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Gryposaurus

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5295: 5290: 1145: 4380: 881: 4394: 1615: 5277: 160: 4387: 1598:, like other cranial modifications in duckbills, may have been used for a variety of social functions, such as identification of sexes or species and social ranking. It could also have functioned as a tool for broadside pushing or butting in social contests, and there may have been inflatable air sacs flanking it for both visual and auditory signaling. The top of the arch is roughened in some specimens, suggesting that it was covered by thick, 5301: 567: 1062: 1133: 708: 136: 5284: 3333: 1101:-shaped scutes upwards of 3.8 centimeters long (1.5 inches) on the flank and tail; uniform polygonal scales on the neck and sides of the body; and pyramidal structures, flattened side-to-side, with fluted sides, longer than tall and found along the top of the back in a single midline row. In 2016, Gregory S. Paul estimated the size of 539:, and for many years the two were thought to be synonyms. It is known from numerous skulls, some skeletons, and even some skin impressions that show it to have had pyramidal scales projecting along the midline of the back. It is most easily distinguished from other duckbills by its narrow arching nasal hump, sometimes described as similar to a " 1472: 823:
had enlarged prongs along its upper margin, where the lower jaw's beak was based. This new species greatly expands the geographic range of this genus, and there may be a second, more lightly built species present as well. Multiple gryposaur species are known from the Kaiparowits Formation,
1970:; Hernández,-Rivera, René; Gates, Terry; Paul, Gregory S.; Nesbitt, Sterling; Serrano-Brañas, Claudia Inés; Garcia-de la Garza, Juan Pablo (2006). "Large hadrosaurine dinosaurs from the latest Campanian of Coahuila, Mexico". In Lucas, Spencer G.; Sullivan Robert M. (eds.). 3221:
Eaton, J.G., 2002. Multituberculate mammals from the Wahweap (Campanian, Aquilan) and Kaiparowits (Campanian, Judithian) formations, within and near Grand Staircase–Escalante National Monument, southern Utah. Miscellaneous Publication 02-4, Utah Geological Survey, 66
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swamps, ponds and lakes, and was bordered by highlands. The climate was wet and humid, and supported an abundant and diverse range of organisms. This formation contains one of the best and most continuous records of Late Cretaceous terrestrial life in the world.
1124:. In profile view, they rise into a rounded hump in front of the eyes, reaching a height as tall as the highest point of the back of the skull. The skeleton is known in great detail, making it a useful point of reference for other duckbill skeletons. 2415: 3307:
Eaton, Jeffrey G.; Cifelli, Richard L.; Hutchinson, J. Howard; Kirkland, James I.; Parrish, J. Michael (1999). "Cretaceous vertebrate faunas from the Kaiparowits Plateau, south-central Utah". In Gillete David D. (ed.).
3209: 2122:(1979). "Upper Cretaceous dinosaurs from the Bearpaw Shale (marine) of south-central Montana with a checklist of Upper Cretaceous dinosaur remains from marine sediments in North America". 5842: 3084: 3210:
40Ar/39Ar age of the Kaiparowits Formation, southern Utah, and correlation of contemporaneous Campanian strata and vertebrate faunas along the margin of the Western Interior Basin
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Lull, Richard Swann; Wright, Nelda E. (1942). Hadrosaurian Dinosaurs of North America. Geological Society of America Special Paper 40. Geological Society of America. pp. 164–172.
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that contained hundreds of teeth, only a relative handful of which were in use at any time. Plant material would have been cropped by its broad beak, and held in the jaws by a
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Titus, Alan L. and Mark A. Loewen (editors). At the Top of the Grand Staircase: The Late Cretaceous of Southern Utah. 2013. Indiana University Press. Hardbound: 634 pp.
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appears to have only existed for part of the duration of time that the rocks were being formed. As the formation was being laid down, it recorded a change to more
2034:(1916). "Contributions to the geology and paleontology of San Juan County, New Mexico. 2. Vertebrate faunas of the Ojo Alamo, Kirtland and Fruitland Formations". 2400:
The hadrosaurian dinosaurs (ornithischia: hadrosauria) of Big Bend National Park, Brewster County, Texas, with implications for late Cretaceous paleozoogeography
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was once thought to fall into this group as well, before it was described (then known under the nickname "Antonio"). A subfamily, Gryposaurinae, was coined by
5872: 5773: 5294: 3031: 788:, from slightly older rocks in Montana than the classic gryposaur localities of Alberta. Based on two parts of a skeleton collected in 1916 for the 2795:(1902). "On Vertebrata of the mid-Cretaceous of the Northwest Territory. 2. New genera and species from the Belly River Series (mid-Cretaceous)". 672:
type specimen is missing the front part of the skull, so the full shape of the nasal arch cannot be seen. The 1942 publication of the influential
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have been unearthed for decades, but none were considered to be identifiable as a determined genus of hadrosaur, but do resemble some species of
4379: 5887: 5289: 3092: 5837: 3366: 3317: 2970: 2908: 2668: 2637: 2601: 2299:(Ornithischia: Hadrosauridae) with descriptions of two new hadrosaurid species and an evaluation of hadrosaurid phylogenetic relationships". 2215: 2187: 2156: 2100: 1881: 1838: 1637:
indicates that the Kaiparowits Formation was deposited between 76.1 and 74.0 million years ago, during the Campanian stage of the Late
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Zanno, Lindsay E.; Sampson, Scott D. (2005). "A new oviraptorosaur (Theropoda; Maniraptora) from the Late Cretaceous (Campanian) of Utah".
1522:-like organ. Its feeding range would have extended from the ground to about 4 m (13 ft) above. The paleontologists who unearthed 3114: 2584:; Weishampel, David B.; Forster, Catherine A (2004). "Hadrosauridae". In Weishampel, David B.; Dodson, Peter; OsmĂłlska Halszka (eds.). 789: 761:
of Montana (which have since fallen out of the literature), by 1990 he had changed his position, and was among the first to again use
3242: 5276: 543:," and which may have been used for species or sexual identification, and/or combat with individuals of the same species. A large 5862: 5852: 5655: 1967: 1907:, a new genus and species of trachodont dinosaur from the Belly River Formation of Alberta, with a description of the skull of 966:, Montana, USA, is known from partial skulls and skeletons from several individuals. Its nasal arch is prominent like that of 5867: 5678: 5665: 2932: 2053: 1802: 645: 147: 1641:
period. During the Late Cretaceous period, the site of the Kaiparowits Formation was located near the western shore of the
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to the east. The plateau where dinosaurs lived was an ancient floodplain dominated by large channels and abundant wetland
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as part of a larger revision that promoted Hadrosaurinae to family status, but is not now in use. A rough equivalent is
1169: 366: 2688: 1144: 618:. This skull was missing the snout, which had eroded into fragments; Brown restored it after the duckbill now known as 5300: 1120:
differ in details of the skull and lower jaw. The prominent nasal arch found in this genus is formed from the paired
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back in the 2000s brought the fact that this creature dined on tough, fibrous plant material which would imply that
624:, which was flat-headed, and believed that some unusual pieces were evidence of compression. Lambe's description of 589: 4386: 1105:
at 7.5 metres (25 ft) in length and 2.5 metric tons (2.8 short tons) in body mass and the other two species (
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based on the phylogenetic analysis conducted by Prieto-Márquez and Wagner in 2012, showing the relationships of
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The Illustrated Encyclopedia of Dinosaurs: An Original and Compelling Insight into Life in the Dinosaur Kingdom
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without hollow head crests. The general term "gryposaur" is sometimes used for duckbills with arched nasals.
2814: 2091:; Horner, Jack R. (1990). "Hadrosauridae". In Weishampel, David B.; Dodson, Peter; OsmĂłlska Halszka (eds.). 2031: 1642: 963: 918:
Dinosaur Park Formation of Alberta, Canada. It is now thought that another species from the same formation,
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Redescription of a remarkably large Gryposaurus notabilis (Dinosauria: Hadrosauridae) from Alberta, Canada
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was a hadrosaurid of typical size and shape; one of the best specimens of this genus, the nearly complete
959: 508: 5799: 5635: 4191: 3493: 1734: 1623: 872:, was unearthed in the Javelina Formation in Texas. Further research is needed to confirm its validity. 808: 516: 2995:, a new species of hadrosaurid dinosaur from the Late Cretaceous of the Pacific coast of North America" 733:
was most common in the late 1970s–early 1980s, and appears in some popular books; one well-known work,
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is incompletely known, lacking most of the bones in front of the eyes, it was very similar to that of
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This situation is made more confusing by old suggestions by some authors, including Jack Horner, that
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present instead. Other dinosaurs known from only the lower part of the formation include the horned
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Hopson, James A. (1975). "The evolution of cranial display structures in hadrosaurian dinosaurs".
1089:) came from an animal about 8.2 meters (27 feet) long. This specimen also has the best example of 930:. The two had been differentiated by the size of the nasal arch (larger and closer to the eyes in 824:
represented by cranial and postcranial remains, and were larger than their northern counterparts.
