211:
68:
43:
682:
653:. Head and body length is 138 to 164 mm (5.4 to 6.5 in). In Osgood's original two specimens, an old female and an adult female, tail length is 188 and 180 mm (7.4 and 7.1 in), respectively; hindfoot length is 30 and 30 mm (1.2 and 1.2 in); and greatest skull length is 37 and 34.7 mm (1.46 and 1.37 in).
850:, the front cusps on the upper and lower first molar, are not divided into smaller outer and inner cusps. Small accessory roots are absent from the molars, so that each of the three upper molars has two roots on the outer side and one on the inner side and each of the lower molars has one root at the front and one at the back.
862:
is confined to a small area in central Peru, at an altitude of 760 to 2,100 m (2,490 to 6,890 ft), but the species may range more widely. It occurs in forest in the dry lowlands of the upper parts of the basin of the
755:(depression). The bony palate is relatively short, with the mesopterygoid fossa extending forward to the end of the molar row or even between the third molars. The roof of the fossa is perforated by large
645:, small structures resembling scales that cover the soles of the hindfeet in many oryzomyines, are well developed. The claw of the first digit extends nearly to the end of the first
834:(low-crowned). On the upper first and second molar, the outer and inner valleys between the cusps and crests do not interpenetrate. Many accessory crests are present, including the
806:, which anchor some of the chewing muscles, usually join into a single crest at a point below the first molar and do not extend forward beyond the molar. There is no distinct
570:
is now one of about 28 genera in the tribe
Oryzomyini, which includes well over a hundred species distributed mainly in South America, including nearby islands such as the
740:, especially in old animals. The parietal bones form part of the roof of the braincase and, unlike in some other rice rats, also extend to the sides of the braincase. The
3304:
503:, the first upper molar has an additional small root at the outer (labial) side; the first lower molar has additional small roots; and the second upper molar has the
560:, was described from southeastern Brazil, and phylogenetic analysis of morphological and molecular data suggested that this animal is the closest known relative of
1417:"A new genus and species of rodent from the Brazilian Atlantic Forest (Rodentia: Cricetidae: Sigmodontinae: Oryzomyini), with comments on oryzomyine biogeography"
3274:
3218:
582:, which encompasses hundreds of species found across South America and into southern North America. Sigmodontinae itself is the largest subfamily of the family
408:) "for convenience". Its relationships remained obscure ever afterward and it was never assigned to any of the several groups of species recognized within
649:
of the second toe and the claw of the fifth toe extends slightly beyond the first phalanx of the fourth toe. As in most oryzomyines, the female has eight
3153:
1493:
495:(a crest at the back) is present on the third upper molar. Two other molecular synapomorphies supported the clade of all members of clade D except
3179:
779:. The condition of various grooves and foramina of the skull indicates that the pattern of the arterial circulation of the head is derived. The
3299:
802:, an opening in the mandible just before the first molar, opens to the outside, not upwards as in a few other oryzomyines. The upper and lower
751:
The incisive foramina are very long, extending well between the molars. The posterolateral palatal pits are well-developed and recessed into a
1451:
Phylogenetic relationships of oryzomyine rodents (Muroidea: Sigmodontinae): separate and combined analyses of morphological and molecular data
1354:
732:
bones, which form the front and back parts of the arch, respectively, do not overlap when seen from the side. The narrowest part of the
518:) group, but instead were found at various positions across the oryzomyine tree, and he suggested that most of these species, including
3294:
736:(located between the eyes) is to the front and the region's margins exhibit strong beading. Various crests develop on the rounded
3289:
3205:
623:). The fur is grayish above and lighter below, where the hairs are gray at the bases but white at the tips. The external ears (
1346:
3184:
522:, should be placed in new genera. Later in 2006, Weksler and others described ten new genera for species formerly placed in
936:
Weksler et al., 2006, p. 10. Percequillo et al., 2011, p. 367, write that the fossa does not reach between the molars in
1486:
927:) as having overlapping squamosals and maxillaries (see character state definitions for character 30, p. 32).
67:
472:
459:) molecular characters and by seven morphological synapomorphies—the tail has a different color above and below; the
1332:
772:
3223:
2692:
543:
3023:
2699:
556:
160:
3279:
3016:
1805:
1690:
1569:
776:
756:
476:
3131:
3045:
2972:
2734:
2678:
2664:
2657:
2650:
2449:
2442:
2369:
2148:
1912:
1479:
436:
3093:
2727:
2685:
2435:
2320:
1962:
1890:
1576:
404:
480:
3052:
2720:
2706:
2636:
2397:
2376:
2327:
2090:
2052:
1883:
1869:
1819:
1784:
1371:
919:
Weksler et al., 2006, p. 10; Percequillo et al., 2011, p. 388. Weksler, 2006, table 5, scores
780:
200:
183:
384:
and M.P. Anderson in 1912. The next year, Osgood described these animals as a new species in the genus
803:
783:, an opening at the back of the skull determined by the shape of the squamosal bone, is large and the
2965:
2943:
2741:
2713:
2671:
2643:
2629:
2600:
2404:
2334:
2155:
2097:
2044:
1976:
1919:
1876:
1862:
1855:
1848:
1812:
1798:
1711:
1683:
571:
455:
to a clade containing all the other species of clade D. Clade D was supported by two shared derived (
432:
332:
A large, long-tailed rice rat, with head and body length of 138 to 164 mm (5.4 to 6.5 in),
156:
689:
3284:
2844:
2815:
2801:
2793:
2749:
2456:
2427:
2390:
2060:
1926:
1841:
1697:
991:
896:
733:
365:
357:
32:
864:
787:
is perforated by a fenestra (opening). The squamosal lacks a suspensory process that contacts the
326:
2607:
2516:
2502:
2362:
2298:
2036:
1990:
1955:
1791:
1676:
741:
716:, is present. The plate's back margin is level with the front of the first upper molar. A strong
637:
223:
62:
1406:
301:
in the strict sense or to any other oryzomyine then known, so that it is now placed in its own
3244:
3140:
2936:
2586:
2537:
2383:
2355:
2348:
2341:
2284:
2270:
2248:
2241:
2104:
1969:
1948:
1704:
1438:
1360:
1350:
760:
712:
extend about as far back as the nasals. The zygomatic notch, an extension at the front of the
658:
3249:
1375:
1340:
2830:
2822:
2593:
2558:
2551:
2530:
2277:
2227:
2220:
2213:
2206:
2199:
2126:
2116:
2068:
1718:
1428:
1401:
986:
807:
464:
424:
3197:
210:
3231:
2808:
2579:
2572:
2544:
2523:
2509:
2291:
2234:
1983:
1471:
880:
792:
713:
666:
662:
624:
627:) are short and the tail is dark above and light below. The hindfeet have well-developed
1082:
Weksler, 2006, table 2, p. 130; Musser and
Carleton, 2005, pp. 1144, 1158
3063:
3006:
2836:
900:
892:
799:
788:
745:
729:
725:
721:
381:
361:
280:
47:
3268:
3192:
2955:
2890:
2880:
1630:
1620:
1548:
1433:
1416:
1392:
1336:
977:
968:
823:
705:
579:
547:
460:
139:
52:
1383:
3145:
3035:
2619:
2138:
1774:
1654:
1643:
1608:
1600:
1589:
879:
is unusual. While there are some similar cases of relationships between Andean and
868:
784:
709:
670:
646:
488:
456:
452:
337:
752:
744:
at the back of the braincase is narrow and wedge-shaped, so that the parietal and
3210:
3166:
3125:
2771:
2761:
2565:
2416:
1831:
1740:
1730:
1415:
Percequillo, Alexandre R.; Weksler, Marcelo; Costa, Leonora P. (February 2011).
847:
764:
628:
538:. In reference to its "isolated distribution", they incorporated the Greek word
492:
313:
3116:
3074:
2994:
2984:
2868:
2857:
2480:
2469:
2310:
1902:
1666:
1559:
1542:
1503:
1328:
843:
839:
831:
717:
701:
599:
583:
504:
448:
420:
416:
294:
276:
272:
129:
1442:
613:
is a large, long-tailed rice rat that in color resembles some North
American
2080:
2025:
1938:
1364:
737:
515:
345:
79:
3236:
3110:
2926:
2914:
2903:
2783:
2260:
2189:
2177:
2167:
2012:
2002:
1524:
835:
827:
575:
484:
394:. Osgood wrote that he was unable to find any species closely related to
386:
285:
99:
3158:
2492:
768:
642:
632:
619:
614:
595:
591:
499:, coupled with three morphological traits—in these species, but not in
578:. Oryzomyini is one of several tribes recognized within the subfamily
554:
may in fact include more than one species. In 2011, a new oryzomyine,
483:, the gap behind the end of the palate) are large; the pattern of the
3171:
1536:
1530:
1518:
650:
468:
349:
341:
268:
119:
109:
89:
3087:
669:
vertebrae; the presence of 12 thoracic vertebrae is a putative
635:
margins and between all of the digits, a character shared only with
475:(perforations of the palate near the third molars) are complex; the
681:
344:. The rostrum (front part of the skull) is long and robust and the
1461:
1450:
883:
animals, these involve inhabitants of humid forests in the Andes;
697:
680:
507:(one of the valleys between the cusps and crests) divided in two.
302:
1342:
Mammal
Species of the World: A Taxonomic and Geographic Reference
1462:
Ten new genera of oryzomyine rodents (Cricetidae: Sigmodontinae)
587:
440:
353:
3091:
1475:
1378:. Field Museum of Natural History, Zoölogical Series 10:93–100.
1270:
Weksler, 2006, pp. 41–42; Weksler et al., 2006, p. 11
1234:
Weksler, 2006, pp. 38–39; Weksler et al., 2006, p. 10
962:
960:
958:
956:
1306:
Musser and
Carleton, 2005, p. 1153; Pacheco et al., 2008
1012:
Pacheco et al., 2008; Musser and
Carleton, 2005, p. 1153
329:
in central Peru, but may yet contain more than one species.
1288:
Weksler, 2006, p. 43; Weksler et al., 2006, p. 11
1261:
Weksler, 2006, p. 44; Weksler et al., 2006, p. 11
891:
is so poorly known, the 2008 IUCN Red List assesses it as "
1382:
Pacheco, V.; Zeballos, H.; Vivar, E.; Weksler, M. (2008).
967:
Pacheco, V.; Zeballos, H.; Vivar, E.; Weksler, M. (2019).
325:
has a limited distribution in the dry upper valley of the
16:
Rodent species in the family
Cricetidae from central Peru
830:(with the cusps higher than the connecting crests) and
336:
has gray fur and short ears. There are well-developed
451:, one of four large groups within Oryzomyini, as the
317:, named in 2011, is probably the closest relative of
1460:
Weksler, M., Percequillo, A.R. and Voss, R.S. 2006.
867:, east of the main mountain range of the Andes. The
279:. Discovered in 1912 and first described in 1913 by
3100:
3062:
3033:
3004:
2982:
2953:
2924:
2900:
2878:
2854:
2781:
2759:
2617:
2490:
2466:
2414:
2308:
2258:
2187:
2165:
2136:
2114:
2078:
2022:
2000:
1936:
1900:
1829:
1772:
1750:
1728:
1664:
1640:
1618:
1586:
1557:
700:, the rostrum (front part) is long and robust. The
1455:Bulletin of the American Museum of Natural History
297:analysis found that it was not closely related to
415:In 2006, Marcelo Weksler published a large-scale
704:are short, not extending further back than the
364:"; it is poorly known but may be threatened by
992:10.2305/IUCN.UK.2019-1.RLTS.T15609A22338308.en
887:, in contrast, lives in an arid area. Because
871:pattern indicated by the relationship between
1487:
899:for cattle pasture and is not known from any
602:, all mainly from Eurasia and North America.
