403:
391:
379:
45:
31:
325:
514:
336:, with adult males measuring 53 to 66 cm (21 to 26 in) in total length, including the 20 to 28 cm (7.9 to 11.0 in) tail, and having an average weight of 1.1 kg (2.4 lb). Females are significantly smaller, measuring 48 to 58 cm (19 to 23 in), including a 17 to 24 cm (6.7 to 9.4 in) tail, and weighing around 0.7 kg (1.5 lb). They have a tapering snout, short legs, and erect ears. They can be distinguished from all other species of quoll by the presence of only four toes, rather than five, on the hind feet, lacking the
1284:
118:
506:
71:
93:
530:) but disease has also been implicated as a potential causative factor of decline. The lack of foxes in Tasmania likely has contributed to the survival of the species there; however, unseasonal weather events and predation by feral cats are thought to have contributed to a possible recent and continuing population declines in Tasmania. The species is currently classified as
250:
484:. Adults also ward off intruders by hissing and making coughing sounds, and also make a sharp shriek that may be an alarm call. If intruders fail to leave quickly, then aggressive action escalates to chasing and wrestling with jaws while standing on their hind legs. Mothers and young have softer calls they use to maintain contact.
468:
Eastern quolls are nocturnal, and spend the day resting in dens, although they may also use natural rock crevices or hollow tree trunks. The dens often consist of no more than a simple, blind-ending tunnel, but are sometimes more complex, including one or more nesting chambers lined with grass. Each
496:
cycle lasts 34 days, although most individuals mate during their first cycle of the year. The female gives birth up to thirty young after a gestation period of 19 to 24 days. Of these, the first to attach themselves to the available teats will be the only survivors. The young remain attached to the
347:
Females possess a relatively shallow fur-lined pouch formed by lateral folds of skin. The pouch becomes enlarged during the breeding season, and includes six to eight teats, which only become elongated and functional if one of the young attaches to them, regressing again after they leave the pouch.
343:
They have a thick coat covered by white spots, that can be either light fawn or near-black, with off-white underparts stretching from the chin to the underside of the tail. Both fawn and black individuals can be born in the same litter, although in surviving populations the former are about three
566:
region of NSW, indicating the species may have survived longer than the generally accepted 1960's decade of its mainland extinction. The NSW National Parks and
Wildlife service reported numerous unconfirmed sightings and the species was reported photographed as recently as 2013 in the Nungatta
612:. Mulligans Flat is a public 485 ha reserve which is surrounded by a predator-proof conservation fence. Founders from the first trial had a survival rate of 28.6%, with the majority of mortalities being associated with males dispersing beyond the predator-proof fence. Adopting an
497:
teat for 60 to 65 days, begin to develop fur at around 51 days, open their eyes at about 79 days, and are fully weaned at 150 to 165 days. They reach sexual maturity in their first year and typically live for 2–3 years, but can live for up to 7 years in captivity.
344:
times more common than the latter. The spots are 5 to 20 mm (0.20 to 0.79 in) in diameter, and are found across the upper body and flanks, from the top of the head to the rump, but, unlike some other species of quoll, do not extend onto the tail.
449:. Within Tasmania, eastern quolls inhabit rainforest, heathland, alpine areas, and scrub below 1,500 m (4,900 ft). However, they prefer dry grassland and forest mosaics, bounded by agricultural land, particularly where pasture grubs are common.
864:
Wilson, Belinda A.; Evans, Maldwyn J.; Batson, William G.; Banks, Sam C.; Gordon, Iain J.; Fletcher, Donald B.; Wimpenny, Claire; Newport, Jenny; Belton, Emily; Rypalski, Annette; Portas, Tim; Manning, Adrian D. (29 June 2020).
1619:
Wilson, Belinda (2021). "Chapter: 6.9 (mammals): Reintroduction of the eastern quoll to
Mulligans Flat Woodland Sanctuary, Australia, using trials, tactics, and adaptive management". In Soorae, Pritpal S. (ed.).
461:. Although the majority of their diet consists of meat, they also eat some vegetable matter, including fruit during the summer, and grass year-round. The eastern quoll is itself prey for Tasmanian devils and
102:
616:
approach, the second and third trials involved only releasing females (preferring those carrying pouch young), which was met with increased survival (76.9% in 2017 and 87.5% in 2018).
402:
457:
The eastern quoll is a solitary predator, hunting at night for its prey of insects, small mammals, birds, and reptiles. They have been known to scavenge food from the much larger
476:
are typically around 35 ha (86 acres) for females, and 44 ha (110 acres) for males, with the latter increasing dramatically during the breeding season. Territories are
1589:
3391:
3196:
644:, and road mortality, no founders survived. A second release of 40 eastern quolls was conducted in 2019, of which none appear to have survived beyond early 2021.
390:
1721:
3261:
378:
3361:
775:
625:
3341:
1481:
3170:
1757:
559:
3209:
867:"Adapting reintroduction tactics in successive trials increases the likelihood of establishment for an endangered carnivore in a fenced sanctuary"
1638:
1003:
571:, Victoria, are likely connected with a nearby wildlife sanctuary, either as direct escapees, or the descendants of escapees from that facility.
3371:
3313:
1690:
3366:
1663:
584:
In 2003, the eastern quoll was reintroduced to a 473ha fox-proof fenced sanctuary at Mt
Rothwell Biodiversity Interpretation Centre at
44:
3386:
3381:
3346:
2053:
30:
941:
694:
605:
2237:
513:
3376:
3235:
1106:
1076:
75:
441:
It likely became functionally extinct across its entire mainland range by the early 1960s, but remains widespread but patchy in
2957:
2948:
2080:
1597:
296:, but were considered extinct on the mainland after 1963. The species has been reintroduced to fox-proof fenced sanctuaries in
324:
257:
Eastern quoll range, not including reintroduced populations in
Victoria, New South Wales and the Australian Capital Territory
3253:
2169:
438:, with a 1923 newspaper article noting its rapid decline and presumed extinction in the area during the preceding ten years.
