31:
75:
196:
50:
318:
has sensitive hearing and sense of smell and good eyesight which helps it to manoeuvre well during flight, specifically at night. What makes it different from other fruit bats is its echolocating ability. It can be distinguished by its grey-brown to brown upperpart which is darker on top of the head
331:
The Sabah Museum specimens had forearm length measurements between 82 and 876 millimetres (3.2 and 34.5 in) for the females and an adult male from
Sarawak had forearms 81 millimetres (3.2 in) long. The external measurements are within the range of 78 to 87 millimetres (3.1 to 3.4 in)
323:
females. The most distinguishable figure of this bat besides producing a distinctive, audible clicking call is its wings. It is attached to the sides of the back and separated by a broad band of fur. The lower incisors are bifid, the canines have a longitudinal groove on the outer surface which is
319:
and paler underpart which is usually grey-brown. It has long pale hairs on the chin and neck despite having short and sparse fur. It sometimes has pale yellow tufts of hair on the side of its neck which occur in adult for this species, especially males. Most males are
30:
388:
Waldien, D.L.; Wilson, Z.; Adleson, S.; Abdul Aziz, S.; Bates, P.J.J.; Bumrungsri, S.; Furey, N.; Ingle, N.R.; Mildenstein, T.; Phelps, K.; Tanalgo, K.; Soisook, P.; Thong, V.D.; Wiantoro, S.; Tsang, S.M. (2019).
488:
3687:
3491:
3556:
3597:
492:
3465:
3100:
3647:
3504:
3677:
3211:
3076:
3067:
541:
360:. This medium-sized bat normally roosts in caves, and feeds on fruit, nectar, and pollen. It roosts dark caves, rock crevices and old tombs.
2993:
3637:
1405:
1396:
3642:
2925:
1450:
3682:
3011:
2961:
1495:
1003:
639:
2943:
1414:
3202:
185:
3530:
3002:
2907:
2898:
2431:
1423:
1021:
675:
3607:
2608:
1805:
1504:
1468:
1441:
738:
693:
3548:
2934:
2599:
1486:
1477:
789:
3509:
3662:
3622:
2154:
2005:
1969:
1856:
1459:
1161:
1012:
666:
3407:
3652:
3617:
2916:
2889:
1978:
1095:
967:
720:
711:
3257:
3692:
3657:
3632:
3543:
3426:
3303:
1565:
1432:
780:
702:
684:
648:
1062:
2952:
2503:
2235:
2100:
1628:
1053:
756:
657:
534:
74:
3561:
3672:
3220:
2768:
1687:
3627:
3612:
2557:
2307:
1996:
1987:
1897:
1528:
1170:
1152:
904:
880:
514:
2512:
2217:
2181:
1119:
976:
3667:
3294:
2404:
2136:
2064:
1945:
1921:
1847:
1737:
1646:
1637:
1595:
1283:
1250:
3369:
2736:
2413:
2316:
2199:
2091:
1604:
1259:
935:
914:
747:
3431:
2485:
2467:
2127:
1728:
1678:
1317:
729:
527:
169:
3336:
1364:
985:
3452:
3312:
2813:
2795:
2494:
2458:
2343:
2271:
2262:
2037:
1888:
299:
345:
2822:
2684:
2530:
2440:
2325:
2109:
1787:
413:
39:
3602:
2759:
2666:
2521:
2449:
2380:
2253:
2163:
2073:
2046:
1719:
1143:
69:
3496:
3569:
3439:
3170:
3161:
3035:
2750:
2539:
2395:
2370:
2352:
2298:
2289:
2280:
2244:
2145:
2118:
1814:
1796:
1342:
622:
320:
3574:
3137:
2777:
2723:
2575:
2566:
2548:
2476:
2422:
2386:
2334:
2208:
2172:
2055:
1823:
994:
847:
408:
364:
303:
195:
3522:
2786:
2675:
2361:
2190:
1871:
1537:
1519:
1307:
1218:
1194:
1071:
856:
509:
295:
519:
3275:
2226:
2082:
587:
243:
3591:
3517:
3327:
3152:
3128:
3050:
2850:
2808:
1086:
895:
771:
399:
390:
349:
59:
54:
3444:
3285:
3118:
3091:
3026:
2862:
2590:
1765:
1702:
1619:
1578:
871:
817:
609:
341:
3535:
3478:
3401:
3185:
2872:
2636:
1233:
1185:
1036:
368:
291:
3392:
2649:
1960:
1838:
1661:
1298:
950:
593:
287:
3238:
2699:
1936:
1355:
1274:
1209:
1134:
830:
357:
279:
275:
227:
146:
86:
3418:
3386:
3248:
2020:
1912:
1778:
1379:
1110:
926:
581:
575:
569:
563:
325:
263:
259:
106:
3470:
2976:
551:
353:
271:
255:
251:
247:
223:
219:
136:
3483:
2718:
283:
116:
96:
3363:
328:
are smaller than second premolars, especially on the upper jaw.
267:
3457:
3367:
2847:
2633:
1762:
1562:
1339:
814:
606:
523:
126:
460:. Washington, D.C.: Conservation International. pp. 151-155.
439:
437:
435:
433:
431:
3376:
3325:
3283:
3274:
3246:
3237:
3183:
3150:
3126:
3117:
3089:
3048:
3024:
2974:
2870:
2861:
2697:
2647:
2588:
2018:
1958:
1934:
1910:
1869:
1836:
1776:
1700:
1659:
1617:
1576:
1517:
1377:
1353:
1296:
1272:
1231:
1207:
1183:
1132:
1108:
1084:
1034:
948:
924:
893:
869:
828:
769:
620:
414:10.2305/IUCN.UK.2019-3.RLTS.T19754A22001514.en
535:
8:
3688:Taxa named by Étienne Geoffroy Saint-Hilaire
344:were collected from coconut plantations on
3364:
3280:
3243:
3123:
2867:
2858:
2844:
2644:
2630:
1773:
1759:
1573:
1559:
1350:
1336:
825:
811:
617:
603:
542:
528:
520:
194:
48:
29:
20:
412:
371:has the largest gathering of these bats.
