4554:
Afanasiev, Gennady; Akmatov, Kunbolot; Aldashev, Almaz; Alpaslan, Ashyk; Baimbetov, Gabit; Bazaliiskii, Vladimir I.; Beisenov, Arman; Boldbaatar, Bazartseren; Boldgiv, Bazartseren; Dorzhu, Choduraa; Ellingvag, Sturla; Erdenebaatar, Diimaajav; Dajani, Rana; Dmitriev, Evgeniy; Evdokimov, Valeriy; Frei, Karin M.; Gromov, Andrey; Goryachev, Alexander; Hakonarson, Hakon; Hegay, Tatyana; Khachatryan, Zaruhi; Khaskhanov, Ruslan; Kitov, Egor; Kolbina, Alina; Kubatbek, Tabaldiev; Kukushkin, Alexey; Kukushkin, Igor; Lau, Nina; Margaryan, Ashot; Merkyte, Inga; Mertz, Ilya V.; Mertz, Viktor K.; Mijiddorj, Enkhbayar; Moiyesev, Vyacheslav; Mukhtarova, Gulmira; Nurmukhanbetov, Bekmukhanbet; Orozbekova, Z.; Panyushkina, Irina; Pieta, Karol; Smrčka, Václav; Shevnina, Irina; Logvin, Andrey; Sjögren, Karl-Göran; Štolcová, Tereza; Taravella, Angela M.; Tashbaeva, Kadicha; Tkachev, Alexander; Tulegenov, Turaly; Voyakin, Dmitriy; Yepiskoposyan, Levon; Undrakhbold, Sainbileg; Varfolomeev, Victor; Weber, Andrzej; Wilson Sayres, Melissa A.; Kradin, Nikolay; Allentoft, Morten E.; Orlando, Ludovic; Nielsen, Rasmus; Sikora, Martin; Heyer, Evelyne; Kristiansen, Kristian; Willerslev, Eske (May 2018).
7161:
Côrte-Real, Helena B.S.M.; de Knijff, Peter; Decorte, Ronny; Dubrova, Yuri E.; Evgrafov, Oleg; Gilissen, Anja; Glisic, Sanja; Gölge, Mukaddes; Hill, Emmeline W.; Jeziorowska, Anna; Kalaydjieva, Luba; Kayser, Manfred; Kivisild, Toomas; Kravchenko, Sergey A.; Krumina, Astrida; Kučinskas, Vaidutis; Lavinha, João; Livshits, Ludmila A.; Malaspina, Patrizia; Maria, Syrrou; McElreavey, Ken; Meitinger, Thomas A.; Mikelsaar, Aavo-Valdur; Mitchell, R. John; Nafa, Khedoudja; Nicholson, Jayne; Nørby, Søren; Pandya, Arpita; Parik, Jüri; Patsalis, Philippos C.; Pereira, Luísa; Peterlin, Borut; Pielberg, Gerli; Prata, Maria João; Previderé, Carlo; Roewer, Lutz; Rootsi, Siiri; Rubinsztein, D.C.; Saillard, Juliette; Santos, Fabrício R.; Stefanescu, Gheorghe; Sykes, Bryan C.; Tolun, Aslihan; Villems, Richard; Tyler-Smith, Chris; Jobling, Mark A. (December 2000).
3477:
1673:
new tree that all agreed to use. Later, a group of citizen scientists with an interest in population genetics and genetic genealogy formed a working group to create an amateur tree aiming at being above all timely. The table below brings together all of these works at the point of the landmark 2002 YCC Tree. This allows a researcher reviewing older published literature to quickly move between nomenclatures.
1394:, E-M34 is found at small frequencies in North Africa and Southern Europe (6.6% in Sicily for example), and has its highest concentration in Ethiopia and the Near East (with highest levels in Oman and Turkey). However, because the diversity is apparently low in Ethiopia, the authors suggest that E-M34 was likely introduced into Ethiopia from the Near East.
7397:
Shen, Peidong; Lavi, Tal; Kivisild, Toomas; Chou, Vivian; Sengun, Deniz; Gefel, Dov; Shpirer, Issac; Woolf, Eilon; Hillel, Jossi; Feldman, Marcus W.; Oefner, Peter J. (September 2004). "Reconstruction of patrilineages and matrilineages of
Samaritans and other Israeli populations from Y-Chromosome and
7279:
Semino, O.; Passarino, G; Oefner, PJ; Lin, AA; Arbuzova, S; Beckman, LE; De
Benedictis, G; Francalacci, P; Kouvatsi, A; Limborska, S; Marcikiae, M; Mika, A; Mika, B; Primorac, D; Santachiara-Benerecetti, AS; Cavalli-Sforza, LL; Underhill, PA (10 November 2000). "The Genetic Legacy of Paleolithic Homo
1672:
Prior to 2002, there were in academic literature at least seven naming systems for the Y-Chromosome
Phylogenetic tree. This led to considerable confusion. In 2002, the major research groups came together and formed the Y-Chromosome Consortium (YCC). They published a joint paper that created a single
6968:
Peričić, Marijana; Lauc, Lovorka Barać; Klarić, Irena
Martinović; Rootsi, Siiri; Janićijević, Branka; Rudan, Igor; Terzić, Rifet; Čolak, Ivanka; Kvesić, Ante; Popović, Dan; Šijački, Ana; Behluli, Ibrahim; Đorđević, Dobrivoje; Efremovska, Ljudmila; Bajec, Đorđe D.; Stefanović, Branislav D.; Villems,
5676:
Cruciani, Fulvio; La Fratta, Roberta; Santolamazza, Piero; Sellitto, Daniele; Pascone, Roberto; Moral, Pedro; Watson, Elizabeth; Guida, Valentina; Colomb, Eliane Beraud; Zaharova, Boriana; Lavinha, João; Vona, Giuseppe; Aman, Rashid; Calì, Francesco; Akar, Nejat; Richards, Martin; Torroni, Antonio;
4503:
Narasimhan, Vagheesh M.; Patterson, Nick; Moorjani, Priya; Rohland, Nadin; Bernardos, Rebecca; Mallick, Swapan; Lazaridis, Iosif; Nakatsuka, Nathan; Olalde, Iñigo; Lipson, Mark; Kim, Alexander M.; Olivieri, Luca M.; Coppa, Alfredo; Vidale, Massimo; Mallory, James; Moiseyev, Vyacheslav; Kitov, Egor;
1549:
remains from
Northern Israel, the Natufians carried the Y-DNA haplogroup E-Z830, a somewhat upwind clade of E-M123 (and therefore ancestral to it). The Natufians were one of the first settled peoples in the world and may have contributed to the domestication of certain crops, and thus the advent of
7360:
Semino, Ornella; Magri, Chiara; Benuzzi, Giorgia; Lin, Alice A.; Al-Zahery, Nadia; Battaglia, Vincenza; MacCioni, Liliana; Triantaphyllidis, Costas; et al. (2004), "Origin, Diffusion, and
Differentiation of Y-Chromosome Haplogroups E and J: Inferences on the Neolithization of Europe and Later
7239:
Scozzari, Rosaria; Cruciani, Fulvio; Pangrazio, Alessandra; Santolamazza, Piero; Vona, Giuseppe; Moral, Pedro; Latini, Veronica; Varesi, Laurent; Memmi, Marc M.; Romano, Valentino; De Leo, Giacomo; Gennarelli, Massimo; Jaruzelska, Jadwiga; Villems, Richard; Parik, Jüri; Macaulay, Vincent; Torroni,
5346:
Bosch, E.; Calafell, F.; Gonzalez-Neira, A.; Flaiz, C.; Mateu, E.; Scheil, H.-G.; Huckenbeck, W.; Efremovska, L.; Mikerezi, I.; Xirotiris, N.; Grasa, C.; Schmidt, H.; Comas, D. (July 2006). "Paternal and maternal lineages in the
Balkans show a homogeneous landscape over linguistic barriers, except
4960:
Adams, Susan M.; Bosch, Elena; Balaresque, Patricia L.; Ballereau, Stéphane J.; Lee, Andrew C.; Arroyo, Eduardo; López-Parra, Ana M.; Aler, Mercedes; Grifo, Marina S. Gisbert; Brion, Maria; Carracedo, Angel; Lavinha, João; Martínez-Jarreta, Begoña; Quintana-Murci, Lluis; Picornell, Antònia; Ramon,
4443:
Skourtanioti, Eirini; Erdal, Yilmaz S.; Frangipane, Marcella; Balossi
Restelli, Francesca; Yener, K. Aslıhan; Pinnock, Frances; Matthiae, Paolo; Özbal, Rana; Schoop, Ulf-Dietrich; Guliyev, Farhad; Akhundov, Tufan; Lyonnet, Bertille; Hammer, Emily L.; Nugent, Selin E.; Burri, Marta; Neumann, Gunnar
4373:
Skourtanioti, Eirini; Erdal, Yilmaz S.; Frangipane, Marcella; Balossi
Restelli, Francesca; Yener, K. Aslıhan; Pinnock, Frances; Matthiae, Paolo; Özbal, Rana; Schoop, Ulf-Dietrich; Guliyev, Farhad; Akhundov, Tufan; Lyonnet, Bertille; Hammer, Emily L.; Nugent, Selin E.; Burri, Marta; Neumann, Gunnar
5632:
Cruciani, Fulvio; Santolamazza, Piero; Shen, Peidong; Macaulay, Vincent; Moral, Pedro; Olckers, Antonel; Modiano, David; Holmes, Susan; Destro-Bisol, Giovanni; Coia, Valentina; Wallace, Douglas C.; Oefner, Peter J.; Torroni, Antonio; Cavalli-Sforza, L. Luca; Scozzari, Rosaria; Underhill, Peter A.
5167:
Behar, Doron M.; Garrigan, Daniel; Kaplan, Matthew E.; Mobasher, Zahra; Rosengarten, Dror; Karafet, Tatiana M.; Quintana-Murci, Lluis; Ostrer, Harry; Skorecki, Karl; Hammer, Michael F. (1 March 2004). "Contrasting patterns of Y chromosome variation in
Ashkenazi Jewish and host non-Jewish European
1651:
A 137-sample study of ancient Eurasian genomes found one Central Scythian who belonged to E-M123* (E-Y31991), in modern northeast Kazakhstan, dated from 800-750BC. According to the BAM file, made available by the authors, he's presumed to be E-Y168273, a clade downstream of PF4428 which is itself
1365:
found 11 E-M123 people in their 1140-person study of Iberia: 1 out of 95 Eastern Andalusians; 1 out of 100 NW Castilians; 1 out of 80 Catalans; 2 out of 52 Extramadurans; 2 out of 60 Northern Portuguese, 1 out of 78 Southern Portuguese, 1 out of 73 Southern Portuguese; 1 out of 73 Valencians; and
5759:
Cruciani, F.; La Fratta, R.; Trombetta, B.; Santolamazza, P.; Sellitto, D.; Colomb, E. B.; Dugoujon, J.-M.; Crivellaro, F.; Benincasa, T.; Pascone, R.; Moral, P.; Watson, E.; Melegh, B.; Barbujani, G.; Fuselli, S.; Vona, G.; Zagradisnik, B.; Assum, G.; Brdicka, R.; Kozlov, A. I.; Efremov, G. D.;
7160:
Rosser, Zoë H.; Zerjal, Tatiana; Hurles, Matthew E.; Adojaan, Maarja; Alavantic, Dragan; Amorim, António; Amos, William; Armenteros, Manuel; Arroyo, Eduardo; Barbujani, Guido; Beckman, Gunhild; Beckman, Lars; Bertranpetit, Jaume; Bosch, Elena; Bradley, Daniel G.; Brede, Gaute; Cooper, Gillian;
5869:
El-Sibai, Mirvat; Platt, Daniel E.; Haber, Marc; Xue, Yali; Youhanna, Sonia C.; Wells, R. Spencer; Izaabel, Hassan; Sanyoura, May F.; Harmanani, Haidar; Bonab, Maziar Ashrafian; Behbehani, Jaafar; Hashwa, Fuad; Tyler-Smith, Chris; Zalloua, Pierre A.; Genographic, Consortium. (November 2009).
