184:
66:
41:
321:. It meets the criteria to be listed as endangered because its extent of occurrence is less than 5,000 km (1,900 sq mi), its range is severely fragmented, and its habitat is projected to decline in extend and quality. It is believed that its population size is declining. Some of the main threats facing this species are
245:
It is a small species of bat. Its forearm is only 30â33 mm (1.2â1.3 in) long. The fur of its back is brackish brown, with conspicuous frosted tips. The fur of its belly is lighter in color than the dorsal surface. The fur is velvety in texture, dense, and short, with individual hairs
410:
Kawai, K., Nikaido, M., Harada, M., Matsumura, S., Lin, L. K., Wu, Y., ... & Okada, N. (2003). The status of the
Japanese and East Asian bats of the genus Myotis (Vespertilionidae) based on mitochondrial sequences. Molecular Phylogenetics and Evolution, 28(2),
420:
Harada, M., & Uchida, T. A. (1982). Karyotype of a rare species, Myotis pruinosus, involving pericentric inversion and duplicated translocation. Cytologia, 47(3-4), 539-543.
289:
It is found most often in mountain forests of lower elevations. It is encountered at elevations of 200â300 m (660â980 ft) above sea level. It has been recorded in the
1700:
318:
1765:
1819:
1674:
1713:
448:
1834:
617:
277:
during the day. It is therefore dependent on large trees with hollows, and is not found in habitats where they aren't any. It is a
230:. Based on its similarity to these two species, it is hypothesized that the ancestor of the frosted myotis evolved to the south of
1739:
45:
1757:
1144:
1718:
246:
approximately 5.5 mm (0.22 in) long. Its wing membranes are dark brown. Their ears are narrow and short, with the
1433:
1187:
1752:
1586:
1074:
1354:
786:
1770:
1408:
1126:
441:
401:
Yoshiyuki, M. (1989). A systematic study of the
Japanese Chiroptera. National Science Museum monographs, 7, 1-242.
1824:
1110:
822:
940:
759:
65:
1442:
1302:
1275:
1196:
974:
1596:
1580:
1614:
1417:
958:
687:
678:
1829:
1537:
1169:
434:
326:
227:
223:
160:
1652:
1661:
1381:
1135:
1058:
1018:
968:
663:
538:
1040:
888:
644:
626:
377:
322:
219:
30:
1519:
1479:
1327:
1154:
696:
314:
60:
1528:
1399:
1345:
1008:
992:
983:
804:
669:
608:
183:
1705:
1791:
1321:
1311:
1293:
1241:
1214:
1034:
1002:
882:
872:
856:
813:
768:
732:
723:
635:
584:
1796:
1427:
1101:
1092:
1083:
1049:
931:
922:
904:
866:
838:
832:
520:
494:
372:
127:
1731:
1778:
1372:
1120:
913:
714:
653:
590:
556:
529:
500:
137:
1363:
1178:
705:
1494:
1485:
1451:
1284:
1266:
1223:
1024:
847:
777:
741:
599:
574:
565:
1813:
1726:
1555:
1513:
1260:
1205:
1160:
1068:
898:
363:
354:
50:
1250:
1232:
750:
547:
247:
1744:
1687:
1646:
1570:
1546:
1503:
1460:
426:
281:
species, with two copies each of 22 chromosomes, for a total of 44 chromosomes.
274:
1469:
1390:
203:
1637:
251:
77:
260:
257:
1783:
1631:
949:
795:
476:
458:
97:
1679:
294:
218:
It was first discovered in August 1969 by Kimio Endo. In 1971, it was
1692:
511:
482:
470:
298:
290:
278:
107:
87:
1608:
302:
231:
207:
1666:
1612:
430:
488:
348:
346:
344:
342:
117:
222:
by Mizuko
Yoshiyuki. It is most closely related to the
1621:
1568:
509:
397:
395:
234:, and its range expanded northwards over time.
378:10.2305/IUCN.UK.2019-3.RLTS.T14192A22066846.en
442:
8:
1609:
449:
435:
427:
182:
39:
20:
376:
338:
237:In Japanese it is known as ăŻăăăȘăăČăłăŠăąăȘ.
250:about half as high as the ear. Their
7:
364:IUCN Red List of Threatened Species
14:
618:Sir David Attenborough's myotis (
1820:IUCN Red List endangered species
1145:Dark-nosed small-footed myotis (
64:
1188:Wall-roosting mouse-eared bat (
1:
1587:Taiwan broad-muzzled myotis (
1075:Eastern small-footed myotis (
313:It is currently evaluated as
1434:Mandelli's mouse-eared bat (
1127:Pallid large-footed myotis (
353:Fukui, D.; Sano, A. (2019).