8:
1109:Weksler, 2006, pp. 75, 77, fig. 42
471:) extend back between the first molars; the
340:of hair on the hindfeet. Females have eight
1407:10.2305/IUCN.UK.2008.RLTS.T15609A4891730.en
1207:Weksler, 2006, pp. 27–28, table 5
3088:
1494:
1480:
1472:
814:shares with only a few other oryzomyines.
209:
41:
20:
1432:
1421:Zoological Journal of the Linnean Society
1405:
990:
510:In Weksler's analysis, species placed in
3305:Taxa named by Alexandre Reis Percequillo
1131:
1129:
1127:
1060:
1058:
1252:Weksler, 2006, p. 41, table 5
1171:Weksler, 2006, p. 17, table 5
1117:
1115:
1064:Musser and Carleton, 2005, p. 1153
1048:
1046:
1044:
1042:
1040:
1038:
1036:
1034:
952:
912:
795:, a defining character of oryzomyines.
463:extends to the side of the skull; the
1024:
1022:
1020:
1018:
7:
3275:IUCN Red List data deficient species
534:; thus, the species is now known as
245:Weksler, Percequillo, and Voss, 2006
1393:IUCN Red List of Threatened Species
978:IUCN Red List of Threatened Species
763:is present; this extension of the
14:
688:occurs in the basin of the upper
586:, other members of which include
550:, citing Pacheco, commented that
1434:10.1111/j.1096-3642.2010.00643.x
1315:Percequillo et al., 2011, p. 379
1297:Percequillo et al., 2011, p. 378
1144:Percequillo et al., 2011, p. 372
1135:Weksler et al., 2006, p. 11
1052:Weksler et al., 2006, p. 10
66:
1121:Weksler et al., 2006, p. 1
1091:Weksler, 2006, figs. 34–40
1347:Johns Hopkins University Press
810:of the lower incisor, a trait
773:masticatory–buccinator foramen
443:gene. In all of his analyses,
283:, it was originally placed in
1:
3300:Taxa named by Marcelo Weksler
1279:Weksler, 2006, pp. 43–44
771:(openings) in the skull, the
1073:Osgood, 1913, pp. 97–98
631:(patches of hair) along the
1189:Steppan, 1995, table 5
1180:Weksler, 2006, table 8
487:circulation in the head is
473:posterolateral palatal pits
376:The first two specimens of
3321:
1331:; Carleton, M. D. (2005).
1100:Weksler, 2006, p. 130
3295:Mammals described in 1913
1513:
1466:American Museum Novitates
1243:Weksler, 2006, p. 40
1225:Weksler, 2006, p. 31
1216:Weksler, 2006, p. 30
1198:Weksler, 2006, p. 52
1162:Weksler, 2006, p. 24
1153:Musser and Carleton, 2005
777:foramen ovale accessorium
724:(cheekbone), so that the
557:Drymoreomys albimaculatus
514:did not form a coherent (
352:is relatively short. The
229:
222:
217:
208:
189:
182:
63:Scientific classification
61:
39:
30:
23:
1339:; Reeder, D. M. (eds.).
1028:Osgood, 1913, p. 97
757:sphenopalatine vacuities
748:bones meet extensively.
542:"lonely place" into the
477:sphenopalatine vacuities
447:was found to be part of
3290:Monotypic rodent genera
895:". It is threatened by
854:Distribution and status
641:among oryzomyines. The
218:Range in northern Peru
1333:"Superfamily Muroidea"
693:
431:belongs. He used both
405:Aegialomys xanthaeolus
311:. The Brazilian genus
985:: e.T15609A22338308.
858:As far as now known,
798:In the mandible, the
781:subsquamosal fenestra
684:
398:and compared it with
348:is rounded. The bony
271:species in the tribe
3075:Ekbletomys hypenemus
1609:A. praeuniversitatis
1400:: e.T15609A4891730.
1376:New Peruvian mammals
665:, and 35 or 36
439:characters from the
259:, also known as the
3132:Eremoryzomys polius
3102:Eremoryzomys polius
1386:Eremoryzomys polius
971:Eremoryzomys polius
897:habitat destruction
860:Eremoryzomys polius
734:interorbital region
611:Eremoryzomys polius
536:Eremoryzomys polius
481:mesopterygoid fossa
423:("rice rats"), the
400:O. xanthaeolus
378:Eremoryzomys polius
366:habitat destruction
360:of the species as "
358:conservation status
256:Eremoryzomys polius
193:Eremoryzomys polius
33:Conservation status
1449:Weksler, M. 2006.
875:and the Brazilian
791:, the roof of the
742:interparietal bone
720:is present in the
694:
638:Sooretamys angouya
380:were collected by
163:, & Voss, 2006
3260:
3259:
3245:Open Tree of Life
3094:Taxon identifiers
3085:
3084:
1507:
1502:Species of tribe
1356:978-0-8018-8221-0
804:masseteric ridges
761:alisphenoid strut
572:Galápagos Islands
479:(openings in the
467:(openings in the
465:incisive foramina
252:
251:
246:
237:
164:
56:
3312:
3253:
3252:
3240:
3239:
3227:
3226:
3214:
3213:
3201:
3200:
3188:
3187:
3175:
3174:
3162:
3161:
3149:
3148:
3136:
3135:
3134:
3121:
3120:
3119:
3089:
2693:O. longicaudatus
2117:Microakodontomys
1741:D. albimaculatus
1691:C. maracajuensis
1570:A. galapagoensis
1505:
1496:
1489:
1482:
1473:
1446:
1436:
1411:
1409:
1368:
1349:. p. 1153.
1345:(3rd ed.).