4553:
Damgaard, Peter de Barros; Marchi, Nina; Rasmussen, Simon; Peyrot, Michaël; Renaud, Gabriel; Korneliussen, Thorfinn; Moreno-Mayar, J. Víctor; Pedersen, Mikkel Winther; Goldberg, Amy; Usmanova, Emma; Baimukhanov, Nurbol; Loman, Valeriy; Hedeager, Lotte; Pedersen, Anders Gorm; Nielsen, Kasper;
6449:
King, R. J.; Özcan, S. S.; Carter, T.; Kalfoğlu, E.; Atasoy, S.; Triantaphyllidis, C.; Kouvatsi, A.; Lin, A. A.; Chow, C-E. T.; Zhivotovsky, L. A.; Michalodimitrakis, M.; Underhill, P. A. (5 February 2008). "Differential Y-chromosome Anatolian Influences on the Greek and Cretan Neolithic".
4136:
K-M2313*, which as yet has no phylogenetic name, has been documented in two living individuals, who have ethnic ties to India and South East Asia. In addition, K-Y28299, which appears to be a primary branch of K-M2313, has been found in three living individuals from India. See: Poznik
1366:
highest levels apparently in the Balearics with 5 out of 37 Minorcans and 4 out of 54 Ibizans. There were none in Majorca (62 people), Gascony (24), Galicia (88), NE Castile (31), Castilla la Mancha (63), The Basque Country (116), the Asturias (20), West Andalucia (73), and Aragon (34).
5307:
Bosch, Elena; Calafell, Francesc; Comas, David; Oefner, Peter J.; Underhill, Peter A.; Bertranpetit, Jaume (2001), "High-resolution analysis of human Y-chromosome variation shows a sharp discontinuity and limited gene flow between north-western Africa and the Iberian Peninsula",
6134:
Gonçalves, Rita; Freitas, Ana; Branco, Marta; Rosa, Alexandra; Fernandes, Ana T.; Zhivotovsky, Lev A.; Underhill, Peter A.; Kivisild, Toomas; Brehm, António (July 2005). "Y-chromosome Lineages from Portugal, Madeira and Açores Record Elements of Sephardim and Berber Ancestry".
129:
The Middle East has been proposed as the most likely origin for this lineage rather than East Africa or North Africa. In fact E-M34 seems to be restricted to Semitic speakers in Ethiopia, as it has not been detected in other populations in the region such as Somalia, Kenya
5503:
Capelli, Cristian; Onofri, Valerio; Brisighelli, Francesca; Boschi, Ilaria; Scarnicci, Francesca; Masullo, Mara; Ferri, Gianmarco; Tofanelli, Sergio; et al. (2009), "Moors and Saracens in Europe: estimating the medieval North African male legacy in southern Europe",
5414:
Capelli, Cristian; Redhead, Nicola; Abernethy, Julia K.; Gratrix, Fiona; Wilson, James F.; Moen, Torolf; Hervig, Tor; Richards, Martin; Stumpf, Michael P.H.; Underhill, Peter A.; Bradshaw, Paul; Shaha, Alom; Thomas, Mark G.; Bradman, Neal; Goldstein, David B. (May 2003).
4504:
Monge, Janet; Adamski, Nicole; Alex, Neel; Broomandkhoshbacht, Nasreen; Candilio, Francesca; Callan, Kimberly; Cheronet, Olivia; Culleton, Brendan J.; Ferry, Matthew; Fernandes, Daniel; Freilich, Suzanne; Gamarra, Beatriz; et al. (5 September 2019).
6018:
Francalacci, P.; Morelli, L.; Underhill, P.A.; Lillie, A.S.; Passarino, G.; Useli, A.; Madeddu, R.; Paoli, G.; et al. (2003), "Peopling of Three Mediterranean Islands (Corsica, Sardinia, and Sicily) Inferred by Y-Chromosome Biallelic Variability",
1534:
announced the discovery of this clade. They found it in two African individuals. The authors warned that they had not yet confirmed that this clade was not a sub-clade or parent clade of either M84 or M290, so the phylogenetic position E1b1b1c1c is
7908:
Zalloua, Pierre A.; Platt, Daniel E.; El Sibai, Mirvat; Khalife, Jade; Makhoul, Nadine; Haber, Marc; Xue, Yali; Izaabel, Hassan; et al. (2008), "Identifying Genetic Traces of Historical Expansions: Phoenician Footprints in the Mediterranean",
1558:
that predates all other clades, which are not known to have existed in the region at the time (10,000 years before present). E-M123 is thought to have a TMRCA about 18,000 years ago, 8,000 years before the Natufian (possibly ancestral) remains are
7584:
Trombetta, Beniamino; Cruciani, Fulvio; Sellitto, Daniele; Scozzari, Rosaria (2011), MacAulay, Vincent (ed.), "A New Topology of the Human Y Chromosome Haplogroup E1b1 (E-P2) Revealed through the Use of Newly Characterized Binary Polymorphisms",
6543:
Lacan, Marie; Keyser, Christine; Ricaut, François-Xavier; Brucato, Nicolas; Tarrús, Josep; Bosch, Angel; Guilaine, Jean; Crubézy, Eric; Ludes, Bertrand (2011), "Ancient DNA suggests the leading role played by men in the Neolithic dissemination",
5084:
Battaglia, Vincenza; Fornarino, Simona; Al-Zahery, Nadia; Olivieri, Anna; Pala, Maria; Myres, Natalie M; King, Roy J; Rootsi, Siiri; et al. (2008), "Y-chromosomal evidence of the cultural diffusion of agriculture in southeast Europe",
4444:
U.; Penske, Sandra; Ingman, Tara; Akar, Murat; Shafiq, Rula; Palumbi, Giulio; Eisenmann, Stefanie; d'Andrea, Marta; Rohrlach, Adam B.; Warinner, Christina; Jeong, Choongwon; Stockhammer, Philipp W.; Haak, Wolfgang; Krause, Johannes (2020).
