37:
518:
Colonies that are grown on malt agar for one month at 25°C appear white to cream-colored, glossy, and smooth. Growth is flat on the edges and slightly raised at the center. The yeast generally does not form
348:. It has been isolated in the wild from soil samples and vineyards. Samples of a variant have been isolated from orange juice and rotten oranges. It has demonstrated potential as an organism to reduce
523:
on potato agar, but some strains have been observed to form either poorly-developed or well-developed pseudohyphae. The yeast has been observed to form one or two sherical and smooth
925:
van Wyk, Niël; Scansani, Stefano; Beisert, Beata; Brezina, Silvia; Fritsch, Stephanie; Semmler, Heike; Pretorius, Isak S.; Rauhut, Doris; von
Walbrunn, Christian (8 July 2022).
612:, have been collected from orange juice in Italy and rotten oranges in Argentina. It is also commonly found in vineyards and the fruit flies that populate vineyards.
1051:
1025:
460:
452:
365:
599:
1112:
515:, which occurs at both poles of the cell. In broth culture, sediment is present, and after one month a thin ring is formed.
574:
is variable. It has a positive growth rate at 30°C, but no growth at 37°C. It can not grow on agar media containing 0.1%
36:
504:
1056:
1102:
619:
for three days followed by the addition of commercial wine yeast resulted in an almost complete elimination of
376:
name had already been used since 1906 for an unrelated organism, Alexander Janke proposed an alternative name,
1117:
967:
559:
500:
131:
1066:
1017:
847:
796:
731:
389:
337:
153:
98:
31:
755:
1074:
999:
904:
839:
788:
723:
681:
673:
361:
88:
1079:
942:
894:
831:
780:
715:
475:. DNA Testing by S.A. Meyer in 1978 conclusively synonymized the anamorphic yeasts in the
1107:
78:
527:
with an equatorial ledge when grown for at least one week on 5% Difco malt extract agar.
1012:
871:
species using multigene sequence analysis with taxonomic implications: descriptions of
672:. Translated by Tanner, Fred Wilbur. New York: John Wiley and Sons, Inc. p. 327.
1096:
624:
575:
444:
were so similar to each other that they should be further combined into one species,
108:
800:
735:
1004:
851:
771:
Miller, M. W.; Phaff, H. J. (1958). "A Comparative Study of the
Apiculate Yeasts".
520:
503:
after 48 hours at 25°C reveals cells that are 1.8 to 6.2 μm by 3 to 11 μm in size,
990:
591:
420:, and Lodder identified as three strains of the same species, consolidated into
384:. Jacomina Lodder in 1934 found the yeast to be identical to another species,
760:(1st ed.). Amsterdam: North-Holland Publishing Company. pp. 601–605.
685:
620:
604:
563:
349:
68:
555:
551:
539:
48:
908:
792:
608:
species in Brazil. Samples of a variant of the species, proposed as var.
931:
in a
Sequential Must Inoculation to Reduce the Malic Acid Content of Wine"
899:
866:
727:
511:
or sometimes spherical, appearing singly or in pairs. Reproduction is by
1038:
984:
843:
571:
341:
947:
926:
1030:
835:
784:
719:
615:
A 2022 study found that
Muscaris wine must aerobically inoculated with
567:
547:
543:
535:
531:
512:
1043:
58:
961:
396:. A similar process occurred with yeasts identified by Klöcker as
578:
and can not utilize 2-keto-d-gluconate as a sole source of carbon.
524:
508:
333:
887:
International
Journal of Systematic and Evolutionary Microbiology
653:
Kurtzman, Cletus P.; Fell, Jack W.; Boekhout, Teun, eds. (2011).
814:
Meyer, S.A.; Smith, M.T.; Simione, F.P. (1978). "Systematics of
677:
595:
965:
388:, also isolated by Klöcker in 1912, and reclassified it as a
865:ÄŚadeĹľ, NeĹľa; Raspor, Peter; Smith, Maudv Th. (1 May 2006).
657:. Vol. 1 (5th ed.). Elsevier. pp. 601–604.