3214:
2246:
652:
3356:
1750:
609:
301:
3248:
3098:
2866:
2219:
2210:
2160:
1718:
563:
312:
The eastern quoll is a member of the family
Dasyuridae, which includes most carnivorous marsupials. Its species name,
352:
of the male bears an unusual fleshy appendage. The large intestine of eastern quolls is relatively simple, having no
3318:
676:
2785:
2201:
767:
1432:
Cunningham, Calum X.; Aandahl, Zach; Jones, Menna E.; Hamer, Rowena; Johnson, Christopher N. (20 September 2022).
3266:
2731:
2228:
636:
baiting to prepare for the eastern quoll reintroduction. However, due to threats including predation by residual
551:
117:
1146:
1129:
418:
The eastern quoll was formerly found across much of southeastern mainland
Australia, from the eastern coasts of
2334:
1511:
Frankham, Greta J.; Thompson, Sean; Ingleby, Sandy; Soderquist, Todd; Eldridge, Mark D. B. (25 November 2016).
797:
Frankham, Greta J.; Thompson, Sean; Ingleby, Sandy; Soderquist, Todd; Eldridge, Mark D. B. (25 November 2016).
3116:
3076:
2722:
2481:
1743:
991:
469:
individual uses a number of dens, usually no more than five, which it alternates between on different days.
239:
3351:
3038:
2939:
2821:
2379:
1489:
633:
3300:
2930:
2912:
2472:
2448:
2062:
1318:
526:
The eastern quoll likely became extinct on mainland
Australia due to predation by introduced predators (
462:
222:
975:
3157:
3085:
2839:
2812:
2803:
2740:
2704:
2677:
2659:
2430:
2421:
2071:
1878:
1869:
1513:"Does the 'extinct' eastern quoll (Dasyurus viverrinus) persist in Barrington Tops, New South Wales?"
1229:
878:
799:"Does the 'extinct' eastern quoll (Dasyurus viverrinus) persist in Barrington Tops, New South Wales?"
589:
423:
297:
2903:
2758:
2686:
2668:
2352:
2343:
2307:
1639:"Fewer than half of quolls survive first three months after landmark return to Australian mainland"
613:
60:
1025:
Runciman, S.I.C.; et al. (1995). "Central cardiovascular shunts in the perinatal marsupial".
745:
2632:
2439:
1571:
1463:
1276:
1260:
1050:
995:
531:
472:
Eastern quolls are solitary, and tend to avoid one another, but can form loose 'neighbourhoods'.
112:
97:
3305:
3201:
600:
In March 2016, a trial reintroduction of 16 eastern quolls from Mount
Rothwell (Victoria), and
3287:
3134:
1984:
1928:
1919:
1532:
1455:
1414:
1268:
1042:
937:
914:
896:
818:
700:
690:
365:
1393:"Evidence of rapid population decline of the eastern quoll (Dasyurus viverrinus) in Tasmania"
3292:
3090:
2921:
2857:
2776:
2593:
2388:
2370:
2361:
2316:
2184:
2104:
2095:
1905:
1860:
1563:
1524:
1445:
1404:
1349:
1252:
1208:
1177:
1141:
1034:
987:
904:
886:
810:
740:
3103:
1664:"A 'risky operation', a group of dead quolls and a plan for the future of Aussie predators"
684:
3274:
3227:
2584:
2412:
2325:
2270:
2143:
1725:
1197:"Diet overlap and abundance of sympatric dasyurid carnivores: a hypothesis of competition"
629:
458:
427:
419:
357:
882:
2848:
2650:
2575:
2002:
1896:
1767:
1623:
Global conservation translocation perspectives 2021: Case studies from around the globe
909:
866:
585:
481:
435:
369:
179:
505:
3335:
3222:
2794:
2519:
1993:
1719:
Department of
Environment and Conservation Threatened Species profile – eastern quoll
1626:(7 ed.). IUCN SSC Conservation Translocation Specialist Group. pp. 194–199.
1575:
1467:
1213:
1196:
999:
731:
680:
477:
80:
1280:
1054:
722:
624:
In March 2018, a pilot release of 20 captive-bred eastern quolls were released into
364:. An unusual feature of eastern quolls is the presence of an opening connecting the
3139:
2532:
2288:
2020:
672:
641:
547:
446:
333:
1322:
891:
3240:
3183:
3070:
2261:
2011:
1798:
1735:
278:
2513:
2463:
2298:
1820:
1814:
1730:
1621:
1310:
836:
651:
within a 400 ha (990 acres; 4.0 km) fox-proof fenced sanctuary, the
648:
473:
189:
3061:
1691:"Record numbers of eastern quoll joeys born at Barrington Wildlife Sanctuary"
1536:
1459:
1418:
1098:
1068:
900:
822:
546:
The last eastern quoll specimen on the mainland was collected as roadkill in
2894:
2884:
1833:
704:
568:
281:
169:
129:
1391:
Fancourt, Bronwyn A.; Hawkins, Clare E.; Nicol, Stewart C. (24 June 2013).
1336:
Fletcher, T.P. (1985). "Aspects of reproduction in the male eastern quoll,
1311:"Ecology of the eastern quoll Dasyurus viverrinus (Dasyrudae: Marsupialia)"
918:
249:
1551:
1512:
1433:
1392:
1340:(Shaw) (Marsupialia: Dasyuridae), with notes on polyestry in the female".
1272:
1046:
1038:
798:
480:, although scats are distributed randomly, rather than placed at specific
3279:
3126:
3055:
2044:
1792:
1786:
601:
442:
431:
293:
285:
149:
3149:
1264:
3175:
2608:
2566:
2551:
2542:
2403:
2128:
2119:
1887:
1843:
689:(3rd ed.). Baltimore: Johns Hopkins University Press. p. 25.