4374:
U.; Penske, Sandra; Ingman, Tara; Akar, Murat; Shafiq, Rula; Palumbi, Giulio; Eisenmann, Stefanie; d'Andrea, Marta; Rohrlach, Adam B.; Warinner, Christina; Jeong, Choongwon; Stockhammer, Philipp W.; Haak, Wolfgang; Krause, Johannes (2020).
5205:
Beleza, Sandra; Gusmao, Leonor; Lopes, Alexandra; Alves, Cintia; Gomes, Iva; Giouzeli, Maria; Calafell, Francesc; Carracedo, Angel; Amorim, Antonio (March 2006). "Micro-Phylogeographic and Demographic History of Portuguese Male Lineages".
6904:
Pelotti; Ceccardi, S; Lugaresi, F; Trane, R; Falconi, M; Bini, C; Willuweit, S; Roewer, L (2007), "Microgeographic genetic variation of Y chromosome in a population sample of Ravenna's area in the Emilia-Romagna region (North of Italy)",
6836:
Onofri, Valerio; Alessandrini, Federica; Turchi, Chiara; Pesaresi, Mauro; Buscemi, Loredana; Tagliabracci, Adriano (February 2006). "Development of multiplex PCRs for evolutionary and forensic applications of 37 human Y chromosome SNPs".
7472:
Shlush; Behar, Doron M.; Yudkovsky, Guennady; Templeton, Alan; Hadid, Yarin; Basis, Fuad; Hammer, Michael; Itzkovitz, Shalev; Skorecki, Karl (2008), Gemmell, Neil John (ed.), "The Druze: A Population Genetic Refugium of the Near East",
6747:
Mendizabal, Isabel; Sandoval, Karla; Berniell-Lee, Gemma; Calafell, Francesc; Salas, Antonio; Martinez-Fuentes, Antonio; Comas, David (2008), "Genetic origin, admixture, and asymmetry in maternal and paternal human lineages in Cuba",
7869:
Zalloua, Pierre A.; Xue, Yali; Khalife, Jade; Makhoul, Nadine; Debiane, Labib; Platt, Daniel E.; Royyuru, Ajay K.; Herrera, Rene J.; et al. (2008), "Y-Chromosomal Diversity in Lebanon Is Structured by Recent Historical Events",
5720:
Cruciani; La Fratta; Torroni; Underhill; Scozzari (2006), "Molecular Dissection of the Y Chromosome Haplogroup E-M78 (E3b1a): A Posteriori Evaluation of a Microsatellite-Network-Based Approach Through Six New Biallelic Markers",
7633:
Underhill, Peter A.; Shen, Peidong; Lin, Alice A.; Jin, Li; Passarino, Giuseppe; Yang, Wei H.; Kauffman, Erin; Bonné-Tamir, Batsheva; et al. (2000), "Y chromosome sequence variation and the history of human populations",
6047:
Fregel, Rosa; Gomes, Verónica; Gusmão, Leonor; González, Ana M; Cabrera, Vicente M; Amorim, António; Larruga, Jose M (2009), "Demographic history of Canary Islands male gene-pool: replacement of native lineages by European",
5123:
Behar, Doron M.; Thomas, Mark G.; Skorecki, Karl; Hammer, Michael F.; Bulygina, Ekaterina; Rosengarten, Dror; Jones, Abigail L.; Held, Karen; Moses, Vivian; Goldstein, David; Bradman, Neil; Weale, Michael E. (October 2003).
6668:
Maca-Meyer N, Sánchez-Velasco P, Flores C, Larruga JM, González AM, Oterino A, Leyva-Cobián F, et al. (2003), "Y Chromosome and Mitochondrial DNA Characterization of Pasiegos, a Human Isolate from Cantabria (Spain)",
6716:
Martinez, Laise; Underhill, Peter A; Zhivotovsky, Lev A; Gayden, Tenzin; Moschonas, Nicholas K; Chow, Cheryl-Emiliane T; Conti, Simon (2007), "Paleolithic Y-haplogroup heritage predominates in a Cretan highland plateau",
5913:
Firasat; Khaliq, Shagufta; Mohyuddin, Aisha; Papaioannou, Myrto; Tyler-Smith, Chris; Underhill, Peter A; Ayub, Qasim (2006), "Y-chromosomal evidence for a limited Greek contribution to the Pathan population of Pakistan",
7070:
Robino, C.; Crobu, F.; Gaetano, C.; Bekada, A.; Benhamamouch, S.; Cerutti, N.; Piazza, A.; Inturri, S.; Torre, C. (2008), "Analysis of Y-chromosomal SNP haplogroups and STR haplotypes in an Algerian population sample",
5542:
Cinnioğlu, Cengiz; King, Roy; Kivisild, Toomas; Kalfoglu, Ersi; Atasoy, Sevil; Cavalleri, Gianpiero L.; Lillie, Anita S.; Roseman, Charles C.; et al. (2004), "Excavating Y-chromosome haplotype strata in Anatolia",
5004:
Alvarez; Santos, Cristina; Montiel, Rafael; Caeiro, Blazquez; Baali, Abdellatif; Dugoujon, Jean-Michel; Aluja, Maria Pilar (2009), "Y-chromosome variation in South Iberia: Insights into the North African contribution",
4432:
Allentoft M.E., Sikora M., Sjogren K.G., Rasmussen S., Rasmussen M., Stenderup J., Damgaard P.B., Schroeder H., Ahlstrom T., Vinner L., et al. 2015 Population genomics of Bronze Age Eurasia. Nature 522(7555), 167-172.