1111:Eastern long-fingered bat (
265:, for a total of 38 teeth.
1851:
1355:Rickett's big-footed bat (
941:Malagasy mouse-eared bat (
787:Csorba's mouse-eared bat (
760:Western small-footed bat (
1835:Mammals described in 1971
1443:Himalayan whiskered bat (
1418:Northern long-eared bat (
1409:Scott's mouse-eared bat (
1303:Beijing mouse-eared bat (
1276:Singapore whiskered bat (
1197:Greater mouse-eared bat (
975:Lesser large-footed bat (
465:
181:
166:
159:
61:Scientific classification
59:
37:
28:
23:
959:Armenian whiskered bat (
823:Kock's mouse-eared bat (
688:Rufous mouse-eared bat (
679:Lesser mouse-eared bat (
1538:Yanbaru whiskered bat (
1170:Burmese whiskered bat (
1382:Thick-thumbed myotis (
1059:Chinese water myotis (
539:Silver-tipped myotis (
206:. It is found only in
1041:Hairy-legged myotis (
645:Southeastern myotis (
627:Southwestern myotis (
457:Species of subfamily
371:: e.T14192A22066846.
228:Burmese whiskered bat
224:Yanbaru whiskered bat
1520:Long-legged myotis (
1328:Flat-headed myotis (
1242:Nimba mountain bat (
697:Far Eastern myotis (
327:forest fragmentation
1312:Eastern water bat (
1294:Peninsular myotis (
1136:Schwartz's myotis (
873:Long-eared myotis (
769:Guatemalan myotis (
733:Long-fingered bat (
724:California myotis (
636:Australian myotis (
323:habitat destruction
31:Conservation status
1420:M. septentrionalis
1102:Little brown bat (
1093:Kashmir cave bat (
1084:Yellowish myotis (
932:Gomantong myotis (
923:Fraternal myotis (
889:Findley's myotis (
839:Dominican myotis (
521:Large-footed bat (
202:) is a species of
1807:
1806:
1792:Open Tree of Life
1615:Taxon identifiers
1606:
1605:
1529:Welwitsch's bat (
1400:Schaub's myotis (
1373:Riparian myotis (
1346:Felten's myotis (
1138:M. martiniquensis
1009:Ikonnikov's bat (
993:Horsfield's bat (
984:Herman's myotis (
934:M. gomantongensis
914:Cinnamon myotis (
805:Daubenton's bat (
715:Bocharic myotis (
670:Bechstein's bat (
654:Chestnut myotis (
647:M. austroriparius
609:Peters's myotis (
591:Hairy-faced bat (
557:Szechwan myotis (
548:Myotis alcathoe (
530:Southern myotis (
285:Range and habitat
190:
189:
152:M. pruinosus
54:
1842:
1825:Mouse-eared bats
1800:
1799:
1787:
1786:
1774:
1773:
1761:
1760:
1748:
1747:
1735:
1734:
1722:
1721:
1709:
1708:
1696:
1695:
1683:
1682:
1670:
1669:
1657:
1656:
1655:
1653:Myotis pruinosus
1642:
1641:
1640:
1623:Myotis pruinosus
1610:
1495:Cape hairy bat (
1486:Fringed myotis (
1452:Velvety myotis (
1337:Frosted myotis (
1285:Montane myotis (
1267:Arizona myotis (
1224:Curacao myotis (
1215:Natterer's bat (
1113:M. macrodactylus
1025:Insular myotis (
1019:M. indochinensis
857:Geoffroy's bat (
848:Elegant myotis (
814:David's myotis (
780:Myotis crypticus
778:Cryptic myotis (
742:Chilean myotis (
620:M. attenboroughi
600:Atacama myotis (
575:Annamit myotis (
566:Anjouan myotis (
495:Vespertilionidae
451:
444:
437:
428:
421:
418:
412:
408:
402:
399:
390:
389:
387:
385:
380:
357:Myotis pruinosus
350:
264:
263:
262:
259:
199:Myotis pruinosus
186:
172:
170:Myotis pruinosus
128:Vespertilionidae
69:
68:
48:
43:
42:
21:
1850:
1849:
1845:
1844:
1843:
1841:
1840:
1839:
1810:
1809:
1808:
1803:
1795:
1790:
1782:
1779:Observation.org
1777:
1769:
1764:
1756:
1751:
1743:
1738:
1730:
1725:
1717:
1712:
1704:
1699:
1691:
1686:
1678:
1673:
1665:
1660:
1651:
1650:
1645:
1636:
1635:
1630:
1617:
1607:
1602:
1564:
1558:Myotis zenatius
1556:Zenati myotis (
1445:M. siligorensis
1296:M. peninsularis
1206:Whiskered bat (
1161:Maluku myotis (
1147:M. melanorhinus
1050:Keen's myotis (
961:M. hajastanicus
905:Hodgson's bat (
841:M. dominicensis
726:M. californicus
568:M. anjouanensis
505:
461:
455:
425:
424:
419:
415:
409:
405:
400:
393:
383:
381:
352:
351:
340:
335:
311:
287:
271:
256:
255:
243:
216:
177:
176:Yoshiyuki, 1971
174:
168:
155:
63:
55:
44:
40:
33:
24:Frosted myotis
17:
12:
11:
5:
1848:
1846:
1838:
1837:
1832:
1827:
1822:
1812:
1811:
1805:
1804:
1802:
1801:
1788:
1775:
1762:
1749:
1736:
1723:
1710:
1697:
1684:
1671:
1658:
1643:
1627:
1625:
1619:
1618:
1613:
1604:
1603:
1601:
1600:
1597:S. moupinensis
1593:
1589:S. latirostris
1584:
1581:S. caliginosus
1576:
1574:
1566:
1565:
1563:
1562:
1553:
1544:
1540:M. yanbarensis
1535:
1531:M. welwitschii
1526:
1517:
1510:
1501:
1492:
1483:
1480:M. taiwanensis
1476:
1467:
1458:
1449:
1440:
1431:
1424:
1415:
1406:
1397:
1388:
1379:
1370:
1364:Ridley's bat (
1361:
1352:
1343:
1334:
1325:
1318:
1309:
1300:
1291:
1282:
1273:
1264:
1257:
1251:Nepal myotis (
1248:
1239:
1233:Black myotis (
1230:
1221:
1212:
1203:
1194:
1185:
1179:Morris's bat (
1176:
1167:
1158:
1155:M. midastactus
1151:
1142:
1133:
1129:M. macrotarsus
1124:
1117:
1108:
1099:
1090:
1081:
1072:
1065:
1056:
1047:
1038:
1031:
1022:
1015:
1006:
999:
995:M. horsfieldii
990:
981:
972:
965:
956:
947:
938:
929:
920:
911:
902:
895:
886:
879:
870:
863:
859:M. emarginatus
854:
845:
836:
829:
820:
811:
807:M. daubentonii
802:
793:
784:
775:
766:
762:M. ciliolabrum
757:
751:Large myotis (
748:
739:
730:
721:
717:M. bucharensis
712:
706:Brandt's bat (
703:
694:
685:
676:
672:M. bechsteinii
667:
660:
651:
642:
633:
624:
615:
606:
602:M. atacamensis
597:
588:
581:
577:M. annamiticus
572:
563:
554:
545:
536:
527:
517:
515:
507:
506:
504:
503:
497:
491:
485:
479:
473:
466:
463:
462:
456:
454:
453:
446:
439:
431:
423:
422:
413:
403:
391:
337:
336:
334:
331:
310:
307:
286:
283:
270:
267:
252:dental formula
242:
239:
215:
212:
194:frosted myotis
188:
187:
179:
178:
175:
164:
163:
157:
156:
149:
147:
143:
142:
135:
131:
130:
125:
121:
120:
115:
111:
110:
105:
101:
100:
95:
91:
90:
85:
81:
80:
75:
71:
70:
57:
56:
38:
35:
34:
29:
26:
25:
16:Species of bat
15:
13:
10:
9:
6:
4:
3:
2:
1847:
1836:
1833:
1831:
1828:
1826:
1823:
1821:
1818:
1817:
1815:
1798:
1793:
1789:
1785:
1780:
1776:
1772:
1767:
1763:
1759:
1754:
1750:
1746:
1741:
1737:
1733:
1728:
1724:
1720:
1715:
1711:
1707:
1702:
1698:
1694:
1689:
1685:
1681:
1676:
1672:
1668:
1663:
1659:
1654:
1648:
1644:
1639:
1633:
1629:
1628:
1626:
1624:
1620:
1616:
1611:
1599:
1598:
1594:
1592:
1590:
1585:
1583:
1582:
1578:
1577:
1575:
1573:
1572:
1567:
1561:
1559:
1554:
1552:
1550:
1549:M. yumanensis
1547:Yuma myotis (
1545:
1543:
1541:
1536:
1534:
1532:
1527:
1525:
1523:
1518:
1516:
1515:
1511:
1509:
1507:
1504:Cave myotis (
1502:
1500:
1498:
1493:
1491:
1489:
1488:M. thysanodes
1484:
1482:
1481:
1477:
1475:
1473:
1468:
1466:
1464:
1461:Indiana bat (
1459:
1457:
1455:
1450:
1448:
1446:
1441:
1439:
1437:
1432:
1430:
1429:
1425:
1423:
1421:
1416:
1414:
1412:
1407:
1405:
1403:
1398:
1396:
1394:
1389:
1387:
1385:
1380:
1378:
1376:
1371:
1369:
1367:
1362:
1360:
1358:
1353:
1351:
1349:
1344:
1342:
1340:
1335:
1333:
1331:
1326:
1324:
1323:
1322:M. phanluongi
1319:
1317:
1315:
1310:
1308:
1306:
1301:
1299:
1297:
1292:
1290:
1288:
1283:
1281:
1279:
1274:
1272:
1270:
1265:
1263:
1262:
1258:
1256:
1254:
1253:M. nipalensis
1249:
1247:
1245:
1244:M. nimbaensis
1240:
1238:
1236:
1231:
1229:
1227:
1222:
1220:
1218:
1213:
1211:
1209:
1208:M. mystacinus
1204:
1202:
1200:
1195:
1193:
1191:
1186:
1184:
1182:
1177:
1175:
1173:
1172:M. montivagus
1168:
1166:
1164:
1163:M. moluccarum
1159:
1157:
1156:
1152:
1150:
1148:
1143:
1141:
1139:
1134:
1132:
1130:
1125:
1123:
1122:
1118:
1116:
1114:
1109:
1107:
1105:
1100:
1098:
1096:
1091:
1089:
1087:
1082:
1080:
1078:
1073:
1071:
1070:
1066:
1064:
1062:
1057:
1055:
1053:
1048:
1046:
1044:
1039:
1037:
1036:
1035:M. izecksohni
1032:
1030:
1028:
1023:
1021:
1020:
1016:
1014:
1012:
1011:M. ikonnikovi
1007:
1005:
1004:
1003:M. hyrcanicus
1000:
998:
996:
991:
989:
987:
982:
980:
978:
973:
971:
970:
966:
964:
962:
957:
955:
953:
952:M. grisescens
948:
946:
944:
939:
937:
935:
930:
928:
926:
921:
919:
917:
912:
910:
908:
903:
901:
900:
896:
894:
892:
887:
885:
884:
883:M. fimbriatus
880:
878:
876:
871:
869:
868:
864:
862:
860:
855:
853:
851:
846:
844:
842:
837:
835:
834:
830:
828:
826:
821:
819:
817:
812:
810:
808:
803:
801:
799:
794:
792:
790:
785:
783:
781:
776:
774:
772:
771:M. cobanensis
767:
765:
763:
758:
756:
754:
749:
747:
745:
744:M. chiloensis
740:
738:
736:
735:M. capaccinii
731:
729:
727:
722:
720:
718:
713:
711:
709:
704:
702:
700:
695:
693:
691:
686:
684:
682:
677:
675:
673:
668:
666:
665:
661:
659:
657:
652:
650:
648:
643:
641:
639:
634:
632:
630:
625:
623:
621:
616:
614:
612:
607:
605:
603:
598:
596:
594:
589:
587:
586:
585:M. annatessae
582:
580:
578:
573:
571:
569:
564:
562:
560:
555:
553:
551:
546:
544:
542:
537:
535:
533:
528:
526:
524:
519:
518:
516:
514:
513:
508:
502:
498:
496:
492:
490:
486:
484:
480:
478:
474:
472:
468:
467:
464:
460:
452:
447:
445:
440:
438:
433:
432:
429:
417:
414:
407:
404:
398:
396:
392:
379:
374:
370:
366:
365:
360:
358:
349:
347:
345:
343:
339:
332:
330:
328:
324:
320:
316:
308:
306:
304:
300:
296:
292:
284:
282:
280:
276:
273:It roosts in
268:
266:
253:
249:
240:
238:
235:
233:
229:
225:
221:
213:
211:
209:
205:
201:
200:
195:
185:
180:
173:
171:
165:
162:
161:Binomial name
158:
154:
153:
148:
145:
144:
141:
140:
136:
133:
132:
129:
126:
123:
122:
119:
116:
113:
112:
109:
106:
103:
102:
99:
96:
93:
92:
89:
86:
83:
82:
79:
76:
73:
72:
67:
62:
58:
52:
47:
36:
32:
27:
22:
19:
1830:Bats of Asia
1622:
1595:
1588:
1579:
1569:
1557:
1548:
1539:
1530:
1521:
1512:
1505:
1496:
1487:
1478:
1471:
1470:Kei myotis (
1462:
1453:
1444:
1435:
1428:M. sibiricus
1426:
1419:
1410:
1401:
1392:
1391:Red myotis (
1383:
1374:
1365:
1356:
1347:
1339:M. pruinosus
1338:
1336:
1330:M. planiceps
1329:
1320:
1313:
1305:M. pequinius
1304:
1295:
1286:
1277:
1268:
1259:
1252:
1243:
1235:M. nigricans
