyrtonyx 3612:Alectura 3576:Cracinae 3568:Cracidae 3445:Oxyurini 3435:Aythyini 3430:Anatinae 3414:Anatidae 3161:Falconry 3124:big year 2979:Seabirds 2889:Foraging 2836:Feathers 2659:17161535 2567:(1982), 2552:24957761 2453:(2000), 2415:(2003), 2381:43465549 2319:citation 2290:(2006), 2261:(1991), 2241:(1887), 2208:(1839), 2159:25686609 2091:16885984 2031:17226774 2023:15353802 1957:Archived 1938:Archived 1915:Archived 1911:BBC News 1884:29170277 1821:25686609 1759:BBC News 1413:archived 1281:11230531 1202:20194171 1146:Archived 1127:25673391 1041:See also 853:Big Bird 757:Geospiza 696:tanagers 677:Taxonomy 589:Polybori 568:Geospiza 560:Geospiza 161:Geospiza 142:Family: 106:Chordata 102:Phylum: 96:Animalia 82:Domain: 5202:Finches 5159:ZooBank 5112:1018917 5086:2491136 4991:1292302 4965:2491688 4921:Avibase 4897:Q375385 4818:1212051 4792:2491661 4748:Avibase 4645:1391207 4619:2491747 4485:7237096 4428:Avibase 4391:Q662854 4326:Commons 3830:Neoaves 3715:Colinus 3681:Guttera 3619:Eulipoa 3440:Mergini 3304:(kiwis) 3288:(rheas) 3099:Ringing 2924:Hybrids 2919:Nesting 2874:Singing 2851:Plumage 2826:Dactyly 2796:Anatomy 2786:Outline 2776:: Aves) 2433:629–631 2425:(ed.), 2361:Bibcode 2167:4462253 2151:1586310 2123:Bibcode 2099:2416057 2083:1586310 2055:Bibcode 2015:1644869 1987:Bibcode 1979:Science 1862:Bibcode 1854:Science 1829:4462253 1793:Bibcode 1569:397–398 1527:379–380 1496:461–462 1367:301–303 1193:2830229 1105:Bibcode 718:Species 653:, then 638:Opuntia 502:(later 456:finches 357:Henslow 355:. From 347:On the 305:During 295:adapted 258:, with 230:tanager 208:species 122:Order: 112:Class: 5151:422526 5125:558067 5030:371380 5004:557498 4870:371389 4857:419366 4831:557477 4684:589958 4658:729196 4537:371375 4511:557700 4446:120618 4338:Portal 4214:(owls) 3688:Numida 3626:Leipoa 3509:Chauna 3502:Anhima 3197:Genera 3171:Imping 3089:Human 2929:Colony 2856:Vision 2841:Flight 2694:et al. 2657:  2639:  2571:Beagle 2550:  2530:Beagle 2480:  2439:  2379:  2356:Nature 2340:  2279:  2269:  2263:Darwin 2165:  2157:  2149:  2141:  2115:Nature 2097:  2089:  2081:  2073:  2047:Nature 2029:  2021:  2013:  2005:  1890:  1882:  1827:  1819:  1811:  1785:Nature 1732:  1640:  1589:  1304:  1279:  1200:  1190:  1125:  1097:Nature 975:Genus 957:Genus 927:Genus 864:Genus 754:Genus 682:Family 544:ends." 454:" and 377:Tahiti 313:Beagle 251:Beagle 233:family 5138:93069 5107:IRMNG 5099:10366 5017:48879 4986:IRMNG 4978:10447 4952:33768 4844:87174 4813:IRMNG 4779:34419 4766:62L3W 4671:48887 4640:IRMNG 4632:71751 4606:33766 4524:48881 4472:36192 3458:swans 3420:ducks 3390:fowls 3185:Lists 2809:Brain 2774:class 2770:Birds 2692:Sato 2655:S2CID 2629:(PDF) 2595:(PDF) 2569:"The 2548:S2CID 2421:, in 2377:S2CID 2163:S2CID 2095:S2CID 2027:S2CID 1888:S2CID 1825:S2CID 1730:S2CID 1458:57–58 1067:Notes 663:niche 385:James 369:Chile 223:tribe 5133:NCBI 5120:ITIS 5081:GBIF 5012:NCBI 4999:ITIS 4960:GBIF 4939:3L8Z 4839:NCBI 4826:ITIS 4805:9469 4787:GBIF 4666:NCBI 4653:ITIS 4614:GBIF 4593:6S57 4519:NCBI 4506:ITIS 4498:9476 4480:GBIF 4459:4N2N 