6222:
Hassan, Hisham Y.; Underhill, Peter A.; Cavalli-Sforza, Luca L.; Ibrahim, Muntaser E. (November 2008). "Y-chromosome variation among Sudanese: Restricted gene flow, concordance with language, geography, and history".
5987:
Flores; Maca-Meyer, Nicole; Larruga, Jose M.; Cabrera, Vicente M.; Karadsheh, Naif; Gonzalez, Ana M. (2005), "Isolates in a corridor of migrations: a high-resolution analysis of Y-chromosome variation in Jordan",
6096:
Gérard, Nathalie; Berriche, Sala; Aouizérate, Annie; Diéterlen, Florent; Lucotte, Gérard (2006). "North African Berber and Arab Influences in the Western Mediterranean Revealed by Y-Chromosome DNA Haplotypes".
5043:
Arredi, B; Poloni, E; Paracchini, S; Zerjal, T; Fathallah, D; Makrelouf, M; Pascali, V; Novelletto, A; Tylersmith, C (2004), "A Predominantly Neolithic Origin for Y-Chromosomal DNA Variation in North Africa",
142:
E-M123 is best known for its major sub-clade E-M34, which dominates this clade. However, earlier studies did not test for E-M34. Looking beyond its geographical patterns, E-M123 is also quite common in many
4619:
7004:
Ramos-Luisa, E.; Blanco-Verea, A.; Brión, M.; Van Huffel, V.; Carracedo, A.; Sánchez-Diz, P. (2009), "Phylogeography of French male lineages (unpublished data 23rd International ISFG Congress)",
3972:
1646:
A study on South Asian history, Narasimhan et al. (2019), found several individuals who belonged to E-Y31991 in Late Bronze Age/Early Iron Age samples in the Swat valley, modern north Pakistan.
1313:
found 12 men out of 121 there were E-M123 positive, while in Egypt there were 7 out of 147. But in that study the Omani E-M123 diversity implied a younger age than the E-M123 found in Egypt. (
7435:
Silva, Dayse A.; Carvalho, Elizeu; Costa, Guilherme; Tavares, Lígia; Amorim, António; Gusmão, Leonor (November 2006). "Y-chromosome genetic variation in Rio De Janeiro population".
5795:
Di Gaetano; Cerutti, Francesca; Crobu, Carlo; Robino (2009), "Differential Greek and northern African migrations to Sicily are supported by genetic evidence from the Y chromosome",
3316:
4256:
Hammer, Michael F.; Behar, Doron M.; Karafet, Tatiana M.; Mendez, Fernando L.; Hallmark, Brian; Erez, Tamar; Zhivotovsky, Lev A.; Rosset, Saharon; Skorecki, Karl (November 2009).
7949:
The use of Y-chromosomal DNA variation to investigate population history; in Papiha SS, Deka R, Chakraborty R (eds): Genomic diversity: applications in human population genetics
7792:
Weale, Michael E; Shah, Tina; Jones, Abigail L; Greenhalgh, John; Wilson, James F; Nymadawa, Pagbajabyn; Zeitlin, David; Connell, Bruce A; Bradman, Neil; Thomas, Mark G (2003).
3306:
7956:
Zhao; Khan, Faisal; Borkar, Minal; Herrera, Rene; Agrawal, Suraksha (2009), "Presence of three different paternal lineages among North Indians: A study of 560 Y chromosomes",
6866:
Paracchini; Pearce, CL; Kolonel, LN; Altshuler, D; Henderson, BE; Tyler-Smith, C (2003), "A Y chromosomal influence on prostate cancer risk: the multi-ethnic cohort study",
4834:
Semino, O.; Passarino, G; Oefner, PJ; Lin, AA; et al. (2000), "The Genetic Legacy of Paleolithic Homo sapiens sapiens in Extant Europeans: A Y Chromosome Perspective",
5952:
Flores, Carlos; Maca-Meyer, Nicole; González, Ana M; Oefner, Peter J; Shen, Peidong; Pérez, Jose A; Rojas, Antonio; Larruga, Jose M; Underhill, Peter A (28 July 2004).
1417:
found about the same levels of E-M34 in Portugal as E-M123*, but E-M34 mainly in Central Portugal (4 people out of 102 tested there) with one more person found in the
1382:
found 10 out of 169 Israelis and Palestinians of various ancestry to be M123+ and M34+, with the highest level group being 4 out of 20 Israeli Jews of Libyan ancestry
6925:
Pereira, Luísa; Černý, Viktor; Cerezo, María; Silva, Nuno M; Hájek, Martin; Vašíková, Alžběta; Kujanová, Martina; Brdička, Radim; Salas, Antonio (17 March 2010).
6796:
Nebel; Filon, D; Brinkmann, B; Majumder, P; Faerman, M; Oppenheim, A (2001), "The Y Chromosome Pool of Jews as Part of the Genetic Landscape of the Middle East",
3999:
Van Oven M, Van Geystelen A, Kayser M, Decorte R, Larmuseau HD (2014). "Seeing the wood for the trees: a minimal reference phylogeny for the human Y chromosome".