424:. Further study by Miller and Phaff in 1958 found that
380:, for the genus in 1923, which he corrected in 1928 to
451:
In 1974, yeast researcher Maudy Th. Smith observed a
594:, and other samples have been isolated from soil in
974:
623:in the wine, and produced a three-fold increase in
479:genus with their teleomorphic counterparts in the
467:yeasts were identified as separate species in the
749:
747:
745:
294:(M.Th. Smith) Tamada, Maeda & Banno 1992
499:Microscopic examination of the yeast cells in
754:Lodder, J.; Kreger-Van Rij, N. J. W. (1952).
8:
920:
918:
697:
695:
648:
646:
644:
642:
640:
962:
590:strain was isolated from a soil sample in
20:
946:
898:
455:stage of the type strain of the original
636:
773:Mycopathologia et Mycologia Applicata
360:The yeast was originally isolated by
160:
7:
1067:db6e41a6-8ad3-447e-aef8-10faa2d4f1b1
602:. It has also been collected from
873:Hanseniaspora pseudoguilliermondii
14:
586:The type species of the original
668:Guilliermond, Alexandre (1920).
588:Pseudosaccharomyces occidentalis
370:Pseudosaccharomyces occidentalis
275:Pseudosaccharomyces occidentalis
35:
471:genus, so it was designated as
368:form in 1912 and classified as
402:Pseudosaccharomyces antillarum
163:Pseudosaccharomyces antillarum
1:
757:The Yeasts: A Taxonomic Study
655:The Yeasts: A Taxonomic Study
434:Pseudosaccharomyces javanicus
259:Pseudosaccharomyces malaianus
222:(Klöcker) Kudryavtsev 1954
203:Pseudosaccharomyces javanicus
182:(Klöcker) Kudryavtsev 1954
227:Pseudosaccharomyces jensenii
867:"Phylogenetic placement of
398:Pseudosaccharomyces indicus
291:Kloeckeraspora occidentalis
243:Pseudosaccharomyces lafarii
187:Pseudosaccharomyces indicus
1134:
976:Hanseniaspora occidentalis
929:Hanseniaspora occidentalis
877:Hanseniaspora occidentalis
704:Hanseniaspora occidentalis
617:Hanseniaspora occidentalis
489:Hanseniaspora occidentalis
473:Hanseniaspora occidentalis
329:Hanseniaspora occidentalis
141:Hanseniaspora occidentalis
25:Hanseniaspora occidentalis
461:As was custom at the time
406:Pseudosaccharomyces willi
299:Pseudosaccharomyces willi
159:
152:
137:
130:
32:Scientific classification
30:
23:
463:, teleomorphic forms of
179:Hanseniaspora antillarum
1113:Fungi described in 1974
869:Hanseniaspora–Kloeckera
824:Antonie van Leeuwenhoek
708:Antonie van Leeuwenhoek
408:, which Janke moved to
310:(Klöcker) Janke 1928
286:(Klöcker) Janke 1928
270:(Klöcker) Janke 1928
254:(Klöcker) Janke 1928
238:(Klöcker) Janke 1928
214:(Klöcker) Janke 1928
198:(Klöcker) Janke 1928
174:(Klöcker) Janke 1928
702:Smith, M. T. (1974). "
530:The yeast can ferment
483:genus, and identified
457:Kloeckera occidentalis
283:Kloeckera occidentalis
219:Hanseniaspora javanica
900:10.1099/ijs.0.64052-0
315:Kloeckeria cacaoicola
442:Kloeckera antillarum
422:Kloeckera antillarum
414:Kloeckera antillarum
352:in wine production.
344:form, it was called
171:Kloeckera antillarum
123:H. occidentalis
948:10.3390/app12146919
570:. Assimilation of
374:Pseudosaccharomyces
147:M. Th. Smith (1974)
836:10.1007/BF00400078
785:10.1007/BF02055037
720:10.1007/BF00399356
562:glucose, sucrose,
485:Kloeckera javanica
446:Kloeckera javanica
430:Kloeckera javanica
426:Kloeckera jensenii
394:Kloeckera jensenii
386:Kloeckera jensenii
346:Kloeckera javanica
338:Saccharomycetaceae
267:Kloeckera malaiana
235:Kloeckera jensenii
211:Kloeckera javanica
99:Saccharomycodaceae
1090:
1089:
1075:Open Tree of Life
968:Taxon identifiers
558:. The yeast can
438:Kloeckera lafarii
325:
324:
319:
311:
303:
295:
287:
279:
271:
263:
255:
251:Kloeckera lafarii
247:
239:
231:
223:
215:
207:
199:
191:
183:
175:
167:
89:Saccharomycetales
1125:
1083:
1082:
1070:
1069:
1060:
1059:
1047:
1046:
1034:
1033:
1021:
1020:
1008:
1007:
995:
994:
993:
963:
953:
952:
950:
935:Applied Sciences
922:
913:
912:
902:
893:(5): 1157–1165.