5407: 5018: 3603: 3289: 3191: 3057: 3012: 2875: 2853: 2767: 2740:"Hadrosaurian dinosaurs from the Maastrichtian Javelina Formation, Big Bend National Park, Texas" 2552: 2469: 2451: 2416:"Hadrosaurian dinosaurs from the Maastrichtian Javelina Formation, Big Bend National Park, Texas" 2379: 2261: 1634: 1003: 828: 524: 408: 381: 154: 2629: 2621: 1933:(1910). "The Cretaceous Ojo Alamo beds of New Mexico with description of the new dinosaur genus 5804: 2593: 5791: 5414: 5153: 5074: 5034: 3918: 3804: 3750: 3721: 3313: 2966: 2914: 2904: 2664: 2633: 2597: 2500: 2443: 2327:(Dinosauria: Hadrosauridae) from the late Campanian Kaiparowits Formation, southern Utah, USA" 2232: 2211: 2183: 2179: 2173: 2152: 2148: 2142: 2096: 1978:. Albuquerque, New Mexico: New Mexico Museum of Natural History and Science. pp. 299–315. 1877: 1834: 5751: 3243:"Presidential Proclamation: Establishment of the Grand Staircase–Escalante National Monument" 2660: 2654: 1097:, showing this dinosaur to have had several different types of scalation: pyramidal, ridged, 608:
shortly thereafter, Lambe drawing attention to its unusual nasal crest. A few years earlier,
5555: 5548: 5358: 5351: 4880: 4791: 4629: 4530: 4516: 4230: 4198: 3833: 3742: 3656: 3591: 3338: 3312:. Miscellaneous Publication 99-1. Salt Lake City: Utah Geological Survey. pp. 345–353. 3281: 3183: 3002: 2759: 2585: 2544: 2435: 2371: 2338: 2013: 1946: 1865: 1782: 1745: 1515: 1158:
was a saurolophine (hadrosaurine of older references) hadrosaurid, a member of the duckbill
1068: 915: 5640: 5435: 5386: 5249: 5180: 5083: 4903: 4622: 4608: 4587: 4205: 4078: 3845: 3613: 3451: 2237:, from the Campanian of the East Coast of North America, with a reevaluation of the genus" 2203: 1719: 1669: 566: 477: 48: 2965:. Vol. Supplement 2. Jefferson, North Carolina: McFarland & Co. pp. 79–80. 1866: 3179: 2755: 2540: 2431: 2009: 891:
As of 2016, there are currently three named species that are recognized as valid today:
5650: 5645: 5583: 5520: 5393: 5242: 5205: 5187: 5132: 4764: 4580: 4573: 4502: 4424: 4411: 4339: 4327: 4296: 4271: 4135: 4057: 4018: 4004: 3881: 3792: 3714: 3692: 3685: 3426: 2958: 2792: 1900: 1861: 1754: 1695: 1680: 1562: 1028: 605: 601: 330: 236: 2056:(1919). "Preliminary description of a new species of trachodont dinosaur of the genus 1113:) at 8 metres (26 ft) in length and 3 metric tons (3.3 short tons) in body mass. 1061: 5821: 5590: 5534: 5513: 5484: 5400: 5337: 5235: 5228: 5214: 5124: 5104: 5097: 4994: 4944: 4937: 4921: 4896: 4869: 4862: 4855: 4826: 4777: 4739: 4704: 4685: 4657: 4601: 4264: 4257: 4212: 4142: 4011: 3870: 3855: 3817: 3728: 3621: 3584: 3577: 3293: 3250: 2771: 2739: 2586: 2455: 2383: 2362:(Dinosauria: Hadrosauridae) from the Upper Campanian Kaiparowits Formation of Utah". 2343: 2322: 1740: 1706: 1700: 1685: 1574: 1389: 1078: 1007: 758: 492: 444: 385: 278: 265: 86: 3195: 3016: 2556: 1829:
Creisler, Benjamin S. (2007). "Deciphering duckbills". In Carpenter, Kenneth (ed.).
17: 5498: 5474: 5462: 5428: 5421: 5372: 5365: 5171: 5115: 5090: 5062: 4951: 4753: 4718: 4664: 4594: 4566: 4452: 4445: 4285: 4250: 4177: 4170: 4092: 4033: 3988: 3890: 3764: 3757: 3706: 3670: 3628: 3478: 3417: 2375: 1989: 1930: 1728: 1724: 1711: 1690: 1676: 1568: 1545: 1365: 1227: 1213: 1173: 1164: 1015: 971: 904: 844: 801: 778: 609: 488: 313: 291: 220: 3139: 3115:"New Dinosaur Species, Gryposaurus Monumentensis, Unearthed In Utah | Science 2.0" 5745: 5687: 5576: 5541: 5379: 5221: 5194: 5146: 4972: 4958: 4930: 4910: 4889: 4833: 4812: 4805: 4798: 4732: 4725: 4678: 4643: 4615: 4551: 4544: 4313: 4163: 4149: 4121: 4099: 4041: 3904: 3735: 3699: 3663: 3559: 3546: 3505: 3375: 2713: 1790: 1715: 1507: 1484: 1406: 1251: 1178: 1132: 1020: 979: 849: 717: 707: 692:. Reviews beginning in the 1990s, however, called into question the identity of 614: 548: 535: 423: 252: 61: 135: 5629: 5604: 5597: 5569: 5562: 5453: 5442: 5328: 5027: 5007: 4965: 4840: 4692: 4559: 4509: 4474: 4467: 4438: 4278: 4156: 4128: 3997: 3970: 3945: 3785: 3778: 3771: 3649: 3328: 3187: 2548: 2504: 1774: 1762: 1650: 1638: 1492: 1121: 1090: 975: 943: 820: 757:
for a partial skull and skeleton and a second less-complete skeleton from the
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provided evidence of a different type of skull configuration, and by 1916 the
544: 106: 71: 31: 5736: 2918: 2447: 1506:. Its skull had special joints that permitted a grinding motion analogous to 527:
in Texas may extend the temporal range of the genus to 66 million years ago.
5611: 5527: 5506: 5344: 5041: 4979: 4488: 4460: 4320: 4184: 4114: 4085: 3911: 3897: 3640: 2692: 2095:(1st ed.). Berkeley: University of California Press. pp. 534–561. 1833:. Bloomington and Indianapolis: Indiana University Press. pp. 185–210. 1786: 1749: 1646: 1603: 1499: 1496: 1471: 1198: 1159: 955: 911: 866: 677: 648:'s naming of a nearly complete skeleton from the Dinosaur Park Formation as 612:
had collected and described a partial skull from New Mexico, which he named
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material. Horner, who described the specimens, considered it to be a less
5730: 5491: 5160: 5048: 4848: 4819: 4746: 4671: 4636: 4537: 4523: 4481: 4305: 4106: 4071: 3977: 3963: 3925: 3435: 3408: 3399: 2827:
Canada Department of Mines Geological Survey Bulletin (Geological Series)
1672: 797: 585: 540: 473: 204: 191: 101: 96: 81: 76: 66: 2763: 2439: 2017: 1994:"Cretaceous Eocene correlation in New Mexico, Wyoming, Montana, Alberta" 696:, which has limited material for comparison with other duckbills. Thus, 5778: 4784: 4219: 2231:
Prieto-Márquez, Alberto; Weishampel, David B.; Horner, John R. (2006).
1599: 774: 512: 504: 455: 116: 91: 2738:
Lehman, Thomas M.; Wick, Steve L.; Wagner, Jonathan R. (1 July 2016).
2527:(Dinosauria, Hadrosauridae), with a taxonomic revision of the genus". 2494: 2414:
Lehman, Thomas M.; Wick, Steven L.; Wagner, Jonathan R. (March 2016).
946:, as well as with two skeletons with skulls that had been assigned to 769:, although known from fragmentary material, can be distinguished from 4650: 4243: 3393: 2898: 1950: 1778: 1770: 1766: 1098: 869: 862: 753:
in a photo caption. Although Horner in 1979 used the new combination
451: 181: 5707: 1993: 819:. Its skull was more robust than that of the other species, and its 2592:(2nd ed.). Berkeley: University of California Press. pp.  1645:, a large inland sea that split North America into two landmasses, 990:
was listed second on the top 10 list of new species in 2008 by the
938:). Ten complete skulls and twelve fragmentary skulls are known for 815:, includes a skull and partial skeleton that represent the species 4495: 4025: 2626:
Dinosaur Provincial Park: A Spectacular Ancient Ecosystem Revealed
1613: 1583: 1553: 1519: 1511: 1503: 1470: 1143: 1131: 1060: 879: 706: 565: 555: 466: 459: 2523:
Prieto–Marquez, Alberto (2010). "The braincase and skull roof of
1753:. Other paleofauna present in the Kaiparowits Formation included 1045:. However, this synonymy was rejected in the 2004 edition of the 5765: 3863: 1758: 1654: 784:
Further research has revealed the presence of a second species,
520: 5711: 5316: 4409: 3544: 3386: 3348: 1186:, but it is slightly younger. Additionally, while the skull of 3344: 1974:. New Mexico Museum of Natural History and Science Bulletin, 507:, Canada, and two formations in the United States: the Lower 3286:
10.1671/0272-4634(2005)025[0897:ANOTMF]2.0.CO;2
982:"Hadrosauravus" is an early, unused name for this species. 970:, but farther forward on the snout, and its teeth are less 934:) and the form of the upper arm (longer and more robust in 632:
skull had been redone with a nasal arch and both Brown and
1872:. Jefferson, North Carolina: McFarland & Co. pp.  700:
has once again been separated, at least temporarily, from
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2278, a skull and partial skeleton collected in 1913 by
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Gates, Terry; Sampson, Scott (2006). "A new species of
986:
is known from a skull and partial skeleton from Utah.