4913:
Underhill, Peter A.; Shen, Peidong; Lin, Alice A.; Jin, Li; et al. (November 2000). "Y chromosome sequence variation and the history of human populations".
1667:
6252:
Henn, B. M.; Gignoux, C.; Lin, A. A.; Oefner, P. J.; Shen, P.; Scozzari, R.; Cruciani, F.; Tishkoff, S. A.; Mountain, J. L.; Underhill, P. A. (4 August 2008).
5270:
Bortolini, Maria Cátira; Thomas, Mark G.; Chikhi, Lourdes; Aguilar, Juan A.; Castro-De-Guerra, Dinorah; Salzano, Francisco M.; Ruiz-Linares, Andres (2004).
5384:
Cadenas; Zhivotovsky, Lev A; Cavalli-Sforza, Luca L; Underhill, Peter A; Herrera, Rene J (2007), "Y-chromosome diversity characterizes the Gulf of Oman",
5581:"Y-Chromosome Based Evidence for Pre-Neolithic Origin of the Genetically Homogeneous but Diverse Sardinian Population: Inference for Association Scans"
7728:
Underhill, Peter A.; Kivisild, Toomas (December 2007). "Use of Y Chromosome and Mitochondrial DNA Population Structure in Tracing Human Migrations".
5467:
Caratti; Gino, S.; Torre, C.; Robino, C. (2009), "Subtyping of Y-chromosomal haplogroup E-M78 (E1b1b1a) by SNP assay and its forensic application",
6625:
Luis, J.R.; Rowold, D.J.; Regueiro, M.; Caeiro, B.; Cinnioğlu, C.; Roseman, C.; Underhill, P.A.; Cavalli-Sforza, L.L.; Herrera, R.J. (March 2004).
1401:
found slightly more E-M34 (29) than E-M78 (26) out of 523 individuals tested (a far different E1b1b population than found in the nearby Balkans).
7034:
Regueiro, M.; Cadenas, A.M.; Gayden, T.; Underhill, P.A.; Herrera, R.J. (2006), "Iran: Tricontinental Nexus for Y-Chromosome Driven Migration",
3977:
7718:
7532:
6515:
Kujanova; Pereira; Fernandes; Pereira; Cerný (2009), "Near Eastern Neolithic Genetic Input in a Small Oasis of the Egyptian Western Desert",
6407:
7666:
UNDERHILL, P. A.; PASSARINO, G.; LIN, A. A.; SHEN, P.; MIRAZON LAHR, M.; FOLEY, R. A.; OEFNER, P. J.; CAVALLI-SFORZA, L. L. (January 2001).
5762:"Tracing Past Human Male Movements in Northern/Eastern Africa and Western Eurasia: New Clues from Y-Chromosomal Haplogroups E-M78 and J-M12"
7240:
Antonio (September 2001). "Human Y-chromosome variation in the Western Mediterranean area: implications for the peopling of the region".
5579:
Contu, Daniela; Morelli, Laura; Santoni, Federico; Foster, Jamie W.; Francalacci, Paolo; Cucca, Francesco; Hawks, John (9 January 2008).
7317:
Semino, Ornella; Santachiara-Benerecetti, A. Silvana; Falaschi, Francesco; Cavalli-Sforza, L. Luca; Underhill, Peter A. (January 2002).
4963:"The Genetic Legacy of Religious Diversity and Intolerance: Paternal Lineages of Christians, Jews, and Muslims in the Iberian Peninsula"
7363:
6798:
5046:
6594:"Y Haplogroups, Archaeological Cultures and Language Families: a Review of the Multidisciplinary Comparisons using the case of E-M35"
1460:
in their study of 275 men in Northern Africa found 2 out of 148 Tunisians from Tunis, 2 out of 19 Algerian Berbers from Tizi Ouzu in
3461:
3311:
3286:
6971:"High-Resolution Phylogenetic Analysis of Southeastern Europe Traces Major Episodes of Paternal Gene Flow Among Slavic Populations"
1510:
based on testing so far (in January 2009) that E-M84 is dominant in 6 out of the 8 clusters of E-M34 which that project identifies.
6927:"Linking the sub-Saharan and West Eurasian gene pools: maternal and paternal heritage of the Tuareg nomads from the African Sahel"
4748:"Patterns of inter- and intra-group genetic diversity in the Vlax Roma as revealed by Y chromosome and mitochondrial DNA lineages"
8005:
1575:
6487:
King; Underhill (2002), "Congruent distribution of Neolithic painted pottery and ceramic figurines with Y-chromosome lineages",
3451:
7709:
Underhill (2002). "Inference of Neolithic Population Histories using Y-chromosome Haplotypes". In Bellwood and Renfrew (ed.).
5954:"Reduced genetic structure of the Iberian peninsula revealed by Y-chromosome analysis: implications for population demography"
5679:"Phylogeographic Analysis of Haplogroup E3b (E-M215) Y Chromosomes Reveals Multiple Migratory Events Within and Out Of Africa"
261:
Y-chromosomal analysis of Greek Cypriots reveals a primarily common pre-Ottoman paternal ancestry with Turkish Cypriots. 2017
5416:
102:
5635:"A Back Migration from Asia to Sub-Saharan Africa Is Supported by High-Resolution Analysis of Human Y-Chromosome Haplotypes"
6400:
In Hot Pursuit of Language in Prehistory: Essays in the Four Fields of Anthropology : in Honor of Harold Crane Fleming
4640:"A Predominantly Indigenous Paternal Heritage for the Austronesian-Speaking Peoples of Insular Southeast Asia and Oceania"
1323:
found 4.66% overall in their 236-person study of Sicily, with higher levels in the east of the island. They found none in
6593:
1372:
found 9 out of 323 people in 3 areas of Sardinia. 4 out of 187 in Cagliari, 1 out of 103 in Sorgono, and 4 out of 86 in
7835:
Y Chromosome Consortium "YCC" (2002), "A Nomenclature System for the Tree of Human Y-Chromosomal Binary Haplogroups",
3256:
6347:
Karafet, T. M.; Mendez, F. L.; Meilerman, M. B.; Underhill, P. A.; Zegura, S. L.; Hammer, M. F. (21 February 2008).