862:
856:
855:
811:
805:
804:
768:
762:
761:
751:
740:
739:
699:
690:
689:
665:
659:
658:
650:
501:YM liquid medium
487:as a synonym of
410:Kloeckera indica
332:is a species of
317:
309:
301:
293:
285:
277:
269:
261:
253:
245:
237:
229:
221:
213:
205:
197:
195:Kloeckera indica
189:
181:
173:
165:
143:
40:
39:
21:
16:Species of yeast
1133:
1132:
1128:
1127:
1126:
1124:
1123:
1122:
1103:Saccharomycetes
1093:
1092:
1091:
1086:
1078:
1073:
1065:
1063:
1055:
1050:
1042:
1037:
1029:
1024:
1016:
1011:
1003:
998:
989:
988:
983:
970:
959:
957:
956:
924:
923:
916:
864:
863:
859:
813:
812:
808:
770:
769:
765:
753:
752:
743:
701:
700:
693:
667:
666:
662:
652:
651:
638:
633:
584:
497:
459:yeast strain.
418:Kloeckera willi
372:. Because the
358:
318:Ciferri 1931
307:Kloeckera willi
302:Klöcker 1912
278:Klöcker 1912
262:Klöcker 1912
246:Klöcker 1912
230:Klöcker 1912
206:Klöcker 1912
190:Klöcker 1912
166:Klöcker 1912
148:
145:
139:
126:
79:Saccharomycetes
34:
17:
12:
11:
5:
1131:
1129:
1121:
1120:
1118:Fungus species
1115:
1110:
1105:
1095:
1094:
1088:
1087:
1085:
1084:
1071:
1061:
1048:
1035:
1022:
1009:
996:
980:
978:
972:
971:
966:
955:
954:
914:
857:
806:
779:(2): 113–141.
763:
741:
714:(3): 441–444.
691:
660:
635:
634:
632:
629:
583:
580:
496:
493:
362:Albert Klöcker
357:
354:
336:in the family
323:
322:
321:
320:
312:
304:
296:
288:
280:
272:
264:
256:
248:
240:
232:
224:
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208:
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184:
176:
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157:
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135:
134:
128:
127:
120:
118:
114:
113:
106:
102:
101:
96:
92:
91:
86:
82:
81:
76:
72:
71:
66:
62:
61:
56:
52:
51:
46:
42:
41:
28:
27:
15:
13:
10:
9:
6:
4:
3:
2:
1130:
1119:
1116:
1114:
1111:
1109:
1106:
1104:
1101:
1100:
1098:
1081:
1076:
1072:
1068:
1062:
1058:
1053:
1049:
1045:
1040:
1036:
1032:
1027:
1023:
1019:
1014:
1010:
1006:
1001:
997:
992:
986:
982:
981:
979:
977:
973:
969:
964:
960:
949:
944:
940:
936:
932:
930:
921:
919:
915:
910:
906:
901:
896:
892:
888:
884:
882:
878:
875:sp. nov. and
874:
870:
861:
858:
853:
849:
845:
841:
837:
833:
829:
825:
821:
817:
816:Hanseniaspora
810:
807:
802:
798:
794:
790:
786:
782:
778:
774:
767:
764:
759:
758:
750:
748:
746:
742:
737:
733:
729:
725:
721:
717:
713:
709:
705:
698:
696:
692:
687:
683:
679:
675:
671:
664:
661:
656:
649:
647:
645:
643:
641:
637:
630:
628:
626:
625:ethyl acetate
622:
618:
613:
611:
607:
606:
601:
597:
593:
589:
581:
579:
577:
576:cycloheximide
573:
569:
565:
561:
557:
553:
549:
545:
541:
537:
533:
528:
526:
522:
516:
514:
510:
506:
502:
494:
492:
490:
486:
482:
481:Hanseniaspora
478:
474:
470:
469:Hanseniaspora
466:
462:
458:
454:
449:
447:
443:
439:
435:
431:
427:
423:
419:
415:
411:
407:
403:
399:
395:
391:
387:
383:
379:
375:
371:
367:
363:
355:
353:
351:
347:
343:
339:
335:
331:
330:
316:
313:
308:
305:
300:
297:
292:
289:
284:
281:
276:
273:
268:
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260:
257:
252:
249:
244:
241:
236:
233:
228:
225:
220:
217:
212:
209:
204:
201:
196:
193:
188:
185:
180:
177:
172:
169:
164:
161:
158:
155:
151:
144:
142:
136:
133:
132:Binomial name
129:
125:
124:
119:
116:
115:
112:
111:
110:Hanseniaspora
107:
104:
103:
100:
97:
94:
93:
90:
87:
84:
83:
80:
77:
74:
73:
70:
67:
64:
63:
60:
57:
54:
53:
50:
47:
44:
43:
38:
33:
29:
26:
22:
19:
975:
958:
941:(14). 6919.