853:. However, in 2016, a possibly forth valid species of 554:
around 8 meters (26 feet) long, it may have preferred
3140:"Toothy dinosaur newest to come out of southern Utah" 1972:
Late Cretaceous vertebrates from the Western Interior
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Horns and Beaks: Ceratopsian and Ornithopod Dinosaurs
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is absent from the upper part of the formation, with
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that lived about 80 to 75 million years ago, in the
392: 373: 358: 346: 320: 5720: 5621: 5472: 5452: 5327: 5204: 5170: 5114: 5073: 5017: 4993: 4920: 4879: 4763: 4702: 4422: 4338: 4295: 4229: 4055: 3987: 3943: 3880: 3844: 3803: 3639: 3602: 3557: 3504: 3477: 3450: 3032:"Plant-eating dinosaur a "Cretaceous weed whacker"" 2714:"Scientists Announce Top 10 New Species; Issue SOS" 2935:(1920). "The osteology of the trachodont dinosaur 2886:. Geological Society of America. pp. 110–117. 2628:. Bloomington: Indiana University Press. pp.  2036:United States Geological Survey Professional Paper 1939:Bulletin of the American Museum of Natural History 644:were one and the same. This idea was reflected in 2903:. Princeton University Press. pp. 337–339. 2718:International Institute for Species Exploration 2624:. In Currie, Phillip J.; Koppelhus Eva (eds.). 992:International Institute for Species Exploration 2882:. Geological Society of America Special Paper 2860:. Geological Society of America Special Paper 2268:. Geological Society of America Special Paper 2210:. New York: Crescent Books. pp. 116–121. 3360: 2941:University of Toronto Studies, Geology Series 2864:. Geological Society of America. p. 226. 2821:, with a description of the type specimen of 835:, indeterminate hadrosaur remains resembling 660:in its own genus). Direct comparison between 8: 2272:. Geological Society of America. p. 21. 688:question for nearly fifty years in favor of 5843:Grand Staircase–Escalante National Monument 2321:Gates, Terry A.; Sampson, Scott D. (2007). 2066:Transactions of the Royal Society of Canada 1622:(left background) and other animals of the 1514:were continually replacing and packed into 813:Grand Staircase–Escalante National Monument 604:. This specimen was described and named by 5833:Late Cretaceous dinosaurs of North America 5708: 5324: 5313: 4999: 4769: 4710: 4430: 4419: 4406: 4235: 4063: 3955: 3951: 3809: 3569: 3565: 3554: 3541: 3483: 3456: 3447: 3383: 3367: 3353: 3345: 2653:Lambert, David; the Diagram Group (1990). 2620:Ryan, Michael J.; Evans, David C. (2005). 2172:Lambert, David; the Diagram Group (1983). 1962: 1960: 745:), but identifies the mounted skeleton of 134: 38: 3006: 2989:Prieto-Márquez, A.; Wagner, J.R. (2011). 2984: 2982: 2470:"Current Research | Big Bend Conservancy" 2342: 2331:Zoological Journal of the Linnean Society 994:. Recently, a possible fourth species of 735:The Illustrated Encyclopedia of Dinosaurs 523:. A possible additional species from the 3161: 3159: 2576: 2574: 2572: 2570: 2568: 2566: 2518: 2516: 2514: 2287: 2285: 2283: 2281: 2279: 454:"; sometimes incorrectly translated as " 5893:Multispecific non-avian dinosaur genera 2880:Hadrosaurian Dinosaurs of North America 2858:Hadrosaurian Dinosaurs of North America 2615: 2613: 2403:(Thesis thesis). Texas Tech University. 2316: 2314: 2266:Hadrosaurian Dinosaurs of North America 2147:. Secaucus, NJ: Citadel Press. p.  2114: 2112: 1856: 1854: 1852: 1850: 1818: 3058:"Huge New Dinosaur Had A Serious Bite" 2900:The Princeton Field Guide to Dinosaurs 2797:Contributions to Canadian Paleontology 2301:Museum of the Rockies Occasional Paper 2083: 2081: 2079: 1998:Geological Society of America Bulletin 1895: 1893: 1824: 1822: 3208:Roberts EM, Deino AL, Chan MA (2005) 1182:has been proposed to be a synonym of 1176:, as used by Alberto Prieto-Márquez. 1033:was considered a possible species of 7: 1136:Size comparison of three species of 1381: 1357: 1298: 1274: 1267: 1243: 1219: 1208: 1010:, along with an unnamed species of 3274:Journal of Vertebrate Paleontology 3212:. Cretaceous Research 26: 307–318. 2529:Journal of Vertebrate Paleontology 2364:Journal of Vertebrate Paleontology 790:American Museum of Natural History 765:in print. Current thought is that 25: 5873:Campanian genus first appearances 865:, who was the last leader of the 668:is hindered by the fact that the 5299: 5293: 5288: 5282: 5275: 4392: 4385: 4378: 3331: 2659:. New York: Avon Books. p.  2397:Wagner, Jonathan R. (May 2001). 2344:10.1111/j.1096-3642.2007.00349.x 2178:. New York: Avon Books. p.  1572:and the hollow-crested duckbill 1548:of the Dinosaur Park Formation, 158: 5656:List of informal dinosaur names 3310:Vertebrate Paleontology in Utah 3030:Dunham, Will (3 October 2007). 2689:"Re: What are these dinosaurs?" 2295:(1992). "Cranial morphology of 1024:, but with a more solid crest. 5666:Timeline of hadrosaur research 3085:"New duck-dinosaur discovered" 2376:10.1080/02724634.2006.10010069 1803:Timeline of hadrosaur research 1720:Parasaurolophus cyrtocristatus 1668:with other dinosaurs, such as 1635:Argon-argon radiometric dating 1594:The distinctive nasal arch of 1205:among the other kritosaurins: 1085:(now regarded as a synonym of 827:In Texas, specifically at the 596:from what is now known as the 148:Natural History Museum of Utah 1: 5888:Fossil taxa described in 1914 2999:Acta Palaeontologica Polonica 2244:Acta Palaeontologica Polonica 5838:Taxa named by Lawrence Lambe 3083:Than, Ker (3 October 2007). 1053:being tabulated as dubious. 1037:, following the synonymy of 960:Lower Two Medicine Formation 847:or at least some species of 5878:Campanian genus extinctions 2963:Dinosaurs: The Encyclopedia 2878:; Wright, Nelda E. (1942). 2856:; Wright, Nelda E. (1942). 2264:; Wright, Nelda E. (1942). 2144:The New Dinosaur Dictionary 1868:Dinosaurs: The Encyclopedia 1116:The three named species of 978:-like characteristics. The 741:for the Canadian material ( 721:is also the same as either 711:Skull, Royal Tyrrell Museum 5909: 3091:. NBC News. Archived from 2474:fossildiscoveryexhibit.com 2206:(1985). "Hadrosaurids I". 2175:A Field Guide to Dinosaurs 1602:skin, or that there was a 1006:, which dates to the late 400:Lehman, Wick, Wagner, 2016 29: 5323: 5312: 5273: 5002: 4772: 4713: 4433: 4418: 4405: 4376: 4238: 4066: 3958: 3954: 3812: 3572: 3568: 3553: 3540: 3486: 3459: 3446: 3382: 3188:10.1017/S0094837300002165 2897:Paul, Gregory S. (2016). 2622:"Ornithischian Dinosaurs" 2549:10.1080/02724631003762971 1741:Kosmoceratops richardsoni 1662:Gryposaurus monumentensis 1403: 1386: 1379: 1362: 1355: 1324:Gryposaurus monumentensis 1320: 1303: 1296: 1279: 1272: 1265: 1248: 1241: 1224: 1217: 1030:Stephanosaurus marginatus 680:on hadrosaurs sealed the 414: 407: 343: 338: 319: 312: 155:Scientific classification 153: 142: 133: 41: 27:Extinct genus of reptiles 2937:Kritosaurus incurvimanus 2141:Glut, Donald F. (1982). 2062:Kritosaurus incurvimanus 1556:-influenced conditions. 1083:Kritosaurus incurvimanus 1027:The dubious hadrosaurid 1018:which closely resembles 924:Gryposaurus incurvimanus 920:Kritosaurus incurvimanus 861:, which was named after 662:Kritosaurus incurvimanus 654:Gryposaurus incurvimanus 650:Kritosaurus incurvimanus 443:(meaning "hooked-nosed ( 30:Not to be confused with 5863:Paleontology in Alberta 5853:Paleontology in Montana 2744:Journal of Paleontology 2420:Journal of Paleontology 2124:Journal of Paleontology 1864:(1997). "Gryposaurus". 1781:were present including 1643:Western Interior Seaway 1002:, was unearthed in the 621:Edmontosaurus annectens 598:Dinosaur Park Formation 501:Dinosaur Park Formation 2817:(1924). "On the genus 2720:. Newswise. 