324:slightly medial to center, and the first
3212:East African little collared fruit bat (
230:. It is one of ten species in the genus
3101:Veldkamp's dwarf epauletted fruit bat (
380:
445:A Field Guide to the Mammals of Borneo
443:Payne, J. & Francis, C.M. (1985).
242:Geoffroy's rousette occurs throughout
3068:Hayman's dwarf epauletted fruit bat (
2926:Lesser Angolan epauletted fruit bat (
491:. National Geographic. Archived from
7:
3077:Peters' dwarf epauletted fruit bat (
2994:Buettikofer's epauletted fruit bat (
1496:New Britain naked-backed fruit bat (
1004:Nusatenggara short-nosed fruit bat (
640:Broad-striped tube-nosed fruit bat (
3598:IUCN Red List least concern species
2944:East African epauletted fruit bat (
1415:Philippine naked-backed fruit bat (
1406:Beaufort's naked-backed fruit bat (
1397:Andersen's naked-backed fruit bat (
400:IUCN Red List of Threatened Species
2432:Little golden-mantled flying fox (
1451:Solomon's naked-backed fruit bat (
1424:Halmahera naked-backed fruit bat (
1022:Indonesian short-nosed fruit bat (
676:Round-eared tube-nosed fruit bat (
447:. Sabah Society: Malaysia. p. 171.
14:
3012:Franquet's epauletted fruit bat (
2962:Wahlberg's epauletted fruit bat (
2609:Sulawesi stripe-faced fruit bat (
1806:Giant golden-crowned flying fox (
1505:Greenish naked-backed fruit bat (
1469:Moluccan naked-backed fruit bat (
1442:Sulawesi naked-backed fruit bat (
739:Philippine tube-nosed fruit bat (
515:Rousettus on Animal Diversity Web
2935:Ethiopian epauletted fruit bat (
2600:Mindoro stripe-faced fruit bat (
1487:Western naked-backed fruit bat (
1478:Panniet naked-backed fruit bat (
790:Steadfast tube-nosed fruit bat (
73:
2908:Peters's epauletted fruit bat (
2899:Ansell's epauletted fruit bat (
2155:New Ireland masked flying fox (
2006:New Georgian monkey-faced Bat (
1970:Bougainville monkey-faced Bat (
1857:Small white-winged flying fox (
1460:Lesser naked-backed fruit bat (
1013:Greater short-nosed fruit bat (
667:Mountain tube-nosed fruit bat (
3648:Mammals of the Solomon Islands
2917:Gambian epauletted fruit bat (
2890:Angolan epauletted fruit bat (
1979:Guadalcanal monkey-faced Bat (
968:Lesser short-nosed fruit bat (
721:Demonic tube-nosed fruit bat (
712:Malaita tube-nosed fruit bat (
694:Keast's tube-nosed fruit bat (
332:recorded in previous studies.
1:
3678:Mammals of Western New Guinea
3203:São Tomé collared fruit bat (
1433:Biak naked-backed fruit bat (
781:Lesser tube-nosed fruit bat (
703:Island tube-nosed fruit bat (
685:Dragon tube-nosed fruit bat (
649:Common tube-nosed fruit bat (
3193:(Little collared fruit bats)
2953:Minor epauletted fruit bat (
2504:Philippine gray flying fox (
2101:Bismarck masked flying fox (
1162:Ratanaworabhan's fruit bat (
757:Umboi tube-nosed fruit bat (
3638:Mammals of Papua New Guinea
3221:Little collared fruit bat (
1879:(Straw-coloured fruit bats)
1688:New Caledonia blossom bat (
1096:Fischer's pygmy fruit bat (
478:(Lekagul and McNeely 1977).
3709:
3643:Mammals of the Philippines
2558:New Caledonia flying fox (
2308:Black-bearded flying fox (
1997:Montane monkey-faced Bat (
1988:Greater monkey-faced Bat (
1898:Straw-coloured fruit bat (
1529:Sulawesi harpy fruit bat (
1171:White-collared fruit bat (
1153:Javan tailless fruit bat (
1063:Rickart's dyak fruit bat (
905:Spotted-winged fruit bat (
881:Mindanao pygmy fruit bat (
489:"Escape from the Bat Cave"
203:Geoffroy's Rousette range
3683:Mammals described in 1810
3408:Rousettus amplexicaudatus
3378:Rousettus amplexicaudatus
3295:Short-palated fruit bat (
2857:
2843:
2643:
2629:
2405:Geelvink Bay flying fox (
2236:Livingstone's fruit bat (
2137:Large Samoan flying fox (
2065:Small Samoan flying fox (
1946:Small-toothed fruit bat (
1922:Fijian monkey-faced Bat (
1848:White-winged flying fox (
1772:
1758:
1738:Moss-forest blossom bat (
1638:Black-bellied fruit bat (
1629:Fardoulis's blossom bat (
1596:Long-tongued nectar bat (
1586:(Long-tongued fruit bats)
1572:
1558:
1349:
1335:
1251:Greater musky fruit bat (
1054:Brooks's dyak fruit bat (
824:
810:
658:Pallas's tube-nosed bat (
616:
602:
558:
550:Extant species of family
456:Bonaccorso, F.J. (1998).