1322:Literature cited
1316:
1313:
1307:
1304:
1298:
1295:
1289:
1286:
1280:
1277:
1271:
1268:
1262:
1259:
1253:
1250:
1244:
1241:
1235:
1232:
1226:
1223:
1217:
1214:
1208:
1205:
1199:
1196:
1190:
1187:
1181:
1178:
1172:
1169:
1163:
1160:
1154:
1151:
1145:
1142:
1136:
1133:
1122:
1119:
1110:
1107:
1101:
1098:
1092:
1089:
1083:
1080:
1074:
1071:
1065:
1062:
1053:
1050:
1029:
1026:
1013:
1010:
1004:
1003:
1001:
999:
994:
964:
941:
934:
928:
917:
808:capsular process
765:alisphenoid bone
574:and some of the
265:Marañon oryzomys
244:
235:
213:
195:
155:
71:
70:
50:
45:
44:
21:
3320:
3319:
3315:
3314:
3313:
3311:
3310:
3309:
3280:Mammals of Peru
3265:
3264:
3261:
3256:
3248:
3243:
3235:
3232:Observation.org
3230:
3222:
3217:
3209:
3204:
3196:
3191:
3183:
3178:
3170:
3165:
3157:
3152:
3144:
3139:
3130:
3129:
3124:
3115:
3114:
3109:
3096:
3086:
3081:
3058:
3029:
3000:
2978:
2949:
2920:
2896:
2874:
2850:
2777:
2755:
2700:O. magellanicus
2613:
2601:O. sydandersoni
2486:
2462:
2410:
2370:N. maculiventer
2304:
2254:
2183:
2161:
2132:
2127:M. transitorius
2110:
2074:
2018:
1996:
1963:H. megacephalus
1932:
1913:H. brasiliensis
1896:
1825:
1768:
1746:
1724:
1660:
1636:
1614:
1582:
1553:
1509:
1500:
1414:
1381:
1357:
1327:
1324:
1319:
1314:
1310:
1305:
1301:
1296:
1292:
1287:
1283:
1278:
1274:
1269:
1265:
1260:
1256:
1251:
1247:
1242:
1238:
1233:
1229:
1224:
1220:
1215:
1211:
1206:
1202:
1197:
1193:
1188:
1184:
1179:
1175:
1170:
1166:
1161:
1157:
1152:
1148:
1143:
1139:
1134:
1125:
1120:
1113:
1108:
1104:
1099:
1095:
1090:
1086:
1081:
1077:
1072:
1068:
1063:
1056:
1051:
1032:
1027:
1016:
1011:
1007:
997:
995:
966:
965:
954:
950:
945:
944:
935:
931:
925:Oryzomys polius
918:
914:
909:
901:protected areas
881:Atlantic Forest
869:biogeographical
856:
820:
793:tympanic cavity
714:zygomatic plate
679:
673:of Oryzomyini.
608:
392:Oryzomys polius
374:
291:Oryzomys polius
233:Oryzomys polius
204:
197:
191:
178:
154:
65:
57:
46:
42:
35:
17:
12:
11:
5:
3318:
3316:
3308:
3307:
3302:
3297:
3292:
3287:
3282:
3277:
3267:
3266:
3258:
3257:
3255:
3254:
3241:
3228:
3215:
3202:
3189:
3176:
3163:
3150:
3137:
3122:
3106:
3104:
3098:
3097:
3092:
3083:
3082:
3080:
3079:
3068:
3066:
3064:Incertae sedis
3060:
3059:
3057:
3056:
3049:
3041:
3039:
3031:
3030:
3028:
3027:
3020:
3012:
3010:
3007:Transandinomys
3002:
3001:
2999:
2998:
2990:
2988:
2980:
2979:
2977:
2976:
2969:
2961:
2959:
2951:
2950:
2948:
2947:
2944:S. ucayalensis
2940:
2932:
2930:
2922:
2921:
2919:
2918:
2909:
2907:
2898:
2897:
2895:
2894:
2886:
2884:
2876:
2875:
2873:
2872:
2863:
2861:
2852:
2851:
2849:
2848:
2841:
2834:
2826:
2819:
2812:
2805:
2797:
2789:
2787:
2779:
2778:
2776:
2775:
2767:
2765:
2757:
2756:
2754:
2753:
2745:
2738:
2731:
2724:
2717:
2710:
2703:
2696:
2689:
2682:
2675:
2668:
2661:
2654:
2647:
2640:
2633:
2625:
2623:
2615:
2614:
2612:
2611:
2604:
2597:
2590:
2583:
2576:
2569:
2562:
2555:
2548:
2541:
2534:
2527:
2520:
2513:
2506:
2498:
2496:
2488:
2487:
2485:
2484:
2475:
2473:
2464:
2463:
2461:
2460:
2453:
2450:N. narboroughi
2446:
2443:N. fernandinae
2439:
2431:
2422:
2420:
2412:
2411:
2409:
2408:
2401:
2394:
2387:
2380:
2373:
2366:
2359:
2352:
2345:
2338:
2331:
2324:
2321:N. albigularis
2316:
2314:
2306:
2305:
2303:
2302:
2295:
2288:
2281:
2274:
2266:
2264:
2256:
2255:
2253:
2252:
2245:
2238:
2231:
2224:
2217:
2210:
2203:
2195:
2193:
2185:
2184:
2182:
2181:
2173:
2171:
2163:
2162:
2160:
2159:
2152:
2144:
2142:
2134:
2133:
2131:
2130:
2122:
2120:
2112:
2111:
2109:
2108:
2101:
2094:
2091:M. caliginosus
2086:
2084:
2076:
2075:
2073:
2072:
2064:
2056:
2053:M. desmarestii
2048:
2040:
2031:
2029:
2020:
2019:
2017:
2016:
2008:
2006:
1998:
1997:
1995:
1994:
1987:
1980:
1973:
1966:
1959:
1952:
1944:
1942:
1934:
1933:
1931:
1930:
1923:
1916:
1908:
1906:
1898:
1897:
1895:
1894:
1887:
1880:
1873:
1866:
1859:
1852:
1845:
1837:
1835:
1827:
1826:
1824:
1823:
1816:
1809:
1806:E. macconnelli
1802:
1795:
1788:
1780:
1778:
1770:
1769:
1767:
1766:
1758:
1756:
1748:
1747:
1745:
1744:
1736:
1734:
1726:
1725:
1723:
1722:
1715:
1708:
1701:
1694:
1687:
1680:
1672:
1670:
1662:
1661:
1659:
1658:
1649:
1647:
1638:
1637:
1635:
1634:
1626:
1624:
1616:
1615:
1613:
1612:
1604:
1595:
1593:
1584:
1583:
1581:
1580:
1577:A. xanthaeolus
1573:
1565:
1563:
1555:
1554:
1552:
1551:
1545:
1539:
1533:
1527:
1521:
1514:
1511:
1510:
1501:
1499:
1498:
1491:
1484:
1476:
1470:
1469:
1458:
1447:
1427:(2): 357–390.