2008-05-23 2656:The Dinosaur Data Book 1626: 1502:, eating a variety of 1480: 1152: 1141: 1071: 888: 807:New material from the 773:by differences in the 755:Hadrosaurus notabilis 712: 694:Kritosaurus navajovius 600:of Alberta, along the 578: 509:Two Medicine Formation 5868:Kaiparowits Formation 5800:Paleobiology Database 5636:Basturs Poble bonebed 3008:10.4202/app.2011.0049 2525:Gryposaurus notabilis 1913:The Ottawa Naturalist 1905:Gryposaurus notabilis 1777:. A variety of early 1735:Nasutoceratops titusi 1712:armored ankylosaurids 1624:Kaiparowits Formation 1617: 1546:bird-hipped dinosaurs 1474: 1307:Gryposaurus notabilis 1147: 1135: 1087:Gryposaurus notabilis 1064: 883: 809:Kaiparowits Formation 710: 666:Gryposaurus notabilis 658:Gryposaurus notabilis 569: 517:Kaiparowits Formation 325:Gryposaurus notabilis 5883:Ornithischian genera 5858:Paleontology in Utah 5658:(contains hadrosaur 5056:Probrachylophosaurus 2089:Weishampel, David B. 1808:2016 in paleontology 1716:duckbilled hadrosaur 1283:Gryposaurus latidens 833:El Picacho Formation 562:History of discovery 18:Gryposaurus latidens 5848:Dinosaur Park fauna 3180:1975Pbio....1...21H 3095:on October 15, 2013 2993:Saurolophus morrisi 2876:Lull, Richard Swann 2854:Lull, Richard Swann 2823:Lambeosaurus lambei 2815:Gilmore, Charles W. 2764:10.1017/jpa.2016.48 2756:2016JPal...90..333L 2541:2010JVPal..30..838P 2493:Bertozzo, Filippo. 2440:10.1017/jpa.2016.48 2432:2016JPal...90..333L 2262:Lull, Richard Swann 2235:Hadrosaurus foulkii 2032:Gilmore, Charles W. 2018:10.1130/gsab-25-355 2010:1914GSAB...25..355B 1757:(sharks and rays), 1750:Hagryphus giganteus 1729:Utahceratops gettyi 1586:-related settings. 1582:may have preferred 1197:The following is a 1014:and an undescribed 1000:Gryposaurus alsatei 926:), is a synonym of 796:is also known from 594:George F. Sternberg 499:are known from the 495:. Named species of 5408:Ornithopodichnites 5019:Brachylophosaurini 3604:Hypsilophodontidae 3247:September 18, 1996 3241:Clinton, William. 2793:Lambe, Lawrence M. 2323:"A new species of 1968:Kirkland, James I. 1909:Chasmosaurus belli 1901:Lambe, Lawrence M. 1691:Ornithomimus velox 1627: 1491:would have been a 1481: 1153: 1142: 1072: 1004:Javelina Formation 889: 829:Javelina Formation 713: 656:(although he left 636:had proposed that 579: 525:Javelina Formation 5815: 5814: 5714:Taxon identifiers 5705: 5704: 5701: 5700: 5697: 5696: 5415:Shiraminesauropus 5308: 5307: 5271: 5270: 5267: 5266: 5263: 5262: 5259: 5258: 5154:Naashoibitosaurus 5075:Austrokritosauria 5035:Brachylophosaurus 4989: 4988: 4401: 4400: 4374: 4373: 4370: 4369: 4366: 4365: 4362: 4361: 4358: 4357: 4051: 4050: 3939: 3938: 3935: 3934: 3919:Transylvanosaurus 3805:Rhabdodontomorpha 3751:Notohypsilophodon 3722:Macrogryphosaurus 3536: 3535: 3532: 3531: 3528: 3527: 3524: 3523: 3319:978-1-55791-634-1 3253:on 28 August 2013 3119:www.science20.com 2972:978-0-89950-917-4 2933:Parks, William A. 2910:978-1-78684-190-2 2685:Olshevsky, George 2670:978-0-380-75896-8 2639:978-0-253-34595-0 2603:978-0-520-24209-8 2370:(3, Suppl): 65A. 2217:978-0-517-46890-6 2204:Norman, David. B. 2189:978-0-380-83519-5 2158:978-0-8065-0782-8 2102:978-0-520-06727-1 2054:Parks, William A. 1883:978-0-89950-917-4 1840:978-0-253-34817-3 1783:multituberculates 1746:oviraptorosaurian 1475:Restored head of 1463: 1462: 1454: 1453: 1445: 1444: 1436: 1435: 1427: 1426: 1418: 1417: 1344: 1343: 1335: 1334: 958:-early Campanian 577:), collected 1918 465:) lizard") was a 436: 435: 430: 401: 389: 370: 355: 16:(Redirected from 5900: 5808: 5807: 5795: 5794: 5782: 5781: 5769: 5768: 5756: 5755: 5754: 5741: 5740: 5739: 5709: 5682:mummy S.M. R4036 5556:Microhadrosaurus 5549:Mandschurosaurus 5359:Camptosaurichnus 5352:Bonaparteichnium 5325: 5318:Related articles 5314: 5303: 5297: 5292: 5287: 5286: 5279: 5000: 4881:Parasaurolophini 4792:Angulomastacator 4770: 4711: 4630:Shuangmiaosaurus 4531:Huehuecanauhtlus 4517:Gongpoquansaurus 4431: 4420: 4407: 4396: 4390: 4389: 4382: 4236: 4231:Hadrosauriformes 4199:Ratchasimasaurus 4064: 3956: 3952: 3834:Muttaburrasaurus 3810: 3743:Muttaburrasaurus 3657:Atlascopcosaurus 3592:Thescelosauridae 3570: 3566: 3555: 3542: 3484: 3457: 3448: 3441: 3440: 3384: 3369: 3362: 3355: 3346: 3341: 3339:Dinosaurs portal 3336: 3335: 3334: 3324: 3323: 3304: 3298: 3297: 3269: 3263: 3262: 3260: 3258: 3249:. Archived from 3238: 3232: 3229: 3223: 3219: 3213: 3206: 3200: 3199: 3163: 3154: 3153: 3151: 3150: 3136: 3130: 3129: 3127: 3126: 3111: 3105: 3104: 3102: 3100: 3080: 3074: 3073: 3071: 3069: 3062:ScienceDaily.com 3054: 3048: 3047: 3045: 3043: 3027: 3021: 3020: 3010: 2986: 2977: 2976: 2955: 2949: 2948: 2929: 2923: 2922: 2894: 2888: 2887: 2872: 2866: 2865: 2850: 2844: 2841: 2835: 2834: 2811: 2805: 2804: 2789: 2783: 2782: 2780: 2778: 2735: 2729: 2728: 2726: 2725: 2710: 2704: 2703: 2701: 2700: 2691:. Archived from 2681: 2675: 2674: 2650: 2644: 2643: 2617: 2608: 2607: 2591: 2578: 2561: 2560: 2520: 2509: 2508: 2490: 2484: 2483: 2481: 2480: 2466: 2460: 2459: 2411: 2405: 2404: 2394: 2388: 2387: 2355: 2349: 2348: 2346: 2318: 2309: 2308: 2289: 2274: 2273: 2258: 2252: 2251: 2241: 2228: 2222: 2221: 2200: 2194: 2193: 2169: 2163: 2162: 2138: 2132: 2131: 2116: 2107: 2106: 2085: 2074: 2073: 2050: 2044: 2043: 2028: 2022: 2021: 1986: 1980: 1979: 1964: 1955: 1954: 1927: 1921: 1920: 1897: 1888: 1887: 1871: 1858: 1845: 1844: 1826: 1666:paleoenvironment 1649:to the west and 1620:G. monumentensis 1516:dental batteries 1382: 1358: 1299: 1275: 1268: 1244: 1220: 1209: 1111:G. monumentensis 1093:impressions for 1069:life restoration 988:G. monumentensis 984:G. monumentensis 916:Upper Cretaceous 901:G. monumentensis 885:G. monumentensis 817:G. monumentensis 729:, or both. This 420: 399: 394: 380: 377:G. monumentensis 375: 365: 360: 353: 348: 322: 303: 290: 277: 264: 251: 235: 219: 163: 162: 146:skeleton in the 144:G. monumentensis 138: 128: 58: 47:Temporal range: 39: 21: 5908: 5907: 5903: 5902: 5901: 5899: 5898: 5897: 5818: 5817: 5816: 5811: 5803: 5798: 5790: 5785: 5777: 5772: 5764: 5759: 5750: 5749: 5744: 5735: 5734: 5729: 5716: 5706: 5693: 5674:mummy AMNH 5060 5641:Dakota (fossil) 5617: 5468: 5448: 5436:Tetrapodosaurus 5387:Hadrosaurichnus 5319: 5304: 5281: 5255: 5250:Shantungosaurus 5200: 5181:Augustynolophus 5166: 5110: 5084:Bonapartesaurus 5069: 5013: 4985: 4916: 4904:Parasaurolophus 4875: 4759: 4707: 4698: 4623:Qianjiangsaurus 4609:Probactrosaurus 4588:Penelopognathus 4427: 4414: 4397: 4384: 4354: 4334: 4291: 4225: 4206:Riabininohadros 4079:Bayannurosaurus 4060: 4047: 3983: 3948: 3931: 3876: 3846:Tenontosauridae 3840: 3799: 3635: 3614:Gideonmantellia 3598: 3562: 3549: 3520: 3500: 3473: 3467:Avemetatarsalia 3452:Avemetatarsalia 3442: 3389: 3388: 3378: 3373: 3337: 3332: 3330: 3327: 3320: 3306: 3305: 3301: 3271: 3270: 3266: 3256: 3254: 3240: 3239: 3235: 3230: 3226: 3220: 3216: 3207: 3203: 3165: 3164: 3157: 3148: 3146: 3138: 3137: 3133: 3124: 3122: 3113: 3112: 3108: 3098: 3096: 3082: 3081: 3077: 3067: 3065: 3064:. Science Daily 3056: 3055: 3051: 3041: 3039: 3029: 3028: 3024: 2988: 2987: 2980: 2973: 2959:Glut, Donald F. 2957: 2956: 2952: 2931: 2930: 2926: 2911: 2896: 2895: 2891: 2874: 2873: 2869: 2852: 2851: 2847: 2842: 2838: 2813: 2812: 2808: 2791: 2790: 2786: 2776: 2774: 2737: 2736: 2732: 2723: 2721: 2712: 2711: 2707: 2698: 2696: 2683: 2682: 2678: 2671: 2652: 2651: 2647: 2640: 2619: 2618: 2611: 2604: 2582:Horner, John R. 2580: 2579: 2564: 2522: 2521: 2512: 2492: 2491: 2487: 2478: 2476: 2468: 2467: 2463: 2413: 2412: 2408: 2396: 2395: 2391: 2357: 2356: 2352: 2320: 2319: 2312: 2293:Horner, John R. 2291: 2290: 2277: 2260: 2259: 2255: 2239: 2230: 2229: 2225: 2218: 2202: 2201: 2197: 2190: 2171: 2170: 2166: 2159: 2140: 2139: 2135: 2120:Horner, John R. 2118: 2117: 2110: 2103: 2087: 2086: 2077: 2052: 2051: 2047: 2030: 2029: 2025: 1988: 1987: 1983: 1966: 1965: 1958: 1945:(24): 267–274. 