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Bottom: Current view. Two simultaneous strand-swaps shown, each depending on the homology of three successive bases at the edge of the (6-8 bp long) spacer. Didactic complications arise from the fact that the model has to account for the fact that the synaptic complex (1) contains both substrates in
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Top: Classical view including strand-exchange followed by branch-migration (proofreading). The mechanism takes within a synaptic complex (1) including both DNA-sites in parallel orientation. While branch-migration provides a straightforward explanation regarding the specific homology requirements an
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Both representations illustrate only half of the respective pathways. Both parts are separated by a
Holliday-junction/isomerization step, before the product (3) is released
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the reversibility of the process, it is difficult to reconcile with the motions the recombinase subunits have to undergo in three dimensions
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