637:
527:
493:
1353:
1181:
1130:"Niche differentiation among sympatric Australian dasyurid carnivores"
3188:
3111:
3009:
3001:
2989:
1952:
1943:
1780:
555:
361:
353:
316:, indicates it is "ferret-like". There are no recognised subspecies.
159:
139:
3032:
1567:
1528:
1450:
1409:
814:
1256:
562:
in 2016 was reported to have been collected in 1989 or 1990 in the
1967:
512:
504:
368:
of the heart in newborn young, in addition to that connecting the
349:
323:
289:
199:
686:
Mammal
Species of the World: A Taxonomic and Geographic Reference
535:
3162:
3036:
2986:
2510:
1811:
1739:
1434:"Regional patterns of continuing decline of the eastern quoll†"
932:
Godsell, J. (1995). "Eastern Quoll". In Strahan, Ronald (ed.).
558:
on 31 January 1963, however a taxidermied specimen provided to
372:
found in all marsupials. Both openings close after a few days.
774:. Department of Agriculture, Water and the Environment. 2021.
716:
714:
567:
district of NSW. Specimens collected in 2005 and 2008 west of
337:
1147:
10.1644/1545-1542(2000)081<0434:NDASAD>2.0.CO;2
1073:
South Australian Register (Adelaide, SA : 1839 - 1900)
292:. Endemic to Australia, they occur on the island state of
1371:. Sydney: in 'The Mammals of Australia'. pp. 70–71.
1164:
Blackhall, S. (1980). "Diet of the eastern native-cat,
992:
10.1644/1545-1410(2001)677<0001:DV>2.0.CO;2
1105:. Adelaide, South Australia. 12 May 1923. p. 16.
647:
In September 2020 eastern quolls were reintroduced to
632:. The National Park was unfenced, but had undergone
3045:
3000:
2892:
2883:
2606:
2564:
2540:
2531:
2461:
2401:
2296:
2287:
2259:
2182:
2141:
2117:
2093:
2042:
1965:
1941:
1917:
1841:
1832:
1550:Hope, Ben; Bilney, Rohan J.; Peterie, Jess (2021).
1731:The Aussie Eastern Quoll Ark Conservation Project
1159:
1157:
492:The breeding season begins in early winter. The
974:Jones, M. E.; Rose, R. K. (26 December 2001).
509:A black eastern quoll photographed in Tasmania
1751:
768:"Dasyurus viverrinus — Eastern Quoll, Luaner"
746:10.2305/IUCN.UK.2016-1.RLTS.T6296A21947190.en
332:Eastern quolls are about the size of a small
8:
1381:"Mammals" Dorling Kindersley Limited, London
1069:"MAMMALS OF THE NEIGHBOUR HOOD OF ADELAIDE"
969:
967:
965:
963:
961:
959:
957:
955:
953:
430:. The species was formerly abundant around
3033:
2997:
2983:
2889:
2537:
2528:
2507:
2293:
1838:
1829:
1808:
1758:
1744:
1736:
248:
91:
69:
43:
29:
20:
1552:"Targeted survey for the eastern quoll ("
1488:. Australian Museum. 2003. Archived from
1449:
1408:
1212:
1145:
908:
890:
744:
1230:"Body temperatures of Tasmanian devils (
664:
560:NSW National Parks and Wildlife Service
374:
3392:Species endangered by invasive species
721:Burbidge, A.A.; Woinarski, J. (2016).
7:
859:
857:
772:Species Profile and Threats Database
288:), and one of six extant species of
1689:Siossian, Emma (10 November 2022).
1590:"Victorian Eastern Quoll Specimens"
1195:Jones, M.E.; Barmuta, L.A. (1998).
1128:Jones, m.E.; barmuta, L.A. (2000).
732:IUCN Red List of Threatened Species
3362:Extinct mammals of South Australia
1369:Eastern Quoll, Dasyurus viverrinus
1315:The Australian National University
14:
2958:Cracking-clay Pilbara planigale (
2949:Orange-headed Pilbara planigale (
2054:Woolley's three-striped dasyure (
1486:Mammals - Sydney mammals database
1290:from the original on 14 July 2021
1228:Jones, M.E.; et al. (1997).
1109:from the original on 14 July 2021
1079:from the original on 14 July 2021
1006:from the original on 14 July 2021
778:from the original on 9 March 2021
606:Mulligans Flat Woodland Sanctuary
3342:IUCN Red List endangered species
2238:Rory Cooper's false antechinus (
2170:Narrow-striped marsupial shrew (
1214:10.1046/j.1365-2656.1998.00203.x
401:
389:
377:
116:
1168:(Shaw), in southern Tasmania".
1075:. 17 November 1890. p. 3.
356:, and not being divided into a
2081:Tate's three-striped dasyure (
936:. Reed Books. pp. 62–65.
517:A dark morph eastern quoll pup
1:
2867:Lesser hairy-footed dunnart (
2220:Alexandria false antechinus (
2211:Fat-tailed false antechinus (
2161:Red-bellied marsupial shrew (
1342:Australian Journal of Zoology
653:Barrington Wildlife Sanctuary
2786:Little long-tailed dunnart (
2247:Woolley's false antechinus (
2202:Sandstone false antechinus (
1637:Coote, Gavin (6 June 2018).
1170:Australian Wildlife Research
1099:"IS THE NATIVE CAT EXTINCT?"
892:10.1371/journal.pone.0234455
610:Australian Capital Territory
596:Australian Capital Territory
426:, to the mid-north coast of
302:Australian Capital Territory
3372:Mammals of Victoria (state)
2732:Boullanger Island dunnart (
2229:Ningbing false antechinus (
1238:) through an alpine winter"
3408:
3367:Mammals of New South Wales
2335:Yellow-footed antechinus (
3387:Taxa named by George Shaw
3382:Mammals described in 1800
3347:EPBC Act endangered biota
2996:
2982:
2723:Kangaroo Island dunnart (
2527:
2506:
2482:Brush-tailed phascogale (
1828:
1807:
1775:
1201:Journal of Animal Ecology
256:
247:
228:
221:
113:Scientific classification
111:
89:
67:
58:
54:both in Dorset, Tasmania
51:
42:
37:
28:
23:
2940:Narrow-nosed planigale (
2822:Slender-tailed dunnart (
2380:Subtropical antechinus (
934:The Mammals of Australia
683:; Reeder, D. M. (eds.).