6319:
Jobling, M.A.; Tyler-Smith, C. (2000), "New uses for new haplotypes the human Y chromosome, disease and selection",
1490:, out of 164 men tested; and 27 Palestinians out of 291 tested. This was apparently higher than the level of E-M78.
1270:
4118:
Haplogroup K2b1 (P397/P399) is also known as Haplogroup MS, but has a broader and more complex internal structure.
1567:
7522:
3640:
1205:
found 1 Tunisian from Tunis in their study of 275 men in Northern Africa, which included 148 people from Tunis.
5244:
3580:
7163:"Y-Chromosomal Diversity in Europe Is Clinal and Influenced Primarily by Geography, Rather than by Language"
5126:"Multiple Origins of Ashkenazi Levites: Y Chromosome Evidence for Both Near Eastern and European Ancestries"
4872:"Y-Chromosome Evidence for a Northward Migration of Modern Humans into Eastern Asia during the Last Ice Age"
4164:
Haplogroup S, as of 2017, is also known as K2b1a. (Previously the name Haplogroup S was assigned to K2b1a4.)
3843:
3570:
3291:
7204:
Sanchez, Juan J; Hallenberg, Charlotte; Børsting, Claus; Hernandez, Alexis; Morling, Niels (9 March 2005).
4173:
Haplogroup M, as of 2017, is also known as K2b1b. (Previously the name Haplogroup M was assigned to K2b1d.)
3667:
3650:
7249:
6678:
4638:
Capelli, Cristian; Wilson, James F.; Richards, Martin; Stumpf, Michael P.H.; et al. (February 2001).
3832:
3824:
3564:
3429:
233:
Comparative Y-chromosome analysis among Cypriots in the context of historical events and migrations. 2021
6182:
Hammer (2003), "Human population structure and its effects on sampling Y chromosome sequence variation",
3658:
2264:
5872:"Geographical Structure of the Y-chromosomal Genetic Landscape of the Levant: A coastal-inland contrast"
3894:
3740:
3271:
5245:"Haplogroup E3b1a2 as a Possible Indicator of Settlement in Roman Britain by Soldiers of Balkan Origin"
4746:
Kaladjieva, Luba; Calafell, Francesc; Jobling, Mark A; Angelicheva, Dora; et al. (February 2001).
3817:
3810:
3803:
3413:
3353:
3338:
3330:
3055:
2874:
2813:
2691:
2447:
2386:
2325:
1507:
7206:"High frequencies of Y chromosome lineages characterized by E3b1, DYS19-11, DYS392-12 in Somali males"
3403:
3122:
The following research teams per their publications were represented in the creation of the YCC tree.
2994:
2508:
2203:
2020:
1898:
7594:
7482:
7289:
7120:
6757:
6553:
6349:"New binary polymorphisms reshape and increase resolution of the human Y chromosomal haplogroup tree"
6265:
6057:
5592:
5431:
4961:
Misericordia; Skorecki, Karl; Behar, Doron M.; Calafell, Francesc; Jobling, Mark A. (December 2008).
4843:
4683:
Hammer, Michael F.; Karafet, Tatiana M.; Redd, Alan J.; Jarjanazi, Hamdi; et al. (1 July 2001).
4567:
4317:
3933:
3852:
3787:
3731:
3716:
3702:
3623:
3595:
3266:
1356:
7254:
6683:
6391:
4316:
Lazaridis, Iosif; et al. (17 June 2016), "The genetic structure of the world's first farmers",
4209:
had records of four E-M123* tests (20 records; 26 May 2017), compared to 93 test results with E-M34.
3926:
3921:
7711:
Examining the farming/language dispersal hypothesis, McDonald Institute for Archaeological Research
3444:
7757:
Weale, Michael E.; Weiss, Deborah A.; Jager, Rolf F.; Bradman, Neil; Thomas, Mark G. (July 2002).
7668:"The phylogeography of Y chromosome binary haplotypes and the origins of modern human populations"
7697:
7655:
7460:
7423:
7096:
7059:
7023:
6704:
6504:
6475:
6437:
6184:
6170:
6122:
5858:
5748:
5568:
5492:
5455:
5372:
5231:
5193:
5032:
4938:
4777:
4601:
4485:
4415:
4024:
3862:
3536:
3519:
1521:
6627:"The Levant versus the Horn of Africa: Evidence for Bidirectional Corridors of Human Migrations"
5272:"Ribeiro's typology, genomes, and Spanish colonialism, as viewed from Gran Canaria and Colombia"
4620:"Genetic Landscape of the West Eurasian Steppe before and after the Scythian Dominance - Page 4"
4258:"Extended Y chromosome haplotypes resolve multiple and unique lineages of the Jewish priesthood"
1550:
agriculture. The discovery of E-Z830 (without other clades) suggests an indigenous presence in
1336:
7983:
7936:
7897:
7858:
7823:
7780:
7745:
7714:
7689:
7647:
7622:
7572:
7528:
7510:
7452:
7415:
7386:
7348:
7305:
7267:
7227:
7192:
7148:
7107:
Rosa, Alexandra; Ornelas, Carolina; Jobling, Mark A; Brehm, António; Villems, Richard (2007).