1929: 1928: 1924: 1899: 1898: 1891: 1884: 1862:Glut, Donald F. 1860: 1859: 1848: 1841: 1828: 1827: 1820: 1816: 1799: 1755:chondrichthyans 1632: 1618:Restoration of 1612: 1592: 1469: 1464: 1455: 1446: 1437: 1428: 1419: 1345: 1336: 1130: 1059: 948:K. incurvimanus 936:K. incurvimanus 922:(also known as 878: 747:K. incurvimanus 634:Charles Gilmore 575:G. incurvimanus 564: 478:Late Cretaceous 428: 398: 379: 364: 352: 334: 328: 301: 288: 275: 262: 249: 233: 217: 157: 129: 127: 126: 125: 124: 119: 114: 109: 104: 99: 94: 89: 84: 79: 74: 69: 64: 53: 52: 49:Late Cretaceous 45: 35: 28: 23: 22: 15: 12: 11: 5: 5906: 5904: 5896: 5895: 5890: 5885: 5880: 5875: 5870: 5865: 5860: 5855: 5850: 5845: 5840: 5835: 5830: 5820: 5819: 5813: 5812: 5810: 5809: 5796: 5783: 5770: 5757: 5742: 5726: 5724: 5718: 5717: 5712: 5703: 5702: 5699: 5698: 5695: 5694: 5692: 5691: 5684: 5676: 5668: 5663: 5653: 5651:Hypsilophodont 5648: 5646:Hadrosaur diet 5643: 5638: 5633: 5625: 5623: 5622:Other articles 5619: 5618: 5616: 5615: 5608: 5601: 5594: 5587: 5584:Stephanosaurus 5580: 5573: 5566: 5559: 5552: 5545: 5538: 5531: 5524: 5521:Fulgurotherium 5517: 5510: 5503: 5495: 5488: 5480: 5478: 5470: 5469: 5467: 5466: 5458: 5456: 5450: 5449: 5447: 5446: 5439: 5432: 5425: 5418: 5411: 5404: 5397: 5394:Iguanodontipus 5390: 5383: 5376: 5369: 5362: 5355: 5348: 5341: 5333: 5331: 5321: 5320: 5317: 5310: 5309: 5306: 5305: 5274: 5272: 5269: 5268: 5265: 5264: 5261: 5260: 5257: 5256: 5254: 5253: 5246: 5243:Laiyangosaurus 5239: 5232: 5225: 5218: 5210: 5208: 5206:Edmontosaurini 5202: 5201: 5199: 5198: 5191: 5188:Prosaurolophus 5184: 5176: 5174: 5168: 5167: 5165: 5164: 5157: 5150: 5143: 5136: 5133:Coahuilasaurus 5129: 5120: 5118: 5112: 5111: 5109: 5108: 5101: 5094: 5087: 5079: 5077: 5071: 5070: 5068: 5067: 5059: 5052: 5045: 5038: 5031: 5023: 5021: 5015: 5014: 5012: 5011: 5003: 4997: 4991: 4990: 4987: 4986: 4984: 4983: 4976: 4969: 4962: 4955: 4948: 4941: 4934: 4926: 4924: 4918: 4917: 4915: 4914: 4907: 4900: 4893: 4885: 4883: 4877: 4876: 4874: 4873: 4866: 4859: 4852: 4845: 4837: 4830: 4823: 4816: 4809: 4802: 4795: 4788: 4781: 4773: 4767: 4765:Lambeosaurinae 4761: 4760: 4758: 4757: 4750: 4743: 4736: 4729: 4722: 4714: 4708: 4703: 4700: 4699: 4697: 4696: 4689: 4682: 4675: 4668: 4661: 4654: 4647: 4640: 4633: 4626: 4619: 4612: 4605: 4598: 4591: 4584: 4581:Nanyangosaurus 4577: 4574:Nanningosaurus 4570: 4563: 4556: 4548: 4541: 4534: 4527: 4520: 4513: 4506: 4503:Gilmoreosaurus 4499: 4492: 4485: 4478: 4471: 4464: 4457: 4449: 4442: 4434: 4428: 4425:Hadrosauroidea 4423: 4416: 4415: 4412:Hadrosauroidea 4410: 4403: 4402: 4399: 4398: 4377: 4375: 4372: 4371: 4368: 4367: 4364: 4363: 4360: 4359: 4356: 4355: 4353: 4352: 4351: 4350: 4344: 4342: 4340:Hadrosauroidea 4336: 4335: 4333: 4332: 4328:Mantellisaurus 4324: 4317: 4310: 4301: 4299: 4297:Iguanodontidae 4293: 4292: 4290: 4289: 4282: 4275: 4272:Mantellisaurus 4268: 4261: 4254: 4247: 4239: 4233: 4227: 4226: 4224: 4223: 4216: 4209: 4202: 4195: 4188: 4181: 4174: 4167: 4160: 4153: 4146: 4139: 4136:Iguanacolossus 4132: 4125: 4118: 4111: 4103: 4096: 4089: 4082: 4075: 4067: 4061: 4058:Ankylopollexia 4056: 4053: 4052: 4049: 4048: 4046: 4045: 4038: 4030: 4022: 4019:Eousdryosaurus 4015: 4008: 4005:Dysalotosaurus 4001: 3993: 3991: 3985: 3984: 3982: 3981: 3974: 3967: 3959: 3949: 3944: 3941: 3940: 3937: 3936: 3933: 3932: 3930: 3929: 3922: 3915: 3908: 3901: 3894: 3886: 3884: 3882:Rhabdodontidae 3878: 3877: 3875: 3874: 3867: 3860: 3851: 3849: 3842: 3841: 3839: 3838: 3830: 3822: 3813: 3807: 3801: 3800: 3798: 3797: 3793:Weewarrasaurus 3789: 3782: 3775: 3768: 3761: 3754: 3747: 3739: 3732: 3725: 3718: 3715:Leaellynasaura 3711: 3703: 3696: 3693:Gasparinisaura 3689: 3686:Galleonosaurus 3682: 3674: 3667: 3660: 3653: 3645: 3643: 3637: 3636: 3634: 3633: 3625: 3618: 3609: 3607: 3600: 3599: 3597: 3596: 3588: 3581: 3573: 3563: 3558: 3551: 3550: 3545: 3538: 3537: 3534: 3533: 3530: 3529: 3526: 3525: 3522: 3521: 3519: 3518: 3517: 3516: 3510: 3508: 3502: 3501: 3499: 3498: 3497: 3496: 3487: 3481: 3475: 3474: 3472: 3471: 3470: 3469: 3460: 3454: 3444: 3443: 3439: 3438: 3429: 3427:Neornithischia 3420: 3411: 3402: 3396: 3387: 3380: 3379: 3374: 3372: 3371: 3364: 3357: 3349: 3343: 3342: 3326: 3325: 3318: 3299: 3280:(4): 897–904. 3264: 3233: 3224: 3214: 3201: 3155: 3131: 3106: 3075: 3049: 3022: 2978: 2971: 2950: 2924: 2909: 2889: 2867: 2845: 2836: 2819:Stephanosaurus 2806: 2784: 2750:(2): 333–356. 2730: 2705: 2687:(1999-11-16). 2676: 2669: 2645: 2638: 2609: 2602: 2588:The Dinosauria 2562: 2535:(3): 838–854. 2510: 2485: 2461: 2426:(2): 333–356. 2406: 2389: 2350: 2337:(2): 351–376. 2310: 2297:Prosaurolophus 2275: 2253: 2233:"The dinosaur 2223: 2216: 2195: 2188: 2164: 2157: 2133: 2108: 2101: 2093:The Dinosauria 2075: 2045: 2023: 2004:(1): 355–380. 1981: 1956: 1922: 1919:(11): 145–155. 1889: 1882: 1846: 1839: 1817: 1815: 1812: 1811: 1810: 1805: 1798: 1795: 1696:tyrannosaurids 1681:Talos sampsoni 1631: 1628: 1611: 1608: 1591: 1588: 1563:Prosaurolophus 1468: 1465: 1461: 1460: 1457: 1456: 1452: 1451: 1448: 1447: 1443: 1442: 1439: 1438: 1434: 1433: 1430: 1429: 1425: 1424: 1421: 1420: 1416: 1415: 1412: 1411: 1402: 1399: 1398: 1395: 1394: 1385: 1380: 1378: 1375: 1374: 1371: 1370: 1361: 1356: 1354: 1351: 1350: 1347: 1346: 1342: 1341: 1338: 1337: 1333: 1332: 1329: 1328: 1319: 1316: 1315: 1312: 1311: 1302: 1297: 1295: 1292: 1291: 1288: 1287: 1278: 1273: 1271: 1266: 1264: 1261: 1260: 1257: 1256: 1247: 1242: 1240: 1237: 1236: 1233: 1232: 1223: 1218: 1216: 1207: 1129: 1128:Classification 1126: 1058: 1055: 1051:Stephanosaurus 964:Pondera County 956:late Santonian 912:late Campanian 877: 874: 606:Lawrence Lambe 602:Red Deer River 563: 560: 533:is similar to 434: 433: 432: 431: 412: 411: 405: 404: 403: 402: 390: 371: 356: 341: 340: 336: 335: 329: 317: 316: 310: 309: 299: 295: 294: 286: 282: 281: 273: 269: 268: 260: 256: 255: 247: 240: 239: 237:Neornithischia 231: 224: 223: 215: 208: 207: 202: 195: 194: 189: 185: 184: 179: 175: 174: 169: 165: 164: 151: 150: 140: 139: 131: 130: 122: 121: 120: 115: 110: 105: 100: 95: 90: 85: 80: 75: 70: 65: 60: 59: 46: 26: 24: 14: 13: 10: 9: 6: 4: 3: 2: 5905: 5894: 5891: 5889: 5886: 5884: 5881: 5879: 5876: 5874: 5871: 5869: 5866: 5864: 5861: 5859: 5856: 5854: 5851: 5849: 5846: 5844: 5841: 5839: 5836: 5834: 5831: 5829: 5828:Saurolophines 5826: 5825: 5823: 5806: 5801: 5797: 5793: 5788: 5784: 5780: 5775: 5771: 5767: 5762: 5758: 5753: 5747: 5743: 5738: 5732: 5728: 5727: 5725: 5723: 5719: 5715: 5710: 5690: 5689: 5685: 5683: 5681: 5680:Edmontosaurus 5677: 5675: 5673: 5672:Edmontosaurus 5669: 5667: 5664: 5661: 5657: 5654: 5652: 5649: 5647: 5644: 5642: 5639: 5637: 5634: 5632: 5631: 5627: 5626: 5624: 5620: 5614: 5613: 5609: 5607: 5606: 5602: 5600: 5599: 5595: 5593: 5592: 5591:Syngonosaurus 5588: 5586: 5585: 5581: 5579: 5578: 5574: 5572: 5571: 5567: 5565: 5564: 5560: 5558: 5557: 5553: 5551: 5550: 5546: 5544: 5543: 5539: 5537: 5536: 5535:Koutalisaurus 5532: 5530: 5529: 5525: 5523: 5522: 5518: 5516: 5515: 5514:Eucercosaurus 5511: 5509: 5508: 5504: 5501: 5500: 5496: 5494: 5493: 5489: 5487: 5486: 5485:Arstanosaurus 5482: 5481: 5479: 5477: 5476: 5471: 5465: 5464: 5460: 5459: 5457: 5455: 5451: 5445: 5444: 5440: 5438: 5437: 5433: 5431: 5430: 5426: 5424: 5423: 5419: 5417: 5416: 5412: 5410: 5409: 5405: 5403: 5402: 5401:Orcauichnites 5398: 5396: 5395: 5391: 5389: 5388: 5384: 5382: 5381: 5377: 5375: 