414:Distribution and habitat
348:As with all quolls, the
273:, formerly known as the
3377:Marsupials of Australia
2931:New Guinean planigale (
2913:Long-tailed planigale (
2473:Red-tailed phascogale (
2449:Broad-striped dasyure (
2063:Three-striped dasyure (
1367:Godsell, Janet (1995).
1309:Godsell, Janet (1983).
2840:Hairy-footed dunnart (
2813:White-footed dunnart (
2741:Grey-bellied dunnart (
2705:White-tailed dunnart (
2678:Stripe-faced dunnart (
2660:Carpentarian dunnart (
2431:Black-tailed dasyure (
2422:Short-furred dasyure (
1879:Crest-tailed mulgara (
1870:Brush-tailed mulgara (
1594:Where Light Meets Dark
1234:) and eastern quolls (
628:on the south coast of
626:Booderee National Park
518:
510:
329:
3301:Paleobiology Database
2904:Paucident planigale (
2759:Long-tailed dunnart (
2687:Red-cheeked dunnart (
2669:Julia Creek dunnart (
2353:Cinnamon antechinus (
2344:Atherton antechinus (
2308:Tropical antechinus (
1724:16 March 2011 at the
1245:Physiological Zoology
1039:10.1002/ar.1092430109
1027:The Anatomical Record
976:"Dasyurus viverrinus"
677:"Dasyurus viverrinus"
516:
508:
408:black and fawn morphs
327:
2633:Fat-tailed dunnart (
2440:Long-nosed dasyure (
2192:(False antechinuses)
1556:Australian Mammalogy
1517:Australian Mammalogy
1492:on 29 September 2007
1438:Australian Mammalogy
1397:Australian Mammalogy
1232:Sarcophilus harrisii
1134:Journal of Mammalogy
803:Australian Mammalogy
739:: e.T6296A21947190.
328:Eastern quoll skull.
277:) is a medium-sized
16:Species of marsupial
3357:Mammals of Tasmania
3117:dasyurus-viverrinus
3104:Dasyurus_viverrinus
3091:Dasyurus_viverrinus
3077:Dasyurus viverrinus
3047:Dasyurus viverrinus
2804:Gilbert's dunnart (
2072:Wallace's dasyure (
1985:New Guinean quoll (
1929:Little red kaluta (
1338:Dasyurus viverrinus
1236:Dasyurus viverrinus
1166:Dasyurus viverrinus
1014:– via BioOne.
883:2020PLoSO..1534455W
725:Dasyurus viverrinus
614:adaptive management
434:, particularly the
300:in 2003 and to the
270:Dasyurus viverrinus
232:Dasyurus viverrinus
61:Conservation status
2922:Common planigale (
2858:Sandhill dunnart (
2777:Chestnut dunnart (
2734:S. boullangerensis
2594:Southern ningaui (
2389:Dusky antechinus (
2371:Brown antechinus (
2362:Swamp antechinus (
2317:Agile antechinus (
2213:P. macdonnellensis
2151:(Marsupial shrews)
2105:Speckled dasyure (
1906:Northern mulgara (
1861:Southern mulgara (
542:Mainland Australia
519:
511:
422:, through most of
330:
275:eastern native cat
214:D. viverrinus
3329:
3328:
3288:Open Tree of Life
3039:Taxon identifiers
3030:
3029:
3026:
3025:
3022:
3021:
2978:
2977:
2974:
2973:
2970:
2969:
2879:
2878:
2619:
2585:Pilbara ningaui (
2502:
2501:
2498:
2497:
2494:
2493:
2413:Habbema dasyure (
2326:Fawn antechinus (
2283:
2282:
2271:Tasmanian devil (
2195:
2154:
1978:
1854:
1354:10.1071/ZO9850101
1182:10.1071/WR9800191
980:Mammalian Species
604:was conducted at
261:
260:
106:
84:
3399:
3322:
3321:
3309:
3308:
3296:
3295:
3283:
3282:
3270:
3269:
3257:
3256:
3244:
3243:
3231:
3230:
3218:
3217:
3205:
3204:
3192:
3191:
3179:
3178:
3166:
3165:
3153:
3152:
3143:
3142:
3130:
3129:
3120:
3119:
3107:
3106:
3094:
3093:
3081:
3080:
3079:
3066:
3065:
3064:
3034:
2998:
2984:
2890:
2849:Ooldea dunnart (
2651:Kakadu dunnart (
2635:S. crassicaudata
2627:S. crassicaudata
2618:
2617:
2613:
2576:Wongai ningaui (
2538:
2529:
2522:
2508:
2294:
2194:
2193:
2189:
2153:
2152:
2148:
2003:Northern quoll (
1987:D. albopunctatus
1977:
1976:
1972:
1897:Little mulgara (
1853:
1852:
1848:
1839:
1830:
1823:
1809:
1760:
1753:
1746:
1737:
1706:
1705:
1703:
1701:
1686:
1680:
1679:
1677:
1675:
1660:
1654:
1653:
1651:
1649:
1634:
1628:
1627:
1616:
1610:
1609:
1607:
1605:
1600:on 15 March 2012
1596:. Archived from
1586:
1580:
1579:
1547:
1541:
1540:
1508:
1502:
1501:
1499:
1497:
1478:
1472:
1471:
1453:
1429:
1423:
1422:
1412:
1388:
1382:
1379:
1373:
1372:
1364:
1358:
1357:
1333:
1327:
1326:
1306:
1300:
1299:
1297:
1295:
1289:
1242:
1225:
1219:
1218:
1216:
1192:
1186:
1185:
1161:
1152:
1151:
1149:
1125:
1119:
1118:
1116:
1114:
1095:
1089:
1088:
1086:
1084:
1065:
1059:
1058:
1022:
1016:
1015:
1013:
1011:
971:
948:
947:
929:
923:
922:
912:
894:
861:
852:
851:
849:
847:
833:
827:
826:
794:
788:
787:
785:
783:
764:
758:
757:
755:
753:
748:
718:
709:
708:
669:
405:
393:
381:
252:
234:
121:
120:
100:
95:
94:
78:
73:
72:
47:
33:
21:
3407:
3406:
3402:
3401:
3400:
3398:
3397:
3396:
3332:
3331:
3330:
3325:
3317:
3312:
3304:
3299:
3291:
3286:
3278:
3275:Observation.org
3273:
3265:
3260:
3252:
3247:
3239:
3234:
3226:
3221:
3213:
3208:
3200:
3195:
3187:
3182:
3174:
3169:
3161:
3156:
3148:
3146:
3138:
3133:
3125:
3123:
3115:
3110:
3102:
3097:
3089:
3084:
3075:
3074:
3069:
3060:
3059:
3054:
3041:
3031:
3018:
2992:
2966:
2942:P. tenuirostris
2933:P. novaeguineae
2875:
2795:Sooty dunnart (
2761:S. longicaudata
2753:S. longicaudata
2743:S. griseoventer
2717:S. griseoventer
2615:
2614:
2612:
2602:
2560:
2523:
2517:
2490:
2457:
2424:M. longicaudata
2397:
2279:
2255:
2191:
2190:
2188:
2185:Pseudantechinus
2178:
2150:
2149:
2147:
2137:
2113:
2089:
2038:
2030:Eastern quoll (
1994:Western quoll (
1974:
1973:
1971:
1961:
1937:
1913:
1850:
1849:
1847:
1824:
1818:
1803:
1771:
1764:
1726:Wayback Machine
1715:
1710:
1709:
1699:
1697:
1688:
1687:
1683:
1673:
1671:
1662:
1661:
1657:
1647:
1645:
1636:
1635:
1631:
1618:
1617:
1613:
1603:
1601:
1588:
1587:
1583:
1568:10.1071/AM20060
1549:
1548:
1544:
1529:10.1071/AM16029
1510:
1509:
1505:
1495:
1493:
1482:"Eastern Quoll"
1480:
1479:
1475:
1451:10.1071/AM22010
1431:
1430:
1426:
1410:10.1071/AM13004
1390:
1389:
1385:
1380:
1376:
1366:
1365:
1361:
1335:
1334:
1330:
1308:
1307:
1303:
1293:
1291:
1287:
1240:
1227:
1226:
1222:
1194:
1193:
1189:
1163:
1162:
1155:
1127:
1126:
1122:
1112:
1110:
1097:
1096:
1092:
1082:
1080:
1067:
1066:
1062:
1024:
1023:
1019:
1009:
1007:
973:
972:
951:
944:
931:
930:
926:
877:(6): e0234455.
863:
862:
855:
845:
843:
837:"Eastern Quoll"
835:
834:
830:
815:10.1071/AM16029
796:
795:
791:
781:
779:
766:
765:
761:
751:
749:
720:
719:
712:
697:
671:
670:
666:
661:
630:New South Wales
622:
620:New South Wales
598:
582:
577:
575:Reintroductions
544:
524:
503:
490:
459:Tasmanian devil
455:
428:New South Wales
420:South Australia
416:
409:
406:
397:
394:
385:
382:
322:
310:
243:
236:
230:
217:
115:
107:
96:
92:
85:
74:
70:
63:
53:
17:
12:
11:
5:
3405:
3403:
3395:
3394:
3389:
3384:
3379:
3374:
3369:
3364:
3359:
3354:
3349:
3344:
3334:
3333:
3327:
3326:
3324:
3323:
3310:
3297:
3284:
3271:
3258:
3245:
3232:
3219:
3206:
3193:
3180:
3167:
3154:
3144:
3131:
3121:
3108:
3095:
3082:
3067:
3051:
3049:
3043:
3042:
3037:
3028:
3027:
3024:
3023:
3020:
3019:
3017:
3016:
3006:
3004:
2994:
2993:
2987:
2980:
2979:
2976:
2975:
2972:
2971:
2968:
2967:
2965:
2964:
2955:
2946:
2937:
2928:
2919:
2910:
2900:
2898:
2887:
2881:
2880:
2877:
2876:
2874:
2873:
2864:
2860:S. psammophila
2855:
2846:
2834:S. psammophila
2829:
2828:
2819:
2810:
2801:
2792:
2783:
2766:
2765:
2748:
2747:
2738:
2729:
2712:
2711:
2694:
2693:
2684:
2675:
2666:
2657:
2640:
2639:
2622:
2620:
2604:
2603:
2601:
2600:
2591:
2582:
2572:
2570:
2562:
2561:
2559:
2558:
2548:
2546:
2535:
2525:
2524:
2511:
2504:
2503:
2500:
2499:
2496:
2495:
2492:
2491:
2489:
2488:
2479:
2469:
2467:
2459:
2458:
2456:
2455:
2451:M. rothschildi
2446:
2437:
2428:
2419:
2409:
2407:
2399:
2398:
2396:
2395:
2386:
2382:A. subtropicus
2377:
2368:
2359:
2350:
2341:
2332:
2323:
2314:
2304:
2302:
2291:
2285:
2284:
2281:
2280:
2278:
2277:
2267:
2265:
2257:
2256:
2254:
2253:
2244:
2235:
2226:
2217:
2208:
2198:
2196:
2180:
2179:
2177:
2176:
2167:
2157:
2155:
2139:
2138:
2136:
2135:
2125:
2123:
2115:
2114:
2112:
2111:
2101:
2099:
2091:
2090:
2088:
2087:
2078:
2069:
2060:
2050:
2048:
2040:
2039:
2037:
2036:
2027:
2021:Bronze quoll (
2018:
2009:
2000:
1991:
1981:
1979:
1963:
1962:
1960:
1959:
1949:
1947:
1939:
1938:
1936:
1935:
1925:
1923:
1915:
1914:
1912:
1911:
1903:
1894:
1885:
1881:D. cristicauda
1876:
1867:
1857:
1855:
1836:
1826:
1825:
1812:
1805:
1804:
1802:
1801:
1795:
1789:
1783:
1776:
1773:
1772:
1768:Dasyuromorphia
1765:
1763:
1762:
1755:
1748:
1740:
1734:
1733:
1728:
1714:
1713:External links
1711:
1708:
1707:
1681:
1655:
1629:
1611:
1581:
1542:
1523:(2): 243–247.