393:Rousettus amplexicaudatus
321:substantially larger than
215:Rousettus amplexicaudatus
202:
193:
179:Rousettus amplexicaudatus
175:
168:
70:Scientific classification
68:
46:
37:
28:
23:
3258:D'Anchieta's fruit bat (
2769:Leschenault's rousette (
2414:Grey-headed flying fox (
2317:Black-eared flying fox (
2200:Ontong Java flying fox (
2092:Ashy-headed flying fox (
1669:(Long-tailed fruit bats)
1605:Long-tongued fruit bat (
1387:(Bare-backed fruit bats)
1260:Lesser musky fruit bat (
936:Black-capped fruit bat (
748:Eastern tube-nosed Bat (
458:Bats of Papua New Guinea
3304:Campo-Ma’an fruit bat (
2513:Temminck's flying fox (
2486:Little red flying fox (
2468:Madagascan flying fox (
2218:Andersen's flying fox (
2182:Gilliard's flying fox (
2128:Spectacled flying fox (
1729:Halmahera blossom bat (
1679:Long-tailed fruit bat (
1120:Salim Ali's fruit bat (
977:Horsfield's fruit bat (
730:Lesser tube-nosed bat (
630:(tube-nosed fruit bats)
314:Like other fruit bats,
161:R. amplexicaudatus
3608:Bats of Southeast Asia
3058:(Dwarf epauleted bats)
2880:(Epauleted fruit bats)
2814:Long-haired rousette (
2796:Bare-backed rousette (
2495:Seychelles fruit bat (
2459:Rodrigues flying fox (
2344:Mauritian flying fox (
2272:Big-eared flying fox (
2038:Admiralty flying fox (
1889:Madagascan fruit bat (
1647:Woodford's fruit bat (
1284:Blanford's fruit bat (
958:(Dog-faced fruit bats)
915:Balionycteris seimundi
2823:Madagascan rousette (
2742:Geoffroy's rousette (
2707:(rousette fruit bats)
2685:Philippine dawn bat (
2531:Vanikoro flying fox (
2441:Solomons flying fox (
2326:Caroline flying fox (
2110:Moluccan flying fox (
1788:Sulawesi flying fox (
407:: e.T19754A22001514.
352:, while the one from
3663:Mammals of Singapore
3623:Mammals of Indonesia
3337:Zenker's fruit bat (
3003:Dobson's fruit bat (
2667:Greater nectar bat (
2522:Insular flying fox (
2450:Rennell flying fox (
2254:Okinawa flying fox (
2164:Nicobar flying fox (
2074:Vanuatu flying fox (
2047:Aldabra flying fox (
1720:Common blossom bat (
1365:Bulmer's fruit bat (
1144:Tailless fruit bat (
986:Peters's fruit bat (
469:(Payne et al. 1985).
348:and the highland of
300:Bismarck Archipelago
24:Geoffroy's rousette
3653:Mammals of Thailand
3618:Mammals of Cambodia
3313:Pohle's fruit bat (
3162:Azagnyi fruit bat (
3036:Hammer-headed Bat (
2825:R. madagascariensis
2760:Egyptian rousette (
2751:Sulawesi rousette (
2540:Kosrae flying fox (
2396:Masked flying fox (
2371:Ornate flying fox (
2353:Temotu flying fox (
2299:Indian flying fox (
2290:Mariana fruit bat (
2281:Lesser flying fox (
2263:Lyle's flying fox (
2245:Lombok flying fox (
2146:Ryukyu flying fox (
2119:Makira flying fox (
1859:D. microleucopterus
1815:Palawan fruit bat (
1797:Talaud flying fox (
336:Biology and ecology
210:Geoffroy's rousette
40:Conservation status
3693:Bats of New Guinea
3658:Mammals of Vietnam
3633:Mammals of Myanmar
3138:Angolan rousette (
2778:Linduan Rousette (
2744:R. amplexicaudatus
2724:Manado fruit bat (
2576:Dwarf flying fox (
2567:Pemba flying fox (
2549:Large flying fox (
2477:Samoa flying fox (
2423:Bonin flying fox (
2387:Pelew flying fox (
2335:Great flying fox (
2209:Small flying fox (
2173:Banks flying fox (
2056:Black flying fox (
1824:Sunda flying fox (
1241:(Musky fruit bats)
1044:(Dayak fruit bats)
995:Minute fruit bat (
848:Borneo fruit bat (
838:(Pygmy fruit bats)
316:R. amplexicaudatus
3585:
3584:
3570:Open Tree of Life
3370:Taxon identifiers
3361:
3360:
3357:
3356:
3353:
3352:
3349:
3348:
3306:C. campomaanensis
3270:
3269:
3233:
3232:
3196:
3113:
3112:
3061:
2987:
2883:
2839:
2838:
2835:
2834:
2787:Comoro rousette (
2710:
2676:Cave nectar bat (
2660:
2657:(Dawn fruit bats)
2625:
2624:
2621:
2620:
2362:Ceram fruit bat (
2191:Gray flying fox (
2130:P. conspicillatus
2031:
1882:
1754:
1753:
1750:
1749:
1713:
1672:
1589:
1554:
1553:
1550:
1549:
1538:Harpy fruit bat (
1390:
1343:Harpyionycterinae
1331:
1330:
1327:
1326:
1308:Swift fruit bat (
1244:
1219:Dusky fruit bat (
1197:O. cartilagonodus
1195:Luzon fruit bat (
1072:Dayak fruit bat (
1047:
1024:C. titthaecheilus
961:
938:C. melanocephalus
857:Pygmy fruit bat (
841:
806:
805:
802:
801:
633:
495:on April 7, 2011.