4441:BOLD 2831:Eggs 2819:milk 2814:Crop 2804:Beak 2721:Fink 2687:PNAS 2478:ISBN 2437:ISBN 2338:ISBN 2325:link 2277:OCLC 2267:ISBN 2155:PMID 2147:OCLC 2139:ISSN 2087:PMID 2079:OCLC 2071:ISSN 2019:PMID 2011:OCLC 2003:ISSN 1880:PMID 1817:PMID 1809:ISSN 1658:link 1638:ISBN 1607:link 1587:ISBN 1302:ISBN 1277:PMID 1198:PMID 1123:PMID 1059:and 1034:ALX1 1023:and 643:aril 467:wren 311:HMS 116:Aves 4947:EoL 4934:CoL 4774:EoL 4761:CoL 4735:ADW 4601:EoL 4588:CoL 4467:EoL 4454:CoL 4415:ADW 2647:doi 2540:doi 2369:doi 2131:doi 2119:518 2063:doi 2051:442 1995:doi 1983:305 1870:doi 1858:359 1801:doi 1789:518 1720:doi 1553:380 1511:462 1384:.; 1382:xix 1322:300 1267:doi 1234:doi 1188:PMC 1180:doi 1176:365 1113:doi 1101:518 527:of 450:, " 340:by 336:of 309:of 221:or 210:of 5183:: 5161:: 5148:: 5135:: 5122:: 5109:: 5096:: 5083:: 5068:: 5053:: 5027:: 5014:: 5001:: 4988:: 4975:: 4962:: 4949:: 4936:: 4923:: 4908:: 4893:: 4867:: 4854:: 4841:: 4828:: 4815:: 4802:: 4789:: 4776:: 4763:: 4750:: 4737:: 4722:: 4707:: 4681:: 4668:: 4655:: 4642:: 4629:: 4616:: 4603:: 4590:: 4575:: 4560:: 4534:: 4521:: 4508:: 4495:: 4482:: 4469:: 4456:: 4443:: 4430:: 4417:: 4402:: 4387:: 2653:, 2645:, 2631:, 2575:, 2546:, 2534:, 2501:). 2435:, 2392:, 2375:, 2367:, 2321:}} 2317:{{ 2294:, 2275:, 2257:; 2183:. 2161:, 2153:, 2145:, 2137:, 2129:, 2117:, 2093:. 2085:. 2077:. 2069:. 2061:. 2049:. 2025:. 2017:. 2009:. 2001:. 1993:. 1981:. 1955:. 1936:. 1932:. 1913:. 1909:. 1886:. 1878:. 1868:. 1856:. 1852:. 1823:. 1815:. 1807:. 1799:. 1787:. 1783:. 1757:. 1728:. 1718:. 1714:. 1687:50 1654:}} 1650:{{ 1603:}} 1599:{{ 1444:^ 1411:, 1407:, 1340:^ 1275:. 1263:18 1261:. 1257:. 1230:54 1228:. 1224:. 1210:^ 1196:. 1186:. 1174:. 1170:. 1121:. 1111:. 1099:. 1095:. 744:, 738:, 732:, 609:: 344:. 60:, 52:, 3559:) 3552:- 3548:( 3422:) 3418:( 3392:) 3388:( 3384:e 3382:a 3380:r 3378:e 3376:s 3374:n 3372:a 3370:o 3368:l 3366:l 3364:a 3362:G 3349:e 3347:a 3345:h 3343:t 3341:a 3339:n 3337:g 3335:o 3333:e 3331:N 2772:( 2762:e 2755:t 2748:v 2649:: 2542:: 2371:: 2363:: 2327:) 2133:: 2125:: 2101:. 2065:: 2057:: 2033:. 1997:: 1989:: 1966:. 1894:. 1872:: 1864:: 1837:. 1803:: 1795:: 1767:. 1742:. 1722:: 1660:) 1646:. 1609:) 1595:. 1571:. 1555:. 1513:. 1308:. 1283:. 1269:: 1242:. 1236:: 1204:. 1182:: 1155:. 1129:. 1115:: 1107:: 988:) 984:( 970:) 966:( 952:) 948:( 942:) 938:( 922:) 918:( 905:( 899:) 895:( 889:) 885:( 879:) 875:( 849:) 845:( 839:) 835:( 829:) 825:( 819:) 815:( 809:) 805:( 799:) 795:( 789:) 785:( 779:) 775:( 769:) 765:( 243:( 20:)

Index

Darwin's Finches
Large ground finch, Medium ground finch Small tree finch, Green warbler-finch
Large ground finch
Medium ground finch
Small tree finch
Green warbler-finch
Scientific classification
Edit this classification
Eukaryota
Animalia
Chordata
Aves
Passeriformes
Emberizoidea
Thraupidae
Geospiza
Camarhynchus
Platyspiza
Certhidea
Pinaroloxias
species
passerine
subfamily
tribe
tanager
family
true finches
dull-coloured grassquit
the second voyage of the Beagle
Galápagos Islands

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