1503:
1473:
1444:(2): 151–159.
1424:
1403:(2): 195–205.
1383:
1374:
1359:
1348:(2): 101–110.
1328:
1301:
1257:10.1086/639541
1220:
1207:(3): 410–421.
1187:
1176:(2): 191–197.
1153:
1140:(2): 434–447.
1120:
1103:The Advertiser
1090:
1060:
1017:
949:
942:
924:
853:
828:
809:(2): 243–247.
789:
759:
710:
695:
663:
662:
660:
657:
621:
618:
597:
594:
586:Mount Rothwell
581:
578:
576:
573:
543:
540:
523:
520:
502:
499:
489:
486:
454:
451:
436:Adelaide Hills
415:
412:
411:
410:
407:
400:
398:
395:
388:
386:
383:
376:
321:
318:
309:
306:
259:
258:
254:
253:
245:
244:
237:
226:
225:
219:
218:
211:
209:
205:
204:
197:
193:
192:
187:
183:
182:
180:Dasyuromorphia
177:
173:
172:
167:
163:
162:
157:
153:
152:
147:
143:
142:
137:
133:
132:
127:
123:
122:
109:
108:
90:
87:
86:
68:
65:
64:
59:
56:
55:
49:
48:
40:
39:
35:
34:
26:
25:
24:Eastern quoll
15:
13:
10:
9:
6:
4:
3:
2:
3404:
3393:
3390:
3388:
3385:
3383:
3380:
3378:
3375:
3373:
3370:
3368:
3365:
3363:
3360:
3358:
3355:
3353:
3352:Dasyuromorphs
3350:
3348:
3345:
3343:
3340:
3339:
3337:
3320:
3315:
3311:
3307:
3302:
3298:
3294:
3289:
3285:
3281:
3276:
3272:
3268:
3263:
3259:
3255:
3250:
3246:
3242:
3237:
3233:
3229:
3224:
3220:
3216:
3211:
3207:
3203:
3198:
3194:
3190:
3185:
3181:
3177:
3172:
3168:
3164:
3159:
3155:
3151:
3145:
3141:
3136:
3132:
3128:
3122:
3118:
3113:
3109:
3105:
3100:
3096:
3092:
3087:
3083:
3078:
3072:
3068:
3063:
3057:
3053:
3052:
3050:
3048:
3044:
3040:
3035:
3015:
3013:
3008:
3007:
3005:
3003:
2999:
2995:
2991:
2990:Myrmecobiidae
2985:
2981:
2963:
2961:
2956:
2954:
2952:
2947:
2945:
2943:
2938:
2936:
2934:
2929:
2927:
2925:
2920:
2918:
2916:
2911:
2909:
2907:
2902:
2901:
2899:
2897:
2896:
2891:
2888:
2886:
2882:
2872:
2870:
2865:
2863:
2861:
2856:
2854:
2852:
2847:
2845:
2843:
2837:
2836:species-group
2835:
2831:
2830:
2827:
2825:
2820:
2818:
2816:
2811:
2809:
2807:
2802:
2800:
2798:
2797:S. fulginosus
2793:
2791:
2789:
2784:
2782:
2780:
2774:
2773:species-group
2772:
2768:
2767:
2764:
2762:
2756:
2755:species-group
2754:
2750:
2749:
2746:
2744:
2739:
2737:
2735:
2730:
2728:
2726:
2720:
2719:species-group
2718:
2714:
2713:
2710:
2708:
2707:S. granulipes
2702:
2701:species-group
2700:
2699:S. granulipes
2696:
2695:
2692:
2690:
2685:
2683:
2681:
2676:
2674:
2672:
2667:
2665:
2663:
2658:
2656:
2654:
2648:
2647:species-group
2646:
2642:
2641:
2638:
2636:
2630:
2629:species-group
2628:
2624:
2623:
2621:
2611:
2610:
2605:
2599:
2597:
2592:
2590:
2588:
2583:
2581:
2579:
2574:
2573:
2571:
2569:
2568:
2563:
2557:
2555:
2550:
2549:
2547:
2545:
2544:
2539:
2536:
2534:
2530:
2526:
2521:
2520:Sminthopsinae
2515:
2509:
2505:
2487:
2485:
2480:
2478:
2476:
2471:
2470:
2468:
2466:
2465:
2460:
2454:
2452:
2447:
2445:
2443:
2438:
2436:
2434:
2429:
2427:
2425:
2420:
2418:
2416:
2411:
2410:
2408:
2406:
2405:
2400:
2394:
2392:
2391:A. swainsonii
2387:
2385:
2383:
2378:
2376:
2374:
2369:
2367:
2365:
2360:
2358:
2356:
2351:
2349:
2347:
2342:
2340:
2338:
2333:
2331:
2329:
2324:
2322:
2320:
2315:
2313:
2311:
2306:
2305:
2303:
2301:
2300:
2295:
2292:
2290:
2286:
2276:
2274:
2269:
2268:
2266:
2264:
2263:
2258:
2252:
2250:
2245:
2243:
2241:
2236:
2234:
2232:
2227:
2225:
2223:
2218:
2216:
2214:
2209:
2207:
2205:
2200:
2199:
2197:
2187:
2186:
2181:
2175:
2173:
2168:
2166:
2164:
2159:
2158:
2156:
2146:
2145:
2144:Phascolosorex
2140:
2134:
2132:
2127:
2126:
2124:
2122:
2121:
2120:Parantechinus
2116:
2110:
2108:
2103:
2102:
2100:
2098:
2097:
2096:Neophascogale
2092:
2086:
2084:
2079:
2077:
2075:
2070:
2068:
2066:
2061:
2059:
2057:
2052:
2051:
2049:
2047:
2046:
2041:
2035:
2033:
2032:D. viverrinus
2028:
2026:
2024:
2019:
2017:
2015:
2012:Tiger quoll (
2010:
2008:
2006:
2005:D. hallucatus
2001:
1999:
1997:
1992:
1990:
1988:
1983:
1982:
1980:
1970:
1969:
1964:
1958:
1956:
1951:
1950:
1948:
1946:
1945:
1940:
1934:
1932:
1931:D. rosamondae
1927:
1926:
1924:
1922:
1921:
1916:
1910:
1909:
1904:
1902:
1900:
1895:
1893:
1891:
1886:
1884:
1882:
1877:
1875:
1873:
1868:
1866:
1864:
1859:
1858:
1856:
1846:
1845:
1840:
1837:
1835:
1831:
1827:
1822:
1816:
1810:
1806:
1800:
1796:
1794:
1790:
1788:
1784:
1782:
1778:
1777:
1774:
1769:
1761:
1756:
1754:
1749:
1747:
1742:
1741:
1738:
1732:
1729:
1727:
1723:
1720:
1717:
1716:
1712:
1696:
1692:
1685:
1682:
1670:. 21 May 2021
1669:
1665:
1659:
1656:
1644:
1640:
1633:
1630:
1625:
1624:
1615:
1612:
1599:
1595:
1591:
1585:
1582:
1577:
1573:
1569:
1565:
1561:
1557:
1553:
1546:
1543:
1538:
1534:
1530:
1526:
1522:
1518:
1514:
1507:
1504:
1491:
1487:
1483:
1477:
1474:
1469:
1465:
1461:
1457:
1452:
1447:
1443:
1439:
1435:
1428:
1425:
1420:
1416:
1411:
1406:
1402:
1398:
1394:
1387:
1384:
1378:
1375:
1370:
1363:
1360:
1355:
1351:
1347:
1343:
1339:
1332:
1329:
1324:
1320:
1316:
1312:
1305:
1302:
1286:
1282:
1278:
1274:
1270:
1266:
1262:
1258:
1254:
1250:
1246:
1239:
1237:
1233:
1224:
1221:
1215:
1210:
1206:
1202:
1198:
1191:
1188:
1183:
1179:
1175:
1171:
1167:
1160:
1158:
1154:
1148:
1143:
1139:
1135:
1131:
1124:
1121:
1108:
1104:
1100:
1094:
1091:
1078:
1074:
1070:
1064:
1061:
1056:
1052:
1048:
1044:
1040:
1036:
1032:
1028:
1021:
1018:
1005:
1001:
997:
993:
989:
985:
981:
977:
970:
968:
966:
964:
962:
960:
958:
956:
954:
950:
945:
943:9781877069253
939:
935:
928:
925:
920:
916:
911:
906:
902:
898:
893:
888:
884:
880:
876:
872:
868:
860:
858:
854:
842:
838:
832:
829:
824:
820:
816:
812:
808:
804:
800:
793:
790:
777:
773:
769:
763:
760:
747:
742:
738:
734:
733:
728:
726:
717:
715:
711:
706:
702:
698:
696:0-801-88221-4
692:
688:
687:
682:
681:Wilson, D. E.
678:
674:
673:Groves, C. P.
668:
665:
658:
656:
654:
650:
645:
643:
642:domestic dogs
639:
635:
631:
627:
619:
617:
615:
611:
607:
603:
595:
593:
591:
587:
579:
574:
572:
570:
565:
561:
557:
553:
549:
541:
539:
537:
533:
529:
521:
515:
507:
500:
498:
495:
487:
485:
483:
479:
475:
470:
466:
464:
460:
452:
450:
448:
444:
439:
437:
433:
429:
425:
421:
413:
404:
399:
392:
387:
380:
375:
373:
371:
367:
363:
359:
355:
351:
345:
341:
339:
335:
326:
319:
317:
315:
307:
305:
303:
299:
295:
291:
287:
283:
280:
276:
272:
271:
266:
265:eastern quoll
255:
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223:Binomial name
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3012:M. fasciatus
3011:
2959:
2951:P. kendricki
2950:
2941:
2932:
2923:
2914:
2905:
2893:
2869:S. youngsoni
2868:
2859:
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2796:
2788:S. dolichura
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2689:S. virginiae
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2587:N. timealeyi
2586:
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2543:Antechinomys
2541:
2533:Sminthopsini
2484:P. tapoatafa
2483:
2474:
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2450:
2441:
2433:M. melanurus
2432:
2423:
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2289:Phascogalini
2272:
2260:
2249:P. woolleyae
2248:
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2023:D. spartacus
2022:
2014:D. maculatus
2013:
2004:
1996:D. geoffroii
1995:
1986:
1966:
1954:
1942:
1930:
1918:
1908:D. woolleyae
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1698:. Retrieved
1694:
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1672:. Retrieved
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1602:. Retrieved
1598:the original
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548:Nielsen Park
545:
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501:Conservation
491:
488:Reproduction
478:scent marked
471:
467:
456:
447:Bruny Island
440:
417:
346:
342:
334:domestic cat
331:
313:
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274:
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264:
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231:
229:
213:
212:
200:
166:Infraclass:
18:
3184:iNaturalist
3071:Wikispecies
3002:Myrmecobius
2924:P. maculata
2885:Planigalini
2842:S. hirtipes
2815:S. leucopus
2806:S. gilberti
2680:S. macroura
2671:S. douglasi
2645:S. macroura
2609:Sminthopsis
2373:A. stuartii
2337:A. flavipes
2273:S. harrisii
2262:Sarcophilus
2231:P. ningbing
2172:P. dorsalis
2131:P. apicalis
2107:N. lorentzi
2074:M. wallacii
1944:Dasyuroides
1799:Marsupialia
1797:Infraclass
1700:11 November
1648:11 November
1562:: 153–157.