340:Specimens in the
207:
206:
63:
3700:
3673:Mammals of Timor
3578:
3577:
3565:
3564:
3552:
3551:
3539:
3538:
3526:
3525:
3513:
3512:
3500:
3499:
3487:
3486:
3474:
3473:
3461:
3460:
3448:
3447:
3435:
3434:
3422:
3421:
3412:
3411:
3410:
3397:
3396:
3395:
3365:
3281:
3244:
3205:M. brachycephala
3195:
3194:
3190:
3171:Woermann's bat (
3124:
3060:
3059:
3055:
2986:
2985:
2984:(Epauleted bats)
2981:
2882:
2881:
2877:
2868:
2859:
2845:
2709:
2708:
2704:
2659:
2658:
2654:
2645:
2631:
2497:P. seychellensis
2416:P. poliocephalus
2337:P. neohibernicus
2227:Kei flying fox (
2112:P. chrysoproctus
2083:Aru flying fox (
2040:P. admiralitatum
2030:
2029:
2025:
1881:
1880:
1876:
1774:
1760:
1712:
1711:
1707:
1690:N. neocaledonica
1671:
1670:
1666:
1588:
1587:
1583:
1574:
1560:
1389:
1388:
1384:
1351:
1337:
1243:
1242:
1238:
1046:
1045:
1041:
960:
959:
955:
840:
839:
835:
826:
812:
632:
631:
627:
618:
604:
544:
537:
530:
521:
497:
496:
485:
479:
476:
470:
467:
461:
454:
448:
441:
426:
425:
423:
421:
416:
385:
367:in the southern
365:Monfort Bat Cave
304:Papua New Guinea
282:, the island of
198:
181:
78:
77:
57:
52:
51:
33:
21:
3708:
3707:
3703:
3702:
3701:
3699:
3698:
3697:
3628:Mammals of Laos
3613:Bats of Oceania
3588:
3587:
3586:
3581:
3573:
3568:
3560:
3555:
3547:
3542:
3534:
3529:
3521:
3516:
3508:
3503:
3495:
3490:
3482:
3477:
3469:
3464:
3456:
3451:
3443:
3438:
3430:
3425:
3417:
3415:
3406:
3405:
3400:
3391:
3390:
3385:
3372:
3362:
3345:
3321:
3266:
3229:
3192:
3191:
3189:
3179:
3146:
3109:
3085:
3057:
3056:
3054:
3044:
3020:
2996:E. buettikoferi
2983:
2982:
2980:
2970:
2879:
2878:
2876:
2853:
2831:
2771:R. leschenaulti
2706:
2705:
2703:
2693:
2656:
2655:
2653:
2639:
2617:
2584:
2533:P. tuberculatus
2355:P. nitendiensis
2238:P. livingstonii
2184:P. gilliardorum
2027:
2026:
2024:
2014:
1954:
1930:
1906:
1878:
1877:
1875:
1865:
1832:
1768:
1746:
1709:
1708:
1706:
1696:
1668:
1667:
1665:
1655:
1613:
1585:
1584:
1582:
1568:
1546:
1520:Harpyionycteris
1513:
1480:D. pannietensis
1386:
1385:
1383:
1373:
1345:
1323:
1292:
1268:
1240:
1239:
1237:
1227:
1203:
1179:
1128:
1104:
1080:
1043:
1042:
1040:
1030:
1006:C. nusatenggara
957:
956:
954:
944:
920:
889:
883:A. paucidentata
865:
837:
836:
834:
820:
798:
765:
629:
628:
626:
612:
598:
554:
548:
506:
501:
500:
487:
486:
482:
477:
473:
468:
464:
455:
451:
442:
429:
419:
417:
387:
386:
382:
377:
338:
312:
296:Solomon Islands
240:
189:
183:
177:
164:
72:
64:
53:
49:
42:
17:
12:
11:
5:
3706:
3704:
3696:
3695:
3690:
3685:
3680:
3675:
3670:
3668:Bats of Borneo
3665:
3660:
3655:
3650:
3645:
3640:
3635:
3630:
3625:
3620:
3615:
3610:
3605:
3600:
3590:
3589:
3583:
3582:
3580:
3579:
3566:
3553:
3540:
3527:
3514:
3501:
3488:
3475:
3462:
3449:
3436:
3423:
3413:
3398:
3382:
3380:
3374:
3373:
3368:
3359:
3358:
3355:
3354:
3351:
3350:
3347:
3346:
3344:
3343:
3333:
3331:
3323:
3322:
3320:
3319:
3310:
3301:
3291:
3289:
3278:
3276:Scotonycterini
3272:
3271:
3268:
3267:
3265:
3264:
3254:
3252:
3241:
3235:
3234:
3231:
3230:
3228:
3227:
3218:
3209:
3199:
3197:
3181:
3180:
3178:
3177:
3168:
3158:
3156:
3148:
3147:
3145:
3144:
3134:
3132:
3121:
3115:
3114:
3111:
3110:
3108:
3107:
3097:
3095:
3087:
3086:
3084:
3083:
3074:
3070:M. intermedius
3064:
3062:
3046:
3045:
3043:
3042:
3032:
3030:
3022:
3021:
3019:
3018:
3009:
3000:
2990:
2988:
2972:
2971:
2969:
2968:
2959:
2950:
2941:
2932:
2923:
2914:
2905:
2896:
2886:
2884:
2865:
2855:
2854:
2848:
2841:
2840:
2837:
2836:
2833:
2832:
2830:
2829:
2820:
2803:
2802:
2793:
2784:
2775:
2766:
2762:R. aegyptiacus
2757:
2748:
2731:
2730:
2713:
2711:
2695:
2694:
2692:
2691:
2682:
2673:
2663:
2661:
2641:
2640:
2634:
2627:
2626:
2623:
2622:
2619:
2618:
2616:
2615:
2606:
2602:S. mindorensis
2596:
2594:
2586:
2585:
2583:
2582:
2573:
2564:
2555:
2546:
2537:
2528:
2519:
2510:
2501:
2492:
2483:
2474:
2465:
2461:P. rodricensis
2456:
2447:
2438:
2429:
2420:
2411:
2402:
2393:
2384:
2377:
2368:
2359:
2350:
2341:
2332:
2323:
2314:
2310:P. melanopogon
2305:
2296:
2287:
2278:
2269:
2260:
2256:P. loochoensis
2251:
2247:P. lombocensis
2242:
2233:
2224:
2220:P. intermedius
2215:
2211:P. hypomelanus
2206:
2197:
2188:
2179:
2170:
2161:
2152:
2143:
2134:
2125:
2116:
2107:
2103:P. capistratus
2098:
2089:
2080:
2071:
2062:
2053:
2049:P. aldabrensis
2044:
2034:
2032:
2028:(flying foxes)
2016:
2015:
2013:
2012:
2003:
1994:
1985:
1976:
1966:
1964:
1956:
1955:
1953:
1952:
1942:
1940:
1932:
1931:
1929:
1928:
1918:
1916:
1908:
1907:
1905:
1904:
1895:
1885:
1883:
1867:
1866:
1864:
1863:
1854:
1850:D. leucopterus
1844:
1842:
1834:
1833:
1831:
1830:
1821:
1812:
1803:
1794:
1784:
1782:
1770:
1769:
1763:
1756:
1755:
1752:
1751:
1748:
1747:
1745:
1744:
1735:
1726:
1716:
1714:
1710:(blossom bats)
1698:
1697:
1695:
1694:
1685:
1675:
1673:
1657:
1656:
1654:
1653:
1644:
1635:
1625:
1623:
1615:
1614:
1612:
1611:
1602:
1592:
1590:
1570:
1569:
1566:Macroglossinae
1563:
1556:
1555:
1552:
1551:
1548:
1547:
1545:
1544:
1535:
1525:
1523:
1515:
1514:
1512:
1511:
1502:
1493:
1484:
1475:
1471:D. moluccensis
1466:
1457:
1448:
1439:
1430:
1421:
1412:
1403:
1393:
1391:
1375:
1374:
1372:
1371:
1361:
1359:
1347:
1346:
1340:
1333:
1332:
1329:
1328:
1325:
1324:
1322:
1321:
1314:
1304:
1302:
1294:
1293:
1291:
1290:
1280:
1278:
1270:
1269:
1267:
1266:
1257:
1247:
1245:
1229:
1228:
1226:
1225:
1215:
1213:
1205:
1204:
1202:
1201:
1191:
1189:
1181:
1180:
1178:
1177:
1168:
1159:
1150:
1140:
1138:
1130:
1129:
1127:
1126:
1116:
1114:
1106:
1105:
1103:
1102:
1092:
1090:
1082:
1081:
1079:
1078:
1069:
1060:
1050:
1048:
1032:
1031:
1029:
1028:
1019:
1010:
1001:
992:
988:C. luzoniensis
983:
979:C. horsfieldii
974:
964:
962:
946:
945:
943:
942:
932:
930:
922:
921:
919:
918:
911:
901:
899:
891:
890:
888:
887:
877:
875:
867:
866:
864:
863:
854:
844:
842:
822:
821:
815:
808:
807:
804:
803:
800:
799:
797:
796:
787:
777:
775:
767:
766:
764:
763:
754:
745:
736:
727:
718:
714:N. malaitensis
709:
700:
691:
682:
673:
664:
655:
646:
636:
634:
614:
613:
607:
600:
599:
597:
596:
590:
588:Laurasiatheria
584:
578:
572:
566:
559:
556:
555:
549:
547:
546:
539:
532:
524:
518:
517:
512:
505:
504:External links
502:
499:
498:
480:
471:
462:
449:
427:
379:
378:
376:
373:
346:Mantani Island
337:
334:
311:
308:
244:Southeast Asia
239:
236:
205:
204:
200:
199:
191:
190:
184:
173:
172:
166:
165:
158:
156:
152:
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144:
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129:
124:
120:
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110:
109:
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100:
99:
94:
90:
89:
84:
80:
79:
66:
65:
47:
44:
43:
38:
35:
34:
26:
25:
16:Species of bat
15:
13:
10:
9:
6:
4:
3:
2:
3705:
3694:
3691:
3689:
3686:
3684:
3681:
3679:
3676:
3674:
3671:
3669:
3666:
3664:
3661:
3659:
3656:
3654:
3651:
3649:
3646:
3644:
3641:
3639:
3636:
3634:
3631:
3629:
3626:
3624:
3621:
3619:
3616:
3614:
3611:
3609:
3606:
3604:
3601:
3599:
3596:
3595:
3593:
3576:
3571:
3567:
3563:
3558:
3554:
3550:
3545:
3541:
3537:
3532:
3528:
3524:
3519:
3515:
3511:
3506:
3502:
3498:
3493:
3489:
3485:
3480:
3476:
3472:
3467:
3463:
3459:
3454:
3450:
3446:
3441:
3437:
3433:
3428:
3424:
3420:
3414:
3409:
3403:
3399:
3394:
3388:
3384:
3383:
3381:
3379:
3375:
3371:
3366:
3342:
3340:
3335:
3334:
3332:
3330:
3329:
3328:Scotonycteris
3324:
3318:
3316:
3311:
3309:
3307:
3302:
3300:
3298:
3293:
3292:
3290:
3288:
3287:
3282:
3279:
3277:
3273:
3263:
3261:
3256:
3255:
3253:
3251:
3250:
3245:
3242:
3240:
3236:
3226:
3224:
3219:
3217:
3215:
3210:
3208:
3206:
3201:
3200:
3198:
3188:
3187:
3182:
3176:
3174:
3169:
3167:
3165:
3160:
3159:
3157:
3155:
3154:
3153:Megaloglossus
3149:
3143:
3141:
3140:L. angolensis
3136:
3135:
3133:
3131:
3130:
3129:Lissonycteris
3125:
3122:
3120:
3116:
3106:
3104:
3103:N. veldkampii
3099:
3098:
3096:
3094:
3093:
3088:
3082:
3080:
3075:
3073:
3071:
3066:
3065:
3063:
3053:
3052:
3051:Micropteropus
3047:
3041:
3039:
3038:H. monstrosus
3034:
3033:
3031:
3029:
3028:
3023:
3017:
3015:
3010:
3008:
3006:
3001:
2999:
2997:
2992:
2991:
2989:
2979:
2978:
2973:
2967:
2965:
2960:
2958:
2956:
2951:
2949:
2947:
2942:
2940:
2938:
2933:
2931:
2929:
2924:
2922:
2920:
2915:
2913:
2911:
2906:
2904:
2902:
2897:
2895:
2893:
2892:E. angolensis
2888:
2887:
2885:
2875:
2874:
2869:
2866:
2864:
2860:
2856:
2852:
2851:Epomophorinae
2846:
2842:
2828:
2826:
2821:
2819:
2817:
2811:
2810:
2809:Stenonycteris
2805:
2804:
2801:
2799:
2798:R. spinalatus
2794:
2792:
2790:
2789:R. obliviosus
2785:
2783:
2781:
2780:R. linduensis
2776:
2774:
2772:
2767:
2765:
2763:
2758:
2756:
2754:
2753:R. celebensis
2749:
2747:
2745:
2739:
2738:
2733:
2732:
2729:
2727:
2721:
2720:
2715:
2714:
2712:
2702:
2701:
2696:
2690:
2688:
2683:
2681:
2679:
2674:
2672:
2670:
2665:
2664:
2662:
2652:
2651:
2646:
2642:
2638:
2632:
2628:
2614:
2612:
2607:
2605:
2603:
2598:
2597:
2595:
2593:
2592:
2587:
2581:
2579:
2574:
2572:
2570:
2569:P. voeltzkowi
2565:
2563:
2561:
2556:
2554:
2552:
2547:
2545:
2543:
2538:
2536:
2534:
2529:
2527:
2525:
2520:
2518:
2516:
2515:P. temminckii
2511:
2509:
2507:
2502:
2500:
2498:
2493:
2491:
2489:
2488:P. scapulatus
2484:
2482:
2480:
2475:
2473:
2471:
2466:
2464:
2462:
2457:
2455:
2453:
2448:
2446:
2444:
2439:
2437:
2435:
2430:
2428:
2426:
2421:
2419:
2417:
2412:
2410:
2408:
2403:
2401:
2399:
2398:P. personatus
2394:
2392:
2390:
2389:P. pelewensis
2385:
2383:
2382:
2378:
2376:
2374:
2369:
2367:
2365:
2360:
2358:
2356:
2351:
2349:
2347:
2342:
2340:
2338:
2333:
2331:
2329:
2328:P. molossinus
2324:
2322:
2320:
2315:
2313:
2311:
2306:
2304:
2302:
2297:
2295:
2293:
2288:
2286:
2284:
2279:
2277:
2275:
2270:
2268:
2266:
2261:
2259:
2257:
2252:
2250:
2248:
2243:
2241:
2239:
2234:
2232:
2230:
2225:
2223:
2221:
2216:
2214:
2212:
2207:
2205:
2203:
2198:
2196:
2194:
2189:
2187:
2185:
2180:
2178:
2176:
2171:
2169:
2167:
2162:
2160:
2158:
2153:
2151:
2149:
2148:P. dasymallus
2144:
2142:
2140:
2135:
2133:
2131:
2126:
2124:
2122:
2117:
2115:
2113:
2108:
2106:
2104:
2099:
2097:
2095:
2090:
2088:
2086:
2081:
2079:
2077:
2072:
2070:
2068:
2063:
2061:
2059:
2054:
2052:
2050:
2045:
2043:
2041:
2036:
2035:
2033:
2023:
2022:
2017:
2011:
2009:
2004:
2002:
2000:
1995:
1993:
1991:
1986:
1984:
1982:
1977:
1975:
1973:
1968:
1967:
1965:
1963:
1962:
1957:
1951:
1949:
1944:
1943:
1941:
1939:
1938:
1933:
1927:
1925:
1920:
1919:
1917:
1915:
1914:
1909:
1903:
1901:
1896:
1894:
1892:
1887:
1886:
1884:
1874:
1873:
1868:
1862:
1860:
1855:
1853:
1851:
1846:
1845:
1843:
1841:
1840:
1835:
1829:
1827:
1822:
1820:
1818:
1813:
1811:
1809:
1804:
1802:
1800:
1795:
1793:
1791:
1790:A. celebensis
1786:
1785:
1783:
1781:
1780:
1775:
1771:
1767:
1761:
1757:
1743:
1741:
1736:
1734:
1732:
1727:
1725:
1723:
1718:
1717:
1715:
1705:
1704:
1699:
1693:
1691:
1686:
1684:
1682:
1681:N. macdonaldi
1677:
1676:
1674:
1664:
1663:
1658:
1652:
1650:
1645:
1643:
1641:
1636:
1634:
1632:
1631:M. fardoulisi
1627:
1626:
1624:
1622:
1621:
1616:
1610:
1608:
1603:
1601:
1599:
1594:
1593:
1591:
1581:
1580:
1575:
1571:
1567:
1561:
1557:
1543:
1541:
1540:H. whiteheadi
1536:
1534:
1532:
1531:H. celebensis
1527:
1526:
1524:
1522:
1521:
1516:
1510:
1508:
1503:
1501:
1499:
1498:D. praedatrix
1494:
1492:
1490:
1485:
1483:
1481:
1476:
1474:
1472:
1467:
1465:
1463:
1458:
1456:
1454:
1449:
1447:
1445:
1440:
1438:
1436:
1431:
1429:
1427:
1422:
1420:
1418:
1413:
1411:
1409:
1404:
1402:
1400:
1395:
1394:
1392:
1382:
1381:
1376:
1370:
1368:
1363:
1362:
1360:
1358:
1357:
1352:
1348:
1344:
1338:
1334:
1320:
1319:
1318:T. suhaniahae
1315:
1313:
1311:
1310:T. nigrescens
1306:
1305:
1303:
1301:
1300:
1295:
1289:
1287:
1282:
1281:
1279:
1277:
1276:
1271:
1265:
1263:
1258:
1256:
1254:
1249:
1248:
1246:
1236:
1235:
1230:
1224:
1222:
1217:
1216:
1214:
1212:
1211:
1206:
1200:
1198:
1193:
1192:
1190:
1188:
1187:
1182:
1176:
1174:
1169:
1167:
1165:
1160:
1158:
1156:
1151:
1149:
1147:
1142:
1141:
1139:
1137:
1136:
1131:
1125:
1123:
1118:
1117:
1115:
1113:
1112:
1107:
1101:
1099:
1094:
1093:
1091:
1089:
1088:
1087:Haplonycteris
1083:
1077:
1075:
1070:
1068:
1066:
1061:
1059:
1057:
1052:
1051:
1049:
1039:
1038:
1033:
1027:
1025:
1020:
1018:
1016:
1011:
1009:
1007:
1002:
1000:
998:
993:
991:
989:
984:
982:
980:
975:
973:
971:
970:C. brachyotis
966:
965:
963:
953:
952:
947:
941:
939:
934:
933:
931:
929:
928:
923:
917:
916:
912:
910:
908:
903:
902:
900:
898:
897:
896:Balionycteris
892:
886:
884:
879:
878:
876:
874:
873:
868:
862:
860:
855:
853:
851:
846:
845:
843:
833:
832:
827:
823:
819:
813:
809:
795:
793:
788:
786:
784:
779:
778:
776:
774:
773:
772:Paranyctimene
768:
762:
760:
755:
753:
751:
746:
744:
742:
737:
735:
733:
728:
726:
724:
719:
717:
715:
710:
708:
706:
701:
699:
697:
692:
690:
688:
687:N. draconilla
683:
681:
679:
674:
672:
670:
665:
663:
661:
660:N. cephalotes
656:
654:
652:
651:N. albiventer
647:
645:
643:
638:
637:
635:
625:
624:
619:
615:
611:
605:
601:
595:
591:
589:
585:
583:
579:
577:
573:
571:
567:
565:
561:
560:
557:
553:
545:
540:
538:
533:
531:
526:
525:
522:
516:
513:
511:
510:IUCN Red List
508:
507:
503:
494:
490:
484:
481:
475:
472:
466:
463:
459:
453:
450:
446:
440:
438:
436:
434:
432:
428:
415:
410:
406:
402:
401:
396:
394:
384:
381:
374:
372:
370:
366:
361:
359:
355:
351:
350:Crocker Range
347:
343:
335:
333:
329:
327:
322:
317:
309:
307:
305:
301:
297:
293:
289:
285:
281:
277:
273:
269:
265:
261:
257:
253:
249:
245:
237:
235:
233:
229:
226:or Old World
225:
221:
217:
216:
211:
201:
197:
192:
187:
182:
180:
174:
171:
170:Binomial name
167:
163:
162:
157:
154:
153:
150:
149:
145:
142:
141:
138:
135:
132:
131:
128:
125:
122:
121:
118:
115:
112:
111:
108:
105:
102:
101:
98:
95:
92:
91:
88:
85:
82:
81:
76:
71:
67:
61:
56:
55:Least Concern
45:
41:
36:
32:
27:
22:
19:
3377:
3338:
3326:
3314:
3305:
3296:
3286:Casinycteris
3284:
3260:P. anchietae
3259:
3247:
3222:
3213:
3204:
3184:
3173:M. woermanni
3172:
3163:
3151:
3139:
3127:
3119:Myonycterini
3102:
3092:Nanonycteris
3090:
3078:
3069:
3049:
3037:
3027:Hypsignathus
3025:
3014:E. franqueti
3013:
3004:
2995:
2975:
2964:E. wahlbergi
2963:
2954:
2945:
2936:
2927:
2919:E. gambianus
2918:
2910:E. crypturus
2909:
2900:
2891:
2871:
2863:Epomophorini
2824:
2815:
2806:
2797:
2788:
2779:
2770:
2761:
2752:
2743:
2741:
2734:
2725:
2716:
2698:
2686:
2677:
2668:
2648:
2610:
2601:
2591:Styloctenium
2589:
2578:P. woodfordi
2577:
2568:
2559:
2550:
2541:
2532:
2523:
2514:
2506:P. speciosus
2505:
2496:
2487:
2479:P. samoensis
2478:
2469:
2460:
2451:
2442:
2433:
2425:P. pselaphon
2424:
2415:
2406:
2397:
2388:
2381:P. pelagicus
2379:
2372:
2363:
2354:
2345:
2336:
2327:
2319:P. melanotus
2318:
2309:
2300:
2292:P. mariannus
2291:
2283:P. mahaganus
2282:
2273:
2264:
2255:
2246:
2237:
2228:
2219:
2210:
2201:
2192:
2183:
2174:
2165:
2156:
2147:
2138:
2129:
2120:
2111:
2102:
2093:
2084:
2076:P. anetianus
2075:
2067:P. allenorum
2066:
2057:
2048:
2039:
2019:
2007:
1998:
1990:P. flanneryi
1989:
1980:
1971:
1959:
1947:
1935:
1924:M. acrodonta
1923:
1911:
1899:
1891:E. dupreanum
1890:
1870:
1858:
1849:
1837:
1825:
1816:
1807:
1798:
1789:
1777:
1766:Pteropodinae
1739:
1731:S. carolinae
1730:
1722:S. australis
1721:
1703:Syconycteris
1701:
1689:
1680:
1660:
1649:M. woodfordi
1648:
1639:
1630:
1620:Melonycteris
1618:
1606:
1597:
1579:Macroglossus
1577:
1539:
1530:
1518:
1506:
1497:
1488:
1479:
1470:
1461:
1452:
1443:
1434:
1426:D. crenulata
1425:
1416:
1408:D. beauforti
1407:
1399:D. anderseni
1398:
1378:
1366:
1354:
1316:
1309:
1297:
1286:S. blanfordi
1285:
1273:
1261:
1252:
1232:
1220:
1208:
1196:
1184:
1172:
1163:
1154:
1146:M. ecaudatus
1145:
1133:
1122:L. salimalii
1121:
1109:
1097:
1085:
1074:D. spadiceus
1073:
1064:
1055:
1035:
1023:
1014:
1005:
996:
987:
978:
969:
949:
937:
925:
913:
906:
894:
882:
872:Alionycteris
870:
858:
849:
829:
818:Cynopterinae
791:
782:
770:
759:N. vizcaccia
758:
750:N. robinsoni
749:
740:
731:
722:
713:
704:
695:
686:
677:
668:
659:
650:
641:
621:
610:Nyctimeninae
586:Superorder:
580:Infraclass:
552:Pteropodidae
493:the original
483:
474:
465:
457:
452:
444:
418:. Retrieved
404:
398:
392:
383:
362:
342:Sabah Museum
339:
330:
315:
313:
241:
238:Distribution
231:
214:
213:
209:
208:
178:
176:
160:
159:
147:
137:Pteropodidae
18:
3479:iNaturalist
3402:Wikispecies
3315:C. ophiodon
3297:C. argynnis
3223:M. torquata
3186:Myonycteris
3079:M. pusillus
2937:E. labiatus
2873:Epomophorus
2637:Rousettinae
2611:S. wallacei
2551:P. vampyrus
2524:P. tonganus
2452:P. rennelli
2364:P. ocularis
2274:P. macrotis
2229:P. keyensis
2202:P. howensis
2175:P. fundatus
2166:P. faunulus
2121:P. cognatus
2094:P. caniceps
2085:P. aruensis
1839:Desmalopex
1826:A. mackloti
1817:A. leucotis
1640:M. melanops
1607:M. sobrinus
1417:D. chapmani
1367:A. bulmerae
1234:Ptenochirus
1186:Otopteropus
1173:M. wetmorei
1164:M. niphanae
1155:M. kusnotoi
1098:H. fischeri
1065:D. rickarti
1037:Dyacopterus
907:B. maculata
850:A. aequalis
678:N. cyclotis
420:19 November
369:Philippines
310:Description
292:Philippines
246:and in the
186:É. Geoffroy
3592:Categories
3339:S. zenkeri
3214:M. relicta
3164:M. azagnyi
3005:E. dobsoni
2946:E. minimus
2928:E. grandis
2901:E. anselli
2849:Subfamily
2816:R. lanosus
2687:E. robusta
2678:E. spelaea
2650:Eonycteris
2635:Subfamily
2560:P. vetulus
2542:P. ualanus
2443:P. rayneri
2434:P. pumilus
2373:P. ornatus
2193:P. griseus
2157:P. ennisae
1999:P. pulchra
1961:Pteralopex
1808:A. jubatus