5374: 5370: 5368: 5367: 5363: 5361: 5360: 5356: 5354: 5353: 5349: 5347: 5346: 5342: 5340: 5339: 5338:Amblydactylus 5335: 5334: 5332: 5330: 5326: 5322: 5315: 5311: 5302: 5298: 5296: 5291: 5285: 5278: 5252: 5251: 5247: 5245: 5244: 5240: 5238: 5237: 5236:Kundurosaurus 5233: 5231: 5230: 5229:Kerberosaurus 5226: 5224: 5223: 5219: 5217: 5216: 5215:Edmontosaurus 5212: 5211: 5209: 5207: 5203: 5197: 5196: 5192: 5190: 5189: 5185: 5183: 5182: 5178: 5177: 5175: 5173: 5169: 5163: 5162: 5158: 5156: 5155: 5151: 5149: 5148: 5144: 5142: 5141: 5137: 5135: 5134: 5130: 5127: 5126: 5125:Anasazisaurus 5122: 5121: 5119: 5117: 5113: 5107: 5106: 5105:Secernosaurus 5102: 5100: 5099: 5098:Kelumapusaura 5095: 5093: 5092: 5088: 5086: 5085: 5081: 5080: 5078: 5076: 5072: 5065: 5064: 5060: 5058: 5057: 5053: 5051: 5050: 5046: 5044: 5043: 5039: 5037: 5036: 5032: 5030: 5029: 5025: 5024: 5022: 5020: 5016: 5010: 5009: 5005: 5004: 5001: 4998: 4996: 4995:Saurolophinae 4992: 4982: 4981: 4977: 4975: 4974: 4970: 4968: 4967: 4963: 4961: 4960: 4956: 4954: 4953: 4949: 4947: 4946: 4945:Hypacrosaurus 4942: 4940: 4939: 4938:Corythosaurus 4935: 4933: 4932: 4928: 4927: 4925: 4923: 4922:Lambeosaurini 4919: 4913: 4912: 4908: 4906: 4905: 4901: 4899: 4898: 4897:Charonosaurus 4894: 4892: 4891: 4887: 4886: 4884: 4882: 4878: 4872: 4871: 4870:Tsintaosaurus 4867: 4865: 4864: 4863:Pararhabdodon 4860: 4858: 4857: 4856:Nipponosaurus 4853: 4851: 4850: 4846: 4843: 4842: 4838: 4836: 4835: 4831: 4829: 4828: 4827:Jaxartosaurus 4824: 4822: 4821: 4817: 4815: 4814: 4810: 4808: 4807: 4803: 4801: 4800: 4796: 4794: 4793: 4789: 4787: 4786: 4782: 4780: 4779: 4778:Adynomosaurus 4775: 4774: 4771: 4768: 4766: 4762: 4756: 4755: 4751: 4749: 4748: 4744: 4742: 4741: 4740:Lapampasaurus 4737: 4735: 4734: 4730: 4728: 4727: 4723: 4721: 4720: 4716: 4715: 4712: 4709: 4706: 4705:Hadrosauridae 4701: 4695: 4694: 4690: 4688: 4687: 4686:Zhanghenglong 4683: 4681: 4680: 4676: 4674: 4673: 4669: 4667: 4666: 4662: 4660: 4659: 4658:Telmatosaurus 4655: 4653: 4652: 4648: 4646: 4645: 4641: 4639: 4638: 4634: 4632: 4631: 4627: 4625: 4624: 4620: 4618: 4617: 4613: 4611: 4610: 4606: 4604: 4603: 4602:Portellsaurus 4599: 4597: 4596: 4592: 4590: 4589: 4585: 4583: 4582: 4578: 4576: 4575: 4571: 4569: 4568: 4564: 4562: 4561: 4557: 4554: 4553: 4549: 4547: 4546: 4542: 4540: 4539: 4535: 4533: 4532: 4528: 4526: 4525: 4521: 4519: 4518: 4514: 4512: 4511: 4507: 4505: 4504: 4500: 4498: 4497: 4493: 4491: 4490: 4486: 4484: 4483: 4479: 4477: 4476: 4472: 4470: 4469: 4465: 4463: 4462: 4458: 4455: 4454: 4450: 4448: 4447: 4443: 4441: 4440: 4436: 4435: 4432: 4429: 4426: 4421: 4417: 4413: 4408: 4404: 4395: 4391: 4388: 4381: 4348: 4347: 4346: 4345: 4343: 4341: 4337: 4330: 4329: 4325: 4323: 4322: 4318: 4316: 4315: 4311: 4308: 4307: 4303: 4302: 4300: 4298: 4294: 4288: 4287: 4283: 4281: 4280: 4276: 4274: 4273: 4269: 4267: 4266: 4265:Jinzhousaurus 4262: 4260: 4259: 4258:Hypselospinus 4255: 4253: 4252: 4248: 4246: 4245: 4241: 4240: 4237: 4234: 4232: 4228: 4222: 4221: 4217: 4215: 4214: 4213:Theiophytalia 4210: 4208: 4207: 4203: 4201: 4200: 4196: 4194: 4193: 4189: 4187: 4186: 4182: 4180: 4179: 4175: 4173: 4172: 4168: 4166: 4165: 4161: 4159: 4158: 4154: 4152: 4151: 4147: 4145: 4144: 4143:Lanzhousaurus 4140: 4138: 4137: 4133: 4131: 4130: 4126: 4124: 4123: 4119: 4117: 4116: 4112: 4109: 4108: 4104: 4102: 4101: 4097: 4095: 4094: 4090: 4088: 4087: 4083: 4081: 4080: 4076: 4074: 4073: 4069: 4068: 4065: 4062: 4059: 4054: 4044: 4043: 4039: 4036: 4035: 4031: 4028: 4027: 4023: 4021: 4020: 4016: 4014: 4013: 4012:Elrhazosaurus 4009: 4007: 4006: 4002: 4000: 3999: 3995: 3994: 3992: 3990: 3986: 3980: 3979: 3975: 3973: 3972: 3968: 3966: 3965: 3961: 3960: 3957: 3953: 3950: 3947: 3942: 3928: 3927: 3923: 3921: 3920: 3916: 3914: 3913: 3909: 3907: 3906: 3902: 3900: 3899: 3895: 3893: 3892: 3888: 3887: 3885: 3883: 3879: 3873: 3872: 3871:Tenontosaurus 3868: 3866: 3865: 3861: 3858: 3857: 3856:Convolosaurus 3853: 3852: 3850: 3847: 3843: 3836: 3835: 3831: 3828: 3827: 3823: 3820: 3819: 3818:Ampelognathus 3815: 3814: 3811: 3808: 3806: 3802: 3795: 3794: 3790: 3788: 3787: 3783: 3781: 3780: 3776: 3774: 3773: 3769: 3767: 3766: 3762: 3760: 3759: 3755: 3753: 3752: 3748: 3745: 3744: 3740: 3738: 3737: 3733: 3731: 3730: 3729:Mahuidacursor 3726: 3724: 3723: 3719: 3717: 3716: 3712: 3709: 3708: 3704: 3702: 3701: 3697: 3695: 3694: 3690: 3688: 3687: 3683: 3680: 3679: 3675: 3673: 3672: 3668: 3666: 3665: 3661: 3659: 3658: 3654: 3652: 3651: 3647: 3646: 3644: 3642: 3638: 3631: 3630: 3626: 3624: 3623: 3622:Hypsilophodon 3619: 3616: 3615: 3611: 3610: 3608: 3605: 3601: 3594: 3593: 3589: 3587: 3586: 3585:Callovosaurus 3582: 3580: 3579: 3578:Burianosaurus 3575: 3574: 3571: 3567: 3564: 3561: 3556: 3552: 3548: 3543: 3539: 3514: 3513: 3512: 3511: 3509: 3507: 3503: 3495: 3491: 3490: 3489: 3488: 3485: 3482: 3480: 3476: 3468: 3464: 3463: 3462: 3461: 3458: 3455: 3453: 3449: 3445: 3437: 3433: 3430: 3428: 3424: 3421: 3419: 3415: 3412: 3410: 3406: 3403: 3401: 3397: 3395: 3391: 3390: 3385: 3381: 3377: 3370: 3365: 3363: 3358: 3356: 3351: 3350: 3347: 3340: 3329: 3321: 3315: 3311: 3303: 3300: 3295: 3291: 3287: 3283: 3279: 3275: 3268: 3265: 3252: 3248: 3244: 3237: 3234: 3228: 3225: 3218: 3215: 3211: 3205: 3202: 3197: 3193: 3189: 3185: 3181: 3177: 3173: 3169: 3162: 3160: 3156: 3145: 3141: 3135: 3132: 3120: 3116: 3110: 3107: 3094: 3090: 3086: 3079: 3076: 3063: 3059: 3053: 3050: 3037: 3033: 3026: 3023: 3018: 3014: 3009: 3004: 3000: 2996: 2994: 2985: 2983: 2979: 2974: 2968: 2964: 2960: 2954: 2951: 2946: 2942: 2938: 2934: 2928: 2925: 2920: 2916: 2912: 2906: 2902: 2901: 2893: 2890: 2885: 2881: 2877: 2871: 2868: 2863: 2859: 2855: 2849: 2846: 2840: 2837: 2832: 2828: 2824: 2820: 2816: 2810: 2807: 2802: 2798: 2794: 2788: 2785: 2773: 2769: 2765: 2761: 2757: 2753: 2749: 2745: 2741: 2734: 2731: 2719: 2715: 2709: 2706: 2695:on 2008-06-26 2694: 2690: 2686: 2680: 2677: 2672: 2666: 2662: 2658: 2657: 2649: 2646: 2641: 2635: 2631: 2627: 2623: 2616: 2614: 2610: 2605: 2599: 2595: 2590: 2589: 2583: 2577: 2575: 2573: 2571: 2569: 2567: 2563: 2558: 2554: 2550: 2546: 2542: 2538: 2534: 2530: 2526: 2519: 2517: 2515: 2511: 2506: 2502: 2498: 2497: 2489: 2486: 2475: 2471: 2465: 2462: 2457: 2453: 2449: 2445: 2441: 2437: 2433: 2429: 2425: 2421: 2417: 2410: 2407: 2402: 2401: 2393: 2390: 2385: 2381: 2377: 2373: 2369: 2365: 2361: 2354: 2351: 2345: 2340: 2336: 2332: 2328: 2326: 2317: 2315: 2311: 2306: 2302: 2298: 2294: 2288: 2286: 2284: 2282: 2280: 2276: 2271: 2267: 2263: 2257: 2254: 2249: 2245: 2238: 2236: 2227: 2224: 2219: 2213: 2209: 2205: 2199: 2196: 2191: 2185: 2181: 2177: 2176: 2168: 2165: 2160: 2154: 2150: 2146: 2145: 2137: 2134: 2130:(3): 566–577. 