1496:12 February
1113:5 September
1083:8 September
752:19 November
474:Home ranges
463:masked owls
396:black morph
320:Description
279:carnivorous
170:Marsupialia
52:black morph
38:fawn morph
3336:Categories
2915:P. ingrami
2779:S. archeri
2725:S. aitkeni
2662:S. butleri
2616:(Dunnarts)
2596:N. yvonnae
2554:A. laniger
2518:subfamily
2514:Dasyuridae
2464:Phascogale
2415:M. habbema
2364:A. minimus
2346:A. godmani
2310:A. adustus
2299:Antechinus
2222:P. mimulus
2204:P. bilarni
2083:M. wavicus
2056:M. leucura
1920:Dasykaluta
1899:D. marlowi
1890:D. hilleri
1863:D. archeri
1851:(Mulgaras)
1844:Dasycercus
1821:Dasyurinae
1819:subfamily
1815:Dasyuridae
1674:11 October
1323:2598644376
846:11 October
841:Mtrothwell
659:References
649:Aussie Ark
564:Gloucester
532:Endangered
384:fawn morph
366:ventricles
314:viverrinus
190:Dasyuridae
98:Endangered
76:Endangered
2960:P. tealei
2906:P. gilesi
2895:Planigale
2851:S. ooldea
2824:S. murina
2771:S. murina
2552:Kultarr (
2475:P. calura
2328:A. bellus
2319:A. agilis
2163:P. doriae
2129:Dibbler (
1955:D. byrnei
1888:Ampurta (
1872:D. blythi
1834:Dasyurini
1576:234852178
1537:1836-7402
1468:252391044
1460:1836-7402
1419:1836-7402
1294:30 August
1000:198968816
901:1932-6203
823:1836-7402
638:red foxes
569:Melbourne
453:Behaviour
304:in 2016.
282:marsupial
208:Species:
136:Kingdom:
130:Eukaryota
3254:10800025
3202:10218946
3124:BioLib:
3056:Wikidata
3010:Numbat (
2653:S. bindi
2578:N. ridei
2240:P. roryi
2065:M. melas
2045:Myoictis
1975:(Quolls)
1968:Dasyurus
1953:Kowari (
1793:Mammalia
1787:Chordata
1781:Animalia
1779:Kingdom
1722:Archived
1695:ABC News
1668:ABC News
1643:ABC News
1604:15 March
1319:ProQuest
1285:Archived
1281:29736671
1265:30164283
1107:Archived
1077:Archived
1055:34243728
1004:Archived
919:32598368
871:PLOS ONE
776:Archived
705:62265494
675:(2005).
602:Tasmania
590:Victoria
580:Victoria
552:Vaucluse
522:Tasmania
482:latrines
443:Tasmania
432:Adelaide
424:Victoria
308:Taxonomy
298:Victoria
294:Tasmania
286:dasyurid
201:Dasyurus
186:Family:
160:Mammalia
150:Chordata
146:Phylum:
140:Animalia
126:Domain:
103:EPBC Act
81:IUCN 3.1
3241:1000152
3176:2435357
3062:Q194516
2988:Family
2567:Ningaui
2512:Family
2442:M. naso
2404:Murexia
1813:Family
1785:Phylum
1770:species
1766:Extant
1273:9231376
1047:8540634
1010:14 July
986:: 1–9.
910:7323978
879:Bibcode
782:14 July
634:red fox
608:in the
534:by the
528:red fox
494:oestrus
242:, 1800)
196:Genus:
176:Order:
156:Class:
101: (
79: (
3306:235548
3293:393372
3215:552611
3163:323728
3147:ECOS:
3112:ARKive
2355:A. leo
1791:Class
1574:
1535:
1466:
1458:
1417:
1321:
1279:
1271:
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998:
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917:
907:
899:
821:
703:
693:
556:Sydney
362:rectum
354:caecum
338:hallux
290:quolls
3314:SPRAT
3280:83050
3197:IRMNG
3189:40167
3140:6CF7M
3127:33318
1572:S2CID
1464:S2CID
1288:(PDF)
1277:S2CID
1261:JSTOR
1241:(PDF)
1051:S2CID
996:S2CID
679:. In
370:atria
358:colon
350:penis
3267:9279
3262:NCBI
3228:6296
3223:IUCN
3210:ITIS
3171:GBIF
3150:7961
1702:2022
1676:2022
1650:2022
1606:2018
1533:ISSN
1498:2007
1456:ISSN
1415:ISSN
1296:2020
1269:PMID
1115:2020
1085:2020
1043:PMID
1012:2021
938:ISBN
915:PMID
897:ISSN
848:2022
819:ISSN
784:2021
754:2021
737:2016
701:OCLC
691:ISBN
536:IUCN
445:and
360:and
263:The
240:Shaw
3319:333
3249:MSW
3236:MDD
3158:EoL
3135:CoL
3099:AFD
3086:ADW
1564:doi
1525:doi
1446:doi
1405:doi
1350:doi
1253:doi
1209:doi
1178:doi
1142:doi
1035:doi
1031:243
988:doi
984:677
905:PMC
887:doi
811:doi
741:doi
588:in
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