1412:
1379:
1369:
1355:
1323:
1320:
1318:
1317:
1308:
1299:
1290:
1281:
1272:
1263:
1254:
1245:
1236:
1227:
1218:
1209:
1200:
1191:
1182:
1173:
1164:
1155:
1146:
1137:
1123:
1111:
1102:
1093:
1084:
1075:
1066:
1054:
1030:
1014:
1005:
951:
949:
946:
943:
942:
929:
911:
910:
908:
905:
893:Data Deficient
889:E. polius
855:
852:
819:
816:
800:mental foramen
789:tegmen tympani
767:separates two
759:. Usually, an
722:zygomatic arch
710:premaxillaries
678:
675:
661:, 7 or 8
607:
604:
520:O. polius
501:O. polius
497:O. polius
445:O. polius
429:O. polius
396:O. polius
382:Wilfred Osgood
373:
370:
362:Data Deficient
281:Wilfred Osgood
275:of the family
250:
249:
248:
247:
238:
227:
226:
220:
219:
215:
214:
206:
205:
198:
187:
186:
180:
179:
175:E. polius
172:
170:
166:
165:
147:
143:
142:
137:
133:
132:
127:
123:
122:
117:
113:
112:
107:
103:
102:
97:
93:
92:
87:
83:
82:
77:
73:
72:
59:
58:
48:Data Deficient
40:
37:
36:
31:
28:
27:
15:
13:
10:
9:
6:
4:
3:
2:
3317:
3306:
3303:
3301:
3298:
3296:
3293:
3291:
3288:
3286:
3283:
3281:
3278:
3276:
3273:
3272:
3270:
3263:
3251:
3246:
3242:
3238:
3233:
3229:
3225:
3220:
3216:
3212:
3207:
3203:
3199:
3194:
3190:
3186:
3181:
3177:
3173:
3168:
3164:
3160:
3155:
3151:
3147:
3142:
3138:
3133:
3127:
3123:
3118:
3112:
3108:
3107:
3105:
3103:
3099:
3095:
3090:
3078:
3076:
3070:
3069:
3067:
3065:
3061:
3055:
3054:
3050:
3048:
3047:
3046:Z. brevicauda
3043:
3042:
3040:
3038:
3037:
3032:
3026:
3025:
3024:T. talamancae
3021:
3019:
3018:
3014:
3013:
3011:
3009:
3008:
3003:
2997:
2996:
2992:
2991:
2989:
2987:
2986:
2981:
2975:
2974:
2970:
2968:
2967:
2963:
2962:
2960:
2958:
2957:
2956:Sigmodontomys
2952:
2946:
2945:
2941:
2939:
2938:
2934:
2933:
2931:
2929:
2928:
2923:
2917:
2916:
2915:R. primigenus
2911:
2910:
2908:
2906:
2905:
2899:
2893:
2892:
2888:
2887:
2885:
2883:
2882:
2881:Pseudoryzomys
2877:
2871:
2870:
2865:
2864:
2862:
2860:
2859:
2853:
2847:
2846:
2845:O. peninsulae
2842:
2840:
2839:
2838:O. palustris
2835:
2833:
2832:
2827:
2825:
2824:
2820:
2818:
2817:
2816:O. dimidiatus
2813:
2811:
2810:
2806:
2804:
2803:
2802:O. antillarum
2798:
2796:
2795:
2794:O. albiventer
2791:
2790:
2788:
2786:
2785:
2780:
2774:
2773:
2769:
2768:
2766:
2764:
2763:
2758:
2752:
2751:
2746:
2744:
2743:
2739:
2737:
2736:
2735:O. stramineus
2732:
2730:
2729:
2725:
2723:
2722:
2718:
2716:
2715:
2711:
2709:
2708:
2704:
2702:
2701:
2697:
2695:
2694:
2690:
2688:
2687:
2683:
2681:
2680:
2679:O. fulvescens
2676:
2674:
2673:
2669:
2667:
2666:
2665:O. flavescens
2662:
2660:
2659:
2658:O. destructor
2655:
2653:
2652:
2651:O. chacoensis
2648:
2646:
2645:
2641:
2639:
2638:
2634:
2632:
2631:
2627:
2626:
2624:
2622:
2621:
2616:
2610:
2609:
2608:O. trinitatis
2605:
2603:
2602:
2598:
2596:
2595:
2591:
2589:
2588:
2584:
2582:
2581:
2577:
2575:
2574:
2570:
2568:
2567:
2563:
2561:
2560:
2556:
2554:
2553:
2549:
2547:
2546:
2542:
2540:
2539:
2535:
2533:
2532:
2528:
2526:
2525:
2521:
2519:
2518:
2517:O. catherinae
2514:
2512:
2511:
2507:
2505:
2504:
2503:O. auyantepui
2500:
2499:
2497:
2495:
2494:
2489:
2483:
2482:
2477:
2476:
2474:
2472:
2471:
2465:
2459:
2458:
2454:
2452:
2451:
2447:
2445:
2444:
2440:
2438:
2437:
2436:N. indefessus
2432:
2430:
2429:
2424:
2423:
2421:
2419:
2418:
2413:
2407:
2406:
2402:
2400:
2399:
2398:N. pectoralis
2395:
2393:
2392:
2388:
2386:
2385:
2381:
2379:
2378:
2377:N. meridensis
2374:
2372:
2371:
2367:
2365:
2364:
2360:
2358:
2357:
2353:
2351:
2350:
2346:
2344:
2343:
2339:
2337:
2336:
2332:
2330:
2329:
2328:N. auriventer
2325:
2323:
2322:
2318:
2317:
2315:
2313:
2312:
2307:
2301:
2300:
2296:
2294:
2293:
2289:
2287:
2286:
2282:
2280:
2279:
2275:
2273:
2272:
2268:
2267:
2265:
2263:
2262:
2257:
2251:
2250:
2246:
2244:
2243:
2239:
2237:
2236:
2232:
2230:
2229:
2225:
2223:
2222:
2218:
2216:
2215:
2211:
2209:
2208:
2204:
2202:
2201:
2197:
2196:
2194:
2192:
2191:
2186:
2180:
2179:
2175:
2174:
2172:
2170:
2169:
2164:
2158:
2157:
2153:
2151:
2150:
2149:M. altissimus
2146:
2145:
2143:
2141:
2140:
2135:
2129:
2128:
2124:
2123:
2121:
2119:
2118:
2113:
2107:
2106:
2102:
2100:
2099:
2098:M. robustulus
2095:
2093:
2092:
2088:
2087:
2085:
2083:
2082:
2077:
2071:
2070:
2065:
2063:
2062:
2057:
2055:
2054:
2049:
2047:
2046:
2045:M. curazensis
2041:
2039:
2038:
2033:
2032:
2030:
2028:
2027:
2021:
2015:
2014:
2010:
2009:
2007:
2005:
2004:
1999:
1993:
1992:
1988:
1986:
1985:
1981:
1979:
1978:
1977:H. perenensis
1974:
1972:
1971:
1967:
1965:
1964:
1960:
1958:
1957:
1953:
1951:
1950:
1946:
1945:
1943:
1941:
1940:
1935:
1929:
1928:
1924:
1922:
1921:
1917:
1915:
1914:
1910:
1909:
1907:
1905:
1904:
1899:
1893:
1892:
1891:H. saturatior
1888:
1886:
1885:
1881:
1879:
1878:
1874:
1872:
1871:
1867:
1865:
1864:
1860:
1858:
1857:
1853:
1851:
1850:
1846:
1844:
1843:
1839:
1838:
1836:
1834:
1833:
1828:
1822:
1821:
1817:
1815:
1814:
1810:
1808:
1807:
1803:
1801:
1800:
1796:
1794:
1793:
1789:
1787:
1786:
1782:
1781:
1779:
1777:
1776:
1771:
1765:
1764:
1760:
1759:
1757:
1755:
1754:
1749:
1743:
1742:
1738:
1737:
1735:
1733:
1732:
1727:
1721:
1720:
1716:
1714:
1713:
1709:
1707:
1706:
1702:
1700:
1699:
1695:
1693:
1692:
1688:
1686:
1685:
1681:
1679:
1678:
1674:
1673:
1671:
1669:
1668:
1663:
1657:
1656:
1651:
1650:
1648:
1646:
1645:
1639:
1633:
1632:
1628:
1627:
1625:
1623:
1622:
1621:Amphinectomys
1617:
1611:
1610:
1605:
1603:
1602:
1597:
1596:
1594:
1592:
1591:
1585:
1579:
1578:
1574:
1572:
1571:
1567:
1566:
1564:
1562:
1561:
1556:
1550:
1549:Sigmodontinae
1546:
1544:
1540:
1538:
1534:
1532:
1528:
1526:
1522:
1520:
1516:
1515:
1512:
1508:
1497:
1492:
1490:
1485:
1483:
1478:
1477:
1474:
1467:
1463:
1459:
1456:
1452:
1448:
1444:
1440:
1435:
1430:
1426:
1422:
1418:
1413:
1408:
1403:
1399:
1395:
1394:
1389:
1387:
1380:
1377:
1373:
1370:
1366:
1362:
1358:
1352:
1348:
1344:
1343:
1338:
1337:Wilson, D. E.
1334:
1330:
1329:Musser, G. G.