2129: 2125: 2121: 2115: 2113: 2109: 2104: 2098: 2094: 2090: 2084: 2082: 2080: 2076: 2071: 2067: 2063: 2059: 2055: 2049: 2046: 2041: 2037: 2033: 2027: 2024: 2019: 2015: 2011: 2007: 2003: 1999: 1995: 1991: 1990:Brown, Barnum 1985: 1982: 1977: 1973: 1969: 1963: 1961: 1957: 1952: 1948: 1944: 1940: 1936: 1932: 1931:Brown, Barnum 1926: 1923: 1918: 1914: 1910: 1906: 1902: 1896: 1894: 1890: 1885: 1879: 1875: 1870: 1869: 1863: 1857: 1855: 1853: 1851: 1847: 1842: 1836: 1832: 1825: 1823: 1819: 1813: 1809: 1806: 1804: 1801: 1800: 1796: 1794: 1792: 1791:insectivorans 1788: 1784: 1780: 1776: 1772: 1768: 1764: 1760: 1756: 1752: 1751: 1747: 1743: 1742: 1737: 1736: 1731: 1730: 1726: 1722: 1721: 1717: 1713: 1709: 1708: 1707:Teratophoneus 1703: 1702: 1701:Albertosaurus 1697: 1693: 1692: 1687: 1686:ornithomimids 1683: 1682: 1678: 1674: 1671: 1670:dromaeosaurid 1667: 1663: 1659: 1656: 1652: 1648: 1644: 1640: 1636: 1629: 1625: 1621: 1616: 1609: 1607: 1605: 1604:cartilaginous 1601: 1597: 1589: 1587: 1585: 1581: 1577: 1576: 1575:Corythosaurus 1571: 1570: 1565: 1564: 1559: 1555: 1551: 1547: 1542: 1540: 1536: 1532: 1528: 1527:monumentensis 1525: 1521: 1517: 1513: 1509: 1505: 1501: 1498: 1494: 1490: 1486: 1478: 1473: 1466: 1459: 1458: 1450: 1449: 1441: 1440: 1432: 1431: 1423: 1422: 1414: 1413: 1410: 1409: 1408: 1401: 1400: 1397: 1396: 1393: 1392: 1391: 1390:Secernosaurus 1384: 1383: 1377: 1376: 1373: 1372: 1369: 1368: 1367: 1360: 1359: 1353: 1352: 1349: 1348: 1340: 1339: 1331: 1330: 1327: 1326: 1325: 1318: 1317: 1314: 1313: 1310: 1309: 1308: 1301: 1300: 1294: 1293: 1290: 1289: 1286: 1285: 1284: 1277: 1276: 1270: 1269: 1263: 1262: 1259: 1258: 1255: 1254: 1253: 1246: 1245: 1239: 1238: 1235: 1234: 1231: 1230: 1229: 1222: 1221: 1215: 1211: 1210: 1206: 1204: 1200: 1195: 1193: 1189: 1185: 1181: 1180: 1175: 1171: 1167: 1166: 1161: 1157: 1150: 1146: 1139: 1134: 1127: 1125: 1123: 1119: 1114: 1112: 1108: 1104: 1100: 1096: 1092: 1088: 1084: 1080: 1079:type specimen 1076: 1070: 1067: 1063: 1056: 1054: 1052: 1048: 1044: 1040: 1036: 1032: 1031: 1025: 1023: 1022: 1017: 1013: 1009: 1008:Maastrichtian 1005: 1001: 997: 993: 989: 985: 981: 980:informal name 977: 974:, reflecting 973: 969: 965: 961: 957: 953: 949: 945: 941: 937: 933: 929: 925: 921: 917: 913: 909: 906: 902: 898: 894: 886: 882: 875: 873: 871: 868: 864: 860: 856: 852: 851: 846: 842: 838: 834: 830: 825: 822: 818: 814: 810: 805: 803: 799: 795: 791: 787: 782: 780: 776: 772: 768: 764: 760: 759:Bearpaw Shale 756: 752: 748: 744: 740: 736: 732: 728: 724: 720: 719: 709: 705: 703: 699: 695: 691: 687: 683: 679: 675: 671: 667: 663: 659: 655: 651: 647: 646:William Parks 643: 639: 635: 631: 627: 623: 622: 617: 616: 611: 607: 603: 599: 595: 591: 587: 583: 576: 572: 568: 561: 559: 557: 553: 550: 546: 542: 538: 537: 532: 528: 526: 522: 518: 514: 510: 506: 502: 498: 494: 493:North America 490: 487: 483: 479: 475: 472: 468: 464: 461: 457: 453: 449: 446: 442: 441: 429:Lambert, 1990 426: 425: 419: 418:Hadrosauravus 416: 415: 413: 410: 406: 397: 391: 387: 383: 378: 372: 368: 363: 357: 351: 345: 344: 342: 337: 332: 327: 326: 318: 315: 311: 308: 307: 300: 297: 296: 293: 287: 284: 283: 280: 279:Saurolophinae 274: 271: 270: 267: 266:Hadrosauridae 261: 258: 257: 254: 248: 245: 242: 241: 238: 232: 229: 226: 225: 222: 216: 213: 210: 209: 206: 203: 200: 197: 196: 193: 190: 187: 186: 183: 180: 177: 176: 173: 170: 167: 166: 161: 156: 152: 149: 145: 141: 137: 132: 118: 113: 108: 103: 98: 93: 88: 83: 78: 73: 68: 63: 57: 50: 44: 40: 37: 33: 19: 5721: 5686: 5679: 5671: 5659: 5628: 5610: 5603: 5596: 5589: 5582: 5575: 5568: 5561: 5554: 5547: 5540: 5533: 5526: 5519: 5512: 5505: 5499:Claorhynchus 5497: 5490: 5483: 5475:Nomina dubia 5473: 5463:Guegoolithus 5461: 5441: 5434: 5429:Staurichnium 5427: 5422:Sousaichnium 5420: 5413: 5406: 5399: 5392: 5385: 5378: 5373:Dakotasaurus 5371: 5366:Caririchnium 5364: 5357: 5350: 5343: 5336: 5280: 5248: 5241: 5234: 5227: 5220: 5213: 5193: 5186: 5179: 5172:Saurolophini 5159: 5152: 5145: 5139: 5138: 5131: 5123: 5116:Kritosaurini 5103: 5096: 5091:Huallasaurus 5089: 5082: 5063:Wulagasaurus 5061: 5054: 5047: 5040: 5033: 5026: 5006: 4978: 4971: 4964: 4957: 4952:Lambeosaurus 4950: 4943: 4936: 4929: 4909: 4902: 4895: 4888: 4868: 4861: 4854: 4847: 4839: 4832: 4825: 4818: 4811: 4804: 4797: 4790: 4783: 4776: 4754:Yamatosaurus 4752: 4745: 4738: 4731: 4724: 4719:Aquilarhinus 4717: 4691: 4684: 4677: 4670: 4665:Tethyshadros 4663: 4656: 4649: 4642: 4635: 4628: 4621: 4614: 4607: 4600: 4595:Plesiohadros 4593: 4586: 4579: 4572: 4567:Lophorhothon 4565: 4558: 4550: 4543: 4536: 4529: 4522: 4515: 4508: 4501: 4494: 4487: 4480: 4473: 4466: 4459: 4453:Batyrosaurus 4451: 4446:Bactrosaurus 4444: 4437: 4383: 4326: 4319: 4312: 4304: 4286:Ouranosaurus 4284: 4277: 4270: 4263: 4256: 4251:Brighstoneus 4249: 4242: 4218: 4211: 4204: 4197: 4190: 4183: 4178:Osmakasaurus 4176: 4171:Oblitosaurus 4169: 4162: 4155: 4148: 4141: 4134: 4127: 4120: 4113: 4105: 4098: 4093:Camptosaurus 4091: 4084: 4077: 4070: 4040: 4034:Kangnasaurus 4032: 4024: 4017: 4010: 4003: 3996: 3989:Dryosauridae 3976: 3969: 3962: 3924: 3917: 3910: 3903: 3896: 3891:Matheronodon 3889: 3869: 3862: 3854: 3832: 3824: 3816: 3791: 3784: 3777: 3770: 3765:Sektensaurus 3763: 3758:Qantassaurus 3756: 3749: 3741: 3734: 3727: 3720: 3713: 3707:Kangnasaurus 3705: 3698: 3691: 3684: 3676: 3671:Diluvicursor 3669: 3662: 3655: 3648: 3629:Vectidromeus 3627: 3620: 3612: 3590: 3583: 3576: 3494:Ornithischia 3479:Ornithischia 3431: 3422: 3418:Ornithischia 3413: 3404: 3309: 3302: 3277: 3273: 3267: 3255:. Retrieved 3251:the original 3246: 3236: 3227: 3217: 3204: 3174:(1): 21–43. 3171: 3168:Paleobiology 3167: 3147:. Retrieved 3143: 3134: 3123:. Retrieved 3121:. 2014-08-27 3118: 3109: 3097:. Retrieved 3093:the original 3089:NBC News.com 3088: 3078: 3066:. Retrieved 3061: 3052: 3040:. Retrieved 3035: 3025: 3001:. in press. 2998: 2992: 2962: 2953: 2944: 2940: 2936: 2927: 2899: 2892: 2883: 2879: 2870: 2861: 2857: 2848: 2839: 2833:(43): 29–48. 2830: 2826: 2822: 2818: 2809: 2800: 2796: 2787: 2777:21 September 2775:. Retrieved 2747: 2743: 2733: 2722:. Retrieved 2717: 2708: 2697:. Retrieved 2693:the original 2679: 2655: 2648: 2625: 2587: 2532: 2528: 2524: 2495: 2488: 2477:. Retrieved 2473: 2464: 2423: 2419: 2409: 2399: 2392: 2367: 2363: 2359: 2353: 2334: 2330: 2324: 2304: 2300: 2296: 2269: 2265: 2256: 2247: 2243: 2234: 2226: 2207: 2198: 2174: 2167: 2143: 2136: 2127: 2123: 2092: 2069: 2068:. Series 3. 2065: 2061: 2057: 2048: 2039: 2035: 2026: 2001: 1997: 1984: 1975: 1971: 1942: 1938: 1934: 1925: 1916: 1912: 1908: 1904: 1903:(1914). "On 1867: 1830: 1775:crocodilians 1748: 1739: 1733: 1727: 1725:ceratopsians 1718: 1705: 1699: 1689: 1679: 1661: 1660: 1633: 1619: 1610:Paleoecology 1595: 1593: 1579: 1573: 1569:Centrosaurus 1567: 1561: 1557: 1549: 1543: 1530: 1526: 1523: 1488: 1482: 1477:G. notabilis 1476: 1467:Paleobiology 1405: 1404: 1388: 1387: 1366:Aquilarhinus 1364: 1363: 1323: 1322: 1321: 1306: 1305: 1304: 1282: 1281: 1280: 1250: 1249: 1228:Wulagasaurus 1226: 1225: 1214:Kritosaurini 1202: 1196: 1191: 1187: 1183: 1177: 1174:Kritosaurini 1165:Tethyshadros 1163: 1155: 1154: 1148: 1137: 1117: 1115: 1110: 1107:G. notabilis 1106: 1102: 1094: 1086: 1082: 1074: 1073: 1066:G. notabilis 1065: 1050: 1046: 1042: 1038: 1034: 1029: 1026: 1019: 1016:saurolophine 1011: 999: 995: 987: 983: 968:G. notabilis 967: 951: 947: 940:G. notabilis 939: 935: 932:G. notabilis 931: 928:G. notabilis 927: 923: 919: 910:is from the 908:G. notabilis 907: 905:type species 900: 896: 893:G. notabilis 892: 890: 884: 858: 854: 848: 845:Kritosaurini 840: 836: 826: 816: 811:of Utah, in 806: 793: 785: 783: 770: 766: 762: 754: 750: 746: 742: 738: 734: 726: 722: 716: 714: 701: 697: 693: 689: 685: 681: 670:incurvimanus 669: 665: 661: 657: 653: 649: 641: 637: 629: 625: 619: 613: 610:Barnum Brown 581: 580: 574: 571:G. notabilis 570: 534: 530: 529: 496: 462: 447: 439: 438: 437: 422: 417: 395: 376: 361: 350:G. notabilis 349: 324: 323: 314:Type species 305: 304: 292:Kritosaurini 243: 227: 221:Ornithischia 211: 198: 143: 42: 36: 5752:Gryposaurus 5746:Wikispecies 5722:Gryposaurus 5688:Willinakaqe 5660:nomina nuda 5577:Siluosaurus 5542:Loncosaurus 5380:Dinehichnus 5222:Kamuysaurus 5195:Saurolophus 5147:Kritosaurus 5140:Gryposaurus 4973:Sahaliyania 4959:Magnapaulia 4931:Amurosaurus 4911:Tlatolophus 4890:Adelolophus 4834:Kazaklambia 4813:Blasisaurus 4806:Arenysaurus 4799:Aralosaurus 4733:Hadrosaurus 4726:Eotrachodon 4679:Yunganglong 4644:Sirindhorna 4616:Protohadros 4552:Koshisaurus 4545:Jintasaurus 4314:Comptonatus 4164:Napaisaurus 4150:Lurdusaurus 4122:Fukuisaurus 4100:Cedrorestes 4042:Valdosaurus 3905:Pareisactus 3736:Morrosaurus 3700:Isasicursor 3664:Chakisaurus 3560:Ornithopoda 3547:Ornithopoda 3506:Ornithopoda 3376:Ornithopoda 3144:EurekAlert! 3036:Reuters.com 2360:Gryposaurus 2325:Gryposaurus 2250:(1): 77–98. 2072:(4): 51–59. 2058:Kritosaurus 1935:Kritosaurus 1763:salamanders 1664:shared its 1606:extension. 1600:keratinized 1596:Gryposaurus 1580:Gryposaurus 1558:Gryposaurus 1550:Gryposaurus 1544:Like other 1533:was both a 1497:quadrupedal 1489:Gryposaurus 1485:hadrosaurid 1407:Willinakaqe 1252:Kritosaurus 1203:Gryposaurus 1192:Gryposaurus 1188:Kritosaurus 1184:Gryposaurus 1179:Kritosaurus 1170:Jack Horner 1156:Gryposaurus 1151:restoration 1149:G. alstasei 1138:Gryposaurus 1122:nasal bones 1118:Gryposaurus 1103:G. latidens 1095:Gryposaurus 1075:Gryposaurus 1057:Description 1043:Kritosaurus 1039:Gryposaurus 1035:Kritosaurus 1021:Saurolophus 1012:Kritosaurus 996:Gryposaurus 954:, from the 952:G. latidens 942:along with 897:G. latidens 855:Gryposaurus 850:Kritosaurus 841:Gryposaurus 837:Kritosaurus 794:G. latidens 786:G. latidens 771:Gryposaurus 767:Hadrosaurus 763:Gryposaurus 751:Hadrosaurus 743:Gryposaurus 739:Kritosaurus 727:Kritosaurus 723:Gryposaurus 718:Hadrosaurus 702:Kritosaurus 698:Gryposaurus 690:Kritosaurus 686:Gryposaurus 682:Kritosaurus 676:and Wright 642:Kritosaurus 638:Gryposaurus 630:Kritosaurus 626:Gryposaurus 615:Kritosaurus 582:Gryposaurus 549:quadrupedal 536:Kritosaurus 531:Gryposaurus 497:Gryposaurus 440:Gryposaurus 424:nomen nudum 396:G. alsatei? 362:G. latidens 354:Lambe, 1914 306:Gryposaurus 272:Subfamily: 253:Ornithopoda 54:80–75  43:Gryposaurus 5822:Categories 5630:Arkharavia 5605:Tichosteus 5598:Thespesius 5570:Pteropelyx 5563:Orthomerus 5443:Wintonopus 5028:Acristavus 5008:Barsboldia 4966:Olorotitan 4841:Latirhinus 4693:Zuoyunlong 4560:Levnesovia 4510:Gobihadros 4475:Datonglong 4468:Claosaurus 4439:Altirhinus 4349:see below↓ 4279:Morelladon 4157:Magnamanus 4129:Hippodraco 3998:Dryosaurus 3971:Hesperonyx 3946:Dryomorpha 3786:Trinisaura 3779:Tietasaura 3772:Talenkauen 3650:Anabisetia 3515:see below↓ 3409:Dinosauria 3257:9 November 3149:2021-08-22 3125:2021-08-22 2825:, Parks". 2724:2009-06-16 2699:2007-05-22 2505:1026402631 2479:2021-06-20 2042:: 279–302. 1814:References 1787:marsupials 1677:troodontid 1651:Appalachia 1639:Cretaceous 1590:Nasal arch 1531:Gryposaurs 1510:, and its 1047:Dinosauria 976:iguanodont 944:postcrania 859:G. alsatei 821:predentary 731:hypothesis 573:(formerly 558:settings. 541:Roman nose 471:duckbilled 205:Dinosauria 32:Gyposaurus 5612:Trachodon 5528:Glishades 5507:Diclonius 5345:Anomoepus 5329:Ichnotaxa 5042:Maiasaura 4980:Velafrons 4489:Equijubus 4461:Choyrodon 4321:Iguanodon 4185:Planicoxa 4115:Dakotadon 4086:Calvarius 3912:Rhabdodon 3898:Mochlodon 3641:Elasmaria 3392:Kingdom: 3294:131302174 3099:3 October 3068:3 October 3042:3 October 3038:. Reuters 2919:985402380 2772:133329640 2456:133329640 2448:0022-3360 2384:220413406 1951:2246/1398 1673:theropods 1647:Laramidia 1500:herbivore 1199:cladogram 1160:subfamily 867:Mescalero 804:species. 775:upper arm 678:monograph 588:specimen 552:herbivore 486:Campanian 482:Santonian 178:Kingdom: 172:Eukaryota 5731:Wikidata 5492:Cionodon 5161:Rhinorex 5049:Ornatops 4849:Minqaria 4820:Canardia 4747:Malefica 4672:Xuwulong 4637:Siamodon 4538:Jeyawati 4524:Gonkoken 4482:Eolambia 4306:Barilium 4107:Cumnoria 4072:Barilium 3978:Owenodon 3964:Draconyx 3926:Zalmoxes 3826:Fostoria 3678:Fostoria 3436:Cerapoda 3400:Chordata 3398:Phylum: 3394:Animalia 3196:88689241 3017:55969908 2961:(2002). 2803:: 25–81. 2557:83539808 2307:: 1–119. 1992:(1914). 1797:See also 1744:and the 831:and the 586:based on 515:and the 484:to late 474:dinosaur 409:Synonyms 339:Species 259:Family: 192:Chordata 188:Phylum: 182:Animalia 168:Domain: 5792:1095993 5779:4823293 5766:4531707 5737:Q385949 4785:Ajnabia 4220:Uteodon 3176:Bibcode 2947:: 1–76. 2752:Bibcode 2630:312–348 2537:Bibcode 2428:Bibcode 2006:Bibcode 1779:mammals 1771:lizards 1767:turtles 1539:browser 1508:chewing 1493:bipedal 1049:, with 972:derived 876:Species 870:Apaches 802:derived 798:bonebed 737:, uses 545:bipedal 513:Montana 505:Alberta 463:gryphus 456:griffin 386:Sampson 298:Genus: 285:Tribe: 123:↓ 5454:Ootaxa 4651:Tanius 4244:Bolong 3316:  3292:  3194:  3015:  2969:  2917:  2907:  2770:  2667:  2636:  2600:  2596:–463. 2555:  2503:  2454:  2446:  2382:  2214:  2186:  2155:  2099:  1880:  1876:–448. 1837:  1789:, and 1723:, the 1714:, the 1675:, the 1554:marine 1537:and a 1535:grazer 1504:plants 1212:  1099:limpet 903:. The 899:, and 863:Alsate 857:named 652:, not 489:stages 480:(late 452:lizard 448:grypos 388:, 2007 384:& 369:, 1992 367:Horner 333:, 1914 5805:53014 5787:IRMNG 4496:Fylax 4026:Iyuku 3432:Clade 3423:Clade 3414:Clade 3405:Clade 3290:S2CID 3192:S2CID 3013:S2CID 2768:S2CID 2553:S2CID 2452:S2CID 2380:S2CID 2240:(PDF) 1759:frogs 1698:like 1688:like 1584:river 1520:cheek 1512:teeth 1483:As a 1041:with 914:-age 887:skull 779:ilium 556:river 491:) of 467:genus 460:Latin 445:Greek 382:Gates 331:Lambe 244:Clade 228:Clade 212:Clade 199:Clade 5774:GBIF 4192:Proa 3864:Iani 3492:see 3465:see 3314:ISBN 3259:2013 3101:2007 3070:2007 3044:2007 2967:ISBN 2915:OCLC 2905:ISBN 2779:2016 2665:ISBN 2634:ISBN 2598:ISBN 2501:OCLC 2444:ISSN 2212:ISBN 2184:ISBN 2153:ISBN 2097:ISBN 2040:98-Q 1878:ISBN 1835:ISBN 1773:and 1738:and 1704:and 1655:peat 1630:Utah 1109:and 1091:skin 839:and 777:and 674:Lull 664:and 640:and 521:Utah 62:Preęž’ 5761:EoL 3282:doi 3222:pp. 3184:doi 3003:doi 2939:". 2760:doi 2594:438 2545:doi 2436:doi 2372:doi 2339:doi 2335:151 2180:161 2149:158 2064:". 2014:doi 1947:hdl 1937:". 1911:". 1874:445 1081:of 962:of 749:as 590:NMC 584:is 519:of 511:in 503:in 469:of 5824:: 5802:: 5789:: 5776:: 5763:: 5748:: 5733:: 3434:: 3425:: 3416:: 3407:: 3288:. 3278:25 3276:. 3245:. 3190:. 3182:. 3170:. 3158:^ 3142:. 3117:. 3087:. 3060:. 3034:. 3011:. 2997:. 2981:^ 2945:11 2943:. 2913:. 2884:40 2862:40 2831:38 2829:. 2799:. 2766:. 2758:. 2746:. 2742:. 2716:. 2663:. 2661:64 2632:. 2612:^ 2565:^ 2551:. 2543:. 2533:30 2531:. 2513:^ 2499:. 2472:. 2450:. 2442:. 2434:. 2424:90 2422:. 2418:. 2378:. 2368:26 2366:. 2333:. 2329:. 2313:^ 2303:. 2278:^ 2270:40 2248:51 2246:. 2242:. 2182:. 2151:. 2128:53 2126:. 2111:^ 2078:^ 2070:13 2060:, 2038:. 2012:. 2002:25 2000:. 1996:. 1976:35 1959:^ 1943:28 1941:. 1917:27 1915:. 1892:^ 1849:^ 1821:^ 1793:. 1785:, 1769:, 1765:, 1761:, 1732:, 1710:, 1694:, 1684:, 1578:. 1541:. 1524:G. 1487:, 1194:. 998:, 950:. 895:, 792:, 781:. 725:, 704:. 450:) 246:: 230:: 214:: 201:: 112:Pg 56:Ma 51:, 5662:) 5502:? 5128:? 5066:? 4844:? 4555:? 4456:? 4331:? 4309:? 4110:? 4037:? 4029:? 3859:? 3848:? 3837:? 3829:? 3821:? 3796:? 3746:? 3710:? 3681:? 3632:? 3617:? 3606:? 3595:? 3368:e 3361:t 3354:v 3322:. 3296:. 3284:: 3261:. 3198:. 3186:: 3178:: 3172:1 3152:. 3128:. 3103:. 3072:. 3046:. 3019:. 3005:: 2991:" 2975:. 2921:. 2801:3 2781:. 2762:: 2754:: 2748:1 2727:. 2702:. 2673:. 2642:. 2606:. 2559:. 2547:: 2539:: 2507:. 2482:. 2458:. 2438:: 2430:: 2386:. 2374:: 2347:. 2341:: 2305:2 2220:. 2192:. 2161:. 2105:. 2020:. 2016:: 2008:: 1953:. 1949:: 1886:. 1843:. 1495:/ 1479:. 1140:. 684:/ 547:/ 458:( 427:) 421:( 393:† 374:† 359:† 347:† 321:† 302:† 289:† 276:† 263:† 250:† 234:† 218:† 117:N 107:K 102:J 97:T 92:P 87:C 82:D 77:S 72:O 67:ęž’ 34:. 20:)

Index

Gryposaurus latidens
Gyposaurus
Late Cretaceous
Ma
Preęž’
ęž’
O
S
D
C
P
T
J
K
Pg
N

Natural History Museum of Utah
Scientific classification
Edit this classification
Eukaryota
Animalia
Chordata
Dinosauria
Ornithischia
Neornithischia
Ornithopoda
Hadrosauridae
Saurolophinae
Kritosaurini

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