938:
934:
928:
927:"The Use of
890:
886:
880:
876:
872:
868:
860:
830:(1): 79–96.
827:
823:
819:
815:
809:
776:
772:
766:
756:
711:
707:
703:
669:
663:
654:
616:
614:
609:
603:
600:Saint Thomas
587:
585:
529:
521:pseudohyphae
517:
498:
488:
484:
480:
476:
472:
468:
464:
456:
453:teleomorphic
450:
445:
441:
437:
433:
432:(originally
429:
425:
421:
417:
413:
409:
405:
401:
397:
393:
385:
381:
377:
373:
369:
359:
345:
328:
327:
326:
314:
306:
298:
290:
282:
274:
266:
258:
250:
242:
234:
226:
218:
210:
202:
194:
186:
178:
170:
162:
140:
138:
122:
121:
109:
24:
18:
592:Saint Croix
495:Description
1097:Categories
818:Zikes and
706:sp. nov".
686:1158567809
670:The Yeasts
631:References
621:malic acid
605:Drosophila
564:cellobiose
560:assimilate
538:, but not
525:ascospores
366:anamorphic
350:malic acid
340:. In its
69:Ascomycota
65:Division:
991:Q80924735
883:var. nov"
820:Kloeckera
556:trehalose
552:raffinose
540:galactose
505:apiculate
477:Kloeckera
465:Kloeckera
382:Kloeckera
378:Klöckeria
117:Species:
55:Kingdom:
49:Eukaryota
1039:MycoBank
1013:Fungorum
985:Wikidata
909:16627671
822:Janke".
801:26209678
793:13622599
736:44692118
678:20014383
572:glycerol
356:Taxonomy
342:anamorph
154:Synonyms
95:Family:
45:Domain:
1031:3480770
881:citrica
852:9985251
728:4546817
610:citrica
582:Ecology
568:salicin
548:lactose
544:maltose
536:sucrose
532:glucose
513:budding
390:synonym
364:in the
105:Genus:
85:Order:
75:Class:
1108:Yeasts
1080:404539
1064:NZOR:
1044:314870
1018:314870
907:
850:
844:566079
842:
799:
791:
734:
726:
684:
676:
566:, and
440:, and
416:, and
404:, and
1057:56407
1005:3JJYK
879:var.
848:S2CID
797:S2CID
732:S2CID
509:ovoid
334:yeast
59:Fungi
1052:NCBI
1026:GBIF
905:PMID
840:PMID
789:PMID
724:PMID
682:OCLC
674:LCCN
598:and
596:Java
534:and
1000:CoL
943:doi
895:doi
832:doi
781:doi
716:doi
554:or
448:.
436:),
392:of
1099::
1077::
1054::
1041::
1028::
1015::
1002::
987::
939:12
937:.
933:.
917:^
903:.
891:56
889:.
885:.
846:.
838:.
828:44
826:.
795:.
787:.
777:10
775:.
744:^
730:.
722:.
712:40
710:.
694:^
680:.
639:^
627:.
550:,
546:,
542:,
507:,
491:.
428:,
412:,
400:,
951:.
945::
911:.
897::
854:.
834::
803:.
783::
738:.
718::
688:.
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