1799:A. humilis
1764:Subfamily
1662:Notopteris
1598:M. minimus
1564:Subfamily
1507:D. viridis
1489:D. peronii
1453:D. inermis
1444:D. exoleta
1341:Subfamily
1299:Thoopterus
1056:D. brooksi
997:C. minutus
951:Cynopterus
816:Subfamily
732:N. minutus
723:N. masalai
669:N. certans
608:Subfamily
594:Chiroptera
375:References
288:East Timor
250:region of
228:fruit bats
127:Chiroptera
3603:Rousettus
3239:Plerotini
2807:Subgenus
2737:Rousettus
2735:Subgenus
2726:R. bidens
2717:Subgenus
2700:Rousettus
2407:P. pohlei
2301:P. medius
2058:P. alecto
1981:P. atrata
1972:P. anceps
1948:N. frosti
1937:Neopteryx
1900:E. helvum
1740:S. hobbit
1435:D. emersa
1356:Aproteles
1275:Sphaerias
1253:P. jagori
1221:P. lucasi
1210:Penthetor
1135:Megaerops
1015:C. sphinx
859:A. alecto
831:Aethalops
783:P. raptor
741:N. rabori
696:N. keasti
623:Nyctimene
562:Kingdom:
358:Niah Cave
356:was from
326:premolars
280:Indonesia
276:Singapore
232:Rousettus
155:Species:
148:Rousettus
93:Kingdom:
87:Eukaryota
3549:13800404
3497:11397121
3416:BioLib:
3393:Q1443996
3387:Wikidata
3249:Plerotes
2955:E. minor
2669:E. major
2470:P. rufus
2346:P. niger
2265:P. lylei
2021:Pteropus
1913:Mirimiri
1779:Acerodon
1462:D. minor
1380:Dobsonia
1262:P. minor
1111:Latidens
927:Chironax
792:P. tenax
705:N. major
642:N. aello
582:Eutheria
576:Mammalia
570:Chordata
568:Phylum:
564:Animalia
264:Cambodia
260:Thailand
133:Family:
117:Mammalia
107:Chordata
103:Phylum:
97:Animalia
83:Domain:
60:IUCN 3.1
3536:1004540
3471:2432948
2977:Epomops
2139:P. coxi
2008:P. taki
1872:Eidolon
592:Order:
574:Class:
354:Sarawak
272:Vietnam
256:Myanmar
252:Oceania
248:Malesia
224:megabat
220:species
218:) is a
143:Genus:
123:Order:
113:Class:
58: (
3575:164530
3510:631667
3458:323422
2719:Boneia
302:, and
294:, the
290:, the
284:Borneo
188:, 1810
3562:58083
3523:19754
3492:IRMNG
3484:40848
3445:4THCK
3432:15892
3419:34011
254:, in
3557:NCBI
3518:IUCN
3505:ITIS
3466:GBIF
3427:BOLD
422:2021
405:2019
363:The
268:Laos
3544:MSW
3531:MDD
3453:EoL
3440:CoL
409:doi
222:of
3594::
3572::
3559::
3546::
3533::
3520::
3507::
3494::
3481::
3468::
3455::
3442::
3429::
3404::
3389::
2812::
2740::
2722::
430:^
403:.
397:.
306:.
298:,
286:,
278:,
274:,
270:,
266:,
262:,
258:,
234:.
3341:)
3317:)
3308:)
3299:)
3262:)
3225:)
3216:)
3207:)
3175:)
3166:)
3142:)
3105:)
3081:)
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3040:)
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3007:)
2998:)
2966:)
2957:)
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2939:)
2930:)
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2894:)
2827:)
2818:)
2800:)
2791:)
2782:)
2773:)
2764:)
2755:)
2746:)
2728:)
2689:)
2680:)
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2604:)
2580:)
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2553:)
2544:)
2535:)
2526:)
2517:)
2508:)
2499:)
2490:)
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2472:)
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2454:)
2445:)
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2427:)
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2330:)
2321:)
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2123:)
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2078:)
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2060:)
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2042:)
2010:)
2001:)
1992:)
1983:)
1974:)
1950:)
1926:)
1902:)
1893:)
1861:)
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1828:)
1819:)
1810:)
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1792:)
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1733:)
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1692:)
1683:)
1651:)
1642:)
1633:)
1609:)
1600:)
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1500:)
1491:)
1482:)
1473:)
1464:)
1455:)
1446:)
1437:)
1428:)
1419:)
1410:)
1401:)
1369:)
1312:)
1288:)
1264:)
1255:)
1223:)
1199:)
1175:)
1166:)
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1124:)
1100:)
1076:)
1067:)
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1026:)
1017:)
1008:)
999:)
990:)
981:)
972:)
940:)
909:)
885:)
861:)
852:)
794:)
785:)
761:)
752:)
743:)
734:)
725:)
716:)
707:)
698:)
689:)
680:)
671:)
662:)
653:)
644:)
543:e
536:t
529:v
424:.
411::
395:"
391:"
212:(
62:)
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