1234:
1226:M. nesopolus
1225:
1217:M. nattereri
1216:
1207:
1198:
1189:
1180:
1171:
1162:
1153:
1146:
1137:
1128:
1119:
1112:
1104:M. lucifugus
1103:
1094:
1085:
1076:
1067:
1060:
1051:
1042:
1033:
1027:M. insularum
1026:
1017:
1010:
1001:
994:
985:
977:M. hasseltii
976:
967:
960:
951:
942:
933:
924:
916:M. fortidens
915:
906:
897:
890:
881:
874:
867:M. escalerai
865:
858:
849:
840:
833:M. diminutus
831:
824:
815:
806:
798:M. dasycneme
797:
788:
779:
770:
761:
753:M. chinensis
752:
743:
734:
725:
716:
707:
698:
689:
680:
671:
662:
655:
646:
638:M. australis
637:
629:M. auriculus
628:
619:
610:
601:
593:M. annectans
592:
583:
576:
567:
558:
549:
541:M. albescens
540:
531:
522:
510:
416:
406:
382:. Retrieved
368:
362:
356:
312:
309:Conservation
288:
275:tree hollows
272:
244:
236:
217:
198:
197:
193:
191:
169:
167:
151:
150:
138:
18:
1688:iNaturalist
1647:Wikispecies
1571:Submyotodon
1497:M. tricolor
1472:M. stalkeri
1436:M. sicarius
1375:M. riparius
1357:M. ricketti
1269:M. occultus
1190:M. muricola
1121:M. macropus
1095:M. longipes
969:M. handleyi
907:M. formosus
891:M. findleyi
708:M. brandtii
699:M. bombinus
664:M. bartelsi
559:M. altarium
550:M. alcathoe
523:M. adversus
499:Subfamily:
384:15 November
301:islands of
241:Description
1814:Categories
1506:M. velifer
1463:M. sodalis
1402:M. schaubi
1384:M. rosseti
1366:M. ridleyi
1348:M. punicus
1287:M. oxyotus
1181:M. morrisi
1061:M. laniger
986:M. hermani
950:Gray bat (
943:M. goudoti
850:M. elegans
825:M. dieteri
816:M. davidii
796:Pond bat (
789:M. csorbai
690:M. bocagii
681:M. blythii
532:M. aelleni
489:Chiroptera
333:References
315:endangered
204:vesper bat
118:Chiroptera
46:Endangered
1522:M. volans
1514:M. vivesi
1411:M. scotti
1278:M. oreias
1261:M. nyctor
1199:M. myotis
1077:M. leibii
1069:M. lavali
1052:M. keenii
1043:M. keaysi
925:M. frater
899:M. flavus
875:M. evotis
656:M. badius
469:Kingdom:
220:described
146:Species:
84:Kingdom:
78:Eukaryota
1758:13802529
1706:10598067
1632:Wikidata
1454:M. simus
1393:M. ruber
1314:M. petax
1086:M. levis
501:Myotinae
493:Family:
483:Mammalia
477:Chordata
475:Phylum:
471:Animalia
459:Myotinae
411:297-307.
226:and the
214:Taxonomy
124:Family:
108:Mammalia
98:Chordata
94:Phylum:
88:Animalia
74:Domain:
51:IUCN 3.1
1745:1005459
1680:2432426
1638:Q644680
611:M. ater
487:Order:
481:Class:
317:by the
295:Shikoku
279:diploid
269:Biology
261:3.1.3.3
258:2.1.3.3
134:Genus:
114:Order:
104:Class:
49: (
1797:423040
1771:196298
1719:632046
1667:327614
512:Myotis
299:Kyushu
297:, and
291:Honshu
248:tragus
139:Myotis
1784:85641
1732:14192
1701:IRMNG
1693:40302
303:Japan
232:Japan
208:Japan
1766:NCBI
1727:IUCN
1714:ITIS
1675:GBIF
386:2021
369:2019
325:and
319:IUCN
192:The
1753:MSW
1740:MDD
1662:EoL
373:doi
254:is
1816::
1794::
1781::
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1755::
1742::
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1634::
394:^
367:.
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341:^
329:.
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450:e
443:t
436:v
388:.
375::
359:"
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196:(
53:)
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