1326:
1325:
1321:
1312:
1309:
1303:
1300:
1294:
1291:
1285:
1282:
1276:
1273:
1267:
1264:
1258:
1255:
1249:
1246:
1240:
1237:
1231:
1228:
1222:
1219:
1213:
1210:
1204:
1201:
1195:
1192:
1186:
1183:
1177:
1174:
1168:
1165:
1159:
1156:
1150:
1147:
1141:
1138:
1132:
1130:
1128:
1124:
1118:
1116:
1112:
1106:
1103:
1097:
1094:
1088:
1085:
1079:
1076:
1070:
1067:
1061:
1059:
1055:
1049:
1047:
1045:
1043:
1041:
1039:
1037:
1035:
1031:
1025:
1023:
1021:
1019:
1015:
1009:
1006:
993:
988:
984:
980:
979:
974:
972:
963:
961:
959:
957:
953:
947:
939:
933:
930:
926:
922:
916:
913:
906:
904:
902:
898:
894:
890:
886:
882:
878:
874:
870:
866:
865:Marañón River
861:
853:
851:
849:
845:
841:
837:
833:
829:
825:
817:
815:
813:
809:
805:
801:
796:
794:
790:
786:
782:
778:
774:
770:
766:
762:
758:
754:
749:
747:
743:
739:
735:
731:
727:
723:
719:
715:
711:
707:
703:
699:
691:
687:
683:
676:
674:
672:
668:
664:
660:
656:
652:
648:
644:
640:
639:
634:
630:
626:
622:
621:
616:
612:
605:
603:
601:
597:
593:
589:
585:
581:
580:Sigmodontinae
577:
573:
569:
565:
563:
559:
558:
553:
549:
548:IUCN Red List
545:
541:
537:
533:
529:
525:
521:
517:
513:
508:
506:
502:
498:
494:
490:
486:
482:
478:
474:
470:
466:
462:
461:parietal bone
458:
457:synapomorphic
454:
450:
446:
442:
438:
434:
433:morphological
430:
426:
422:
418:
413:
411:
407:
406:
401:
397:
393:
389:
388:
383:
379:
371:
369:
367:
363:
359:
356:assesses the
355:
351:
347:
343:
339:
335:
330:
328:
327:Marañón River
324:
320:
316:
315:
310:
309:
304:
300:
296:
293:. In 2006, a
292:
288:
287:
282:
278:
274:
270:
266:
262:
261:gray rice rat
258:
257:
242:
239:
234:
231:
230:
228:
225:
221:
216:
212:
207:
202:
196:
194:
188:
185:
184:Binomial name
181:
177:
176:
171:
168:
167:
162:
158:
153:
152:
148:
145:
144:
141:
140:Sigmodontinae
138:
135:
134:
131:
128:
125:
124:
121:
118:
115:
114:
111:
108:
105:
104:
101:
98:
95:
94:
91:
88:
85:
84:
81:
78:
75:
74:
69:
64:
60:
54:
49:
38:
34:
29:
26:
22:
19:
3262:
3101:
3072:
3051:
3044:
3036:Zygodontomys
3034:
3022:
3017:T. bolivaris
3015:
3005:
2993:
2983:
2973:S. aphrastus
2971:
2964:
2954:
2942:
2935:
2925:
2913:
2902:
2889:
2879:
2867:
2856:
2843:
2837:
2829:
2821:
2814:
2807:
2800:
2792:
2782:
2772:O. balneator
2770:
2760:
2748:
2740:
2733:
2728:O. rupestris
2726:
2719:
2712:
2705:
2698:
2691:
2686:O. griseolus
2684:
2677:
2670:
2663:
2656:
2649:
2642:
2635:
2628:
2620:Oligoryzomys
2618:
2606:
2599:
2592:
2587:O. speciosus
2585:
2578:
2571:
2564:
2557:
2550:
2543:
2538:O. flavicans
2536:
2529:
2522:
2515:
2508:
2501:
2491:
2481:N. vespuccii
2479:
2468:
2455:
2448:
2441:
2434:
2426:
2415:
2405:N. pirrensis
2403:
2396:
2389:
2382:
2375:
2368:
2361:
2354:
2347:
2340:
2335:N. caracolus
2333:
2326:
2319:
2309:
2299:N. squamipes
2297:
2290:
2283:
2276:
2269:
2259:
2247:
2240:
2233:
2226:
2219:
2212:
2205:
2198:
2188:
2176:
2166:
2154:
2147:
2139:Microryzomys
2137:
2125:
2115:
2103:
2096:
2089:
2079:
2067:
2061:M. georginae
2059:
2051:
2043:
2035:
2024:
2011:
2001:
1989:
1982:
1975:
1968:
1961:
1954:
1947:
1937:
1925:
1920:H. chacarius
1918:
1911:
1901:
1889:
1884:H. rostratus
1882:
1875:
1870:H. melanotis
1868:
1861:
1854:
1847:
1840:
1830:
1818:
1811:
1804:
1797:
1790:
1783:
1775:Euryoryzomys
1773:
1762:
1761:
1753:Eremoryzomys
1752:
1751:
1739:
1729:
1717:
1712:C. subflavus
1710:
1703:
1696:
1689:
1684:C. langguthi
1682:
1675:
1665:
1653:
1644:Carletonomys
1642:
1629:
1619:
1607:
1599:
1590:Agathaeromys
1588:
1575:
1568:
1558:
1465:
1454:
1424:
1420:
1397:
1391:
1385:
1372:Osgood, W.H.
1341:
1311:
1302:
1293:
1284:
1275:
1266:
1257:
1248:
1239:
1230:
1221:
1212:
1203:
1194:
1185:
1176:
1167:
1158:
1149:
1140:
1105:
1096:
1087:
1078:
1069:
1008:
996:. Retrieved
982:
976:
970:
938:Eremoryzomys
937:
932:
924:
921:Eremoryzomys
920:
915:
888:
885:Eremoryzomys
884:
876:
873:Eremoryzomys
872:
859:
857:
821:
812:Eremoryzomys
811:
797:
785:mastoid bone
750:
695:
686:Eremoryzomys
685:
671:synapomorphy
657:has 12
654:
636:
629:ungual tufts
618:
610:
609:
568:Eremoryzomys
567:
566:
562:Eremoryzomys
561:
555:
552:Eremoryzomys
551:
544:generic name
539:
535:
531:
528:Eremoryzomys
527:
526:, including
523:
519:
516:monophyletic
511:
509:
500:
496:
453:sister group
444:
428:
419:analysis of
414:
409:
403:
399:
395:
391:
385:
377:
375:
338:ungual tufts
333:
331:
323:Eremoryzomys
322:
319:Eremoryzomys
318:
312:
308:Eremoryzomys
307:
306:
298:
290:
284:
264:
260:
255:
254:
253:
240:
236:Osgood, 1913
232:
192:
190:
174:
173:
151:Eremoryzomys
150:
149:
25:Eremoryzomys
24:
18:
3167:iNaturalist
3126:Wikispecies
3053:Z. brunneus
2937:S. melanops
2762:Oreoryzomys
2721:O. nigripes
2707:O. microtis
2637:O. arenalis
2594:O. superans
2559:O. phaeotis
2552:O. paricola
2531:O. concolor
2417:Nesoryzomys
2391:N. nimbosus
2285:N. palmipes
2271:N. apicalis
2249:N. tenuipes
2242:N. spinosus
2178:M. hammondi
2037:M. audreyae
1991:H. yunganus
1956:H. laticeps
1927:H. sciureus
1877:H. rhabdops
1863:H. intectus
1856:H. fuscatus
1849:H. chapmani
1832:Handleyomys
1820:E. russatus
1785:E. emmonsae
1731:Drymoreomys
1698:C. marinhus
1601:A. donovani
1547:Subfamily:
1506:(rice rats)
1504:Oryzomyini
998:14 November
877:Drymoreomys
848:anteroconid
840:mesolophids
702:nasal bones
690:Río Marañón
606:Description
546:. The 2008
493:posteroloph
402:(currently
314:Drymoreomys
161:Percequillo
136:Subfamily:
3285:Oryzomyini
3269:Categories
2995:S. angouya
2985:Sooretamys
2891:P. simplex
2869:P. nivalis
2858:Pennatomys
2831:O. nelsoni
2823:O. gorgasi
2742:O. vegetus
2714:O. moojeni
2672:O. fornesi
2644:O. brendae
2630:O. andinus
2580:O. rutilus
2573:O. roberti
2545:O. mamorae
2524:O. cleberi
2510:O. bicolor
2470:Noronhomys
2457:N. swarthi
2428:N. darwini
2363:N. levipes
2311:Nephelomys
2278:N. grandis
2228:N. paracou
2221:N. musseri
2214:N. minutus
2207:N. guianae
2200:N. dubosti
2156:M. minutus
2105:M. zunigae
2013:L. molitor
1970:H. oniscus
1949:H. acritus
1903:Holochilus
1842:H. alfaroi
1813:E. nitidus
1799:E. legatus
1677:C. goytaca
1667:Cerradomys
1631:A. savamis
1560:Aegialomys
1543:Cricetidae
1468:3537:1–29.
1457:296:1–149.
948:References
844:anterocone
832:brachydont
718:jugal bone
708:, and the
584:Cricetidae
505:mesoflexus
491:; and the
421:Oryzomyini
289:and named
277:Cricetidae
273:Oryzomyini
130:Cricetidae
2966:S. alfari
2809:O. couesi
2750:O. victus
2384:N. moerex
2356:N. keaysi
2349:N. devius
2342:N. childi
2292:N. rattus
2235:N. pictus
2081:Melanomys
2069:M. luciae
2026:Megalomys
1939:Hylaeamys
1763:E. polius
1705:C. scotti
1655:C. cailoi
1517:Kingdom:
1443:1096-3642
836:mesolophs
746:occipital
738:braincase
730:squamosal
726:maxillary
706:lacrimals
655:E. polius
437:molecular
435:data and
427:to which
417:cladistic
346:braincase
334:E. polius
295:cladistic
169:Species:
86:Kingdom:
80:Eukaryota
3117:Q1769498
3111:Wikidata
2927:Scolomys
2904:Reigomys
2784:Oryzomys
2261:Nectomys
2190:Neacomys
2168:Mindomys
2003:Lundomys
1984:H. tatei
1792:E. lamia
1719:C. vivoi
1541:Family:
1537:Rodentia
1531:Mammalia
1525:Chordata
1523:Phylum:
1519:Animalia
1365:62265494
828:bunodont
775:and the
769:foramina
659:thoracic
615:woodrats
600:deermice
596:hamsters
592:lemmings
576:Antilles
524:Oryzomys
512:Oryzomys
485:arterial
410:Oryzomys
387:Oryzomys
372:Taxonomy
299:Oryzomys
286:Oryzomys
224:Synonyms
126:Family:
120:Rodentia
110:Mammalia
100:Chordata
96:Phylum:
90:Animalia
76:Domain:
53:IUCN 3.1
3250:1035611
3211:1002531
3159:5786881
2493:Oecomys
1535:Order:
1529:Class:
696:In the
647:phalanx
643:squamae
633:plantar
620:Neotoma
489:derived
449:clade D
267:, is a
263:or the
203:, 1913)
157:Weksler
146:Genus:
116:Order:
106:Class:
51: (
3237:200581
3224:530169
3185:972463
2566:O. rex
1441:
1374:1913.
1363:
1353:
842:. The
824:molars
818:Molars
667:caudal
663:lumbar
651:mammae
625:pinnae
598:, and
540:eremia
532:polius
469:palate
350:palate
342:mammae
269:rodent
241:polius
201:Osgood
3198:15609
3172:74316
3146:6GG6M
1335:. In
907:Notes
753:fossa
698:skull
677:Skull
588:voles
425:tribe
303:genus
3219:NCBI
3193:IUCN
3180:ITIS
3154:GBIF
1439:ISSN
1398:2008
1361:OCLC
1351:ISBN
1000:2021
983:2019
923:(as
846:and
838:and
826:are
822:The
728:and
530:for
441:IRBP
354:IUCN
3206:MDD
3141:CoL
1429:doi
1425:161
1402:doi
987:doi
3271::
3247::
3234::
3221::
3208::
3195::
3182::
3169::
3156::
3143::
3128::
3113::
1464:.
1453:.
1437:.
1423:.
1419:.
1396:.
1390:.
1359:.
1126:^
1114:^
1057:^
1033:^
1017:^
981:.
975:.
955:^
903:.
594:,
590:,
564:.
412:.
390:,
368:.
321:.
305:,
243::
159:,
3077:"
3073:"
3071:†
2912:†
2901:†
2866:†
2855:†
2828:†
2799:†
2747:†
2478:†
2467:†
2433:†
2425:†
2066:†
2058:†
2050:†
2042:†
2034:†
2023:†
1652:†
1641:†
1606:†
1598:†
1587:†
1495:e
1488:t
1481:v
1445:.
1431::
1410:.
1404::
1388:"
1384:"
1367:.
1002:.
989::
973:"
969:"
940:.
692:.
617:(
199:(
55:)
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