7088:
7051:
6992:
6956:
6893:
6854:
6825:
6785:
6736:
6696:
6656:
6581:
6532:
6467:
6403:
6378:
6336:
6293:
6240:
6211:
6162:
6114:
6085:
6036:
6007:
5975:
5941:
5901:
5850:
5822:
5783:
5740:
5708:
5664:
5620:
5560:
5531:
5484:
5447:
5403:
5364:
5335:
5223:
5185:
5155:
5112:
5073:
5024:
4992:
4930:
4901:
4859:
4822:
4791:"Paternal Population History of East Asia: Sources, Patterns, and Microevolutionary Processes"
4769:
4735:
4706:
4671:
4593:
4535:
4477:
4407:
4287:
4093:
4074:
F-Y27277, sometimes known as F2'4, is both the parent clade of F2 and F4 and a child of F-M89.
4016:
3954:
3949:
3940:
3883:
3874:
3794:
3765:
3725:
3683:
3635:
3553:
3418:
3373:
3368:
3343:
3261:
1613:
59:
4446:"Genomic History of Neolithic to Bronze Age Anatolia, Northern Levant, and Southern Caucasus"
4376:"Genomic History of Neolithic to Bronze Age Anatolia, Northern Levant, and Southern Caucasus"
7973:
7965:
7926:
7918:
7887:
7879:
7848:
7840:
7813:
7805:
7770:
7737:
7679:
7639:
7612:
7602:
7562:
7554:
7500:
7490:
7444:
7407:
7376:
7368:
7338:
7330:
7297:
7259:
7217:
7182:
7174:
7138:
7128:
7080:
7043:
7013:
6982:
6946:
6938:
6914:
6883:
6875:
6846:
6815:
6807:
6775:
6765:
6726:
6688:
6646:
6638:
6571:
6561:
6524:
6496:
6459:
6429:
6418:"Genetics, Egypt, and History: Interpreting Geographical Patterns of Y Chromosome Variation"
6395:
6368:
6360:
6328:
6283:
6273:
6232:
6201:
6193:
6152:
6144:
6106:
6075:
6065:
6028:
5997:
5965:
5931:
5923:
5891:
5883:
5842:
5812:
5804:
5773:
5730:
5698:
5690:
5654:
5646:
5610:
5600:
5552:
5521:
5513:
5476:
5439:
5393:
5356:
5325:
5317:
5293:
5283:
5215:
5177:
5145:
5137:
5102:
5094:
5063:
5055:
5014:
4982:
4974:
4922:
4891:
4883:
4851:
4812:
4802:
4759:
4727:
4696:
4661:
4651:
4583:
4575:
4525:
4517:
4467:
4457:
4397:
4387:
4277:
4269:
4008:
3916:
3674:
3628:
3408:
3393:
3388:
3383:
3363:
3358:
3348:
2752:
2630:
2569:
1959:
1837:
1776:
1564:
1332:
1181:
also found two E-M123* individuals in Northern Portugal, out of 101 people, as well as 2 in
144:
7741:
7775:
7758:
4701:
4684:
3688:
3398:
3378:
2142:
2081:
1605:
1408:
1352:
109:
M123. Like its closest relatives within the larger E-M215 haplogroup, E-M123 is found in
17:
7598:
7486:
7293:
7124:
6761:
6557:
6269:
6061:
5596:
5435:
4847:
4571:
3490:
Please help update this article to reflect recent events or newly available information.
1174:
found two E-M123* individuals, both in Northern Portugal out of 109 people tested there.
7978:
7931:
7892:
7818:
7793:
7684:
7667:
7617:
7567:
7542:
7505:
7381:
7187:
7162:
7143:
7109:"Y-chromosomal diversity in the population of Guinea-Bissau: a multiethnic perspective"
7108:
6951:
6926:
6888:
6820:
6780:
6651:
6626:
6576:
6373:
6348:
6288:
6254:"Y-chromosomal evidence of a pastoralist migration through Tanzania to southern Africa"
6253:
6206:
6080:
5936:
5896:
5871:
5817:
5703:
5678:
5615:
5580:
5526:
5330:
5150:
5125:
5107:
5068:
4987:
4962:
4896:
4871:
4817:
4790:
4666:
4639:
4530:
4505:
4282:
4257:
4206:
3904:
3697:
3530:
3437:
1155:
after testing 3401 individuals from five continents (of which 116 were Bulgarian), and
497:
84:
7853:
7343:
7318:
7263:
6332:
5659:
5634:
5443:
4789:
Karafet, Tatiana; Xu, Liping; Du, Ruofu; Wang, William; et al. (September 2001).
4731:
4041:
1503:, defined by SNP mutation M84, with M136 defining a sub-clade as of October 2008. The
1464:(10.5%), and 3 out of 44 North Egyptians, 4 out of 29 South Egyptians (So 9.5% in all
1226:
found none amongst the significant presence of E-M34 they found in their study of the
7999:
7319:"Ethiopians and Khoisan Share the Deepest Clades of the Human Y-Chromosome Phylogeny"
6692:
6508:
6463:
6441:
6148:
5887:
5360:
5219:
4870:
Su, Bing; Xiao, Junhua; Underhill, Peter; Deka, Ranjan; et al. (December 1999).
4489:
4419:
4097:
3779:
3773:
3759:
3748:
3711:
3614:
3609:
3600:
3586:
3547:
1344:
448:
7659:
7464:
7100:
7027:
6850:
6708:
6479:
6126:
5752:
5572:
5376:
5288:
5271:
5197:
4942:
4781:
4605:
4555:
4028:
7701:
7427:
7063:
6174:
5862:
5846:
5496:
5235:
5036:
1604:
A study on the population genomics of Bronze Age Eurasia found in the remains from
1599:
one individual who lived circa 3369-3110 BC, who belonged to E-M84 (E1b1b1b2a1a1~).
1480:
found one in South Iran out of 117 people, and none in North Iran out of 33 people.
1258:
E-M123 has sometimes been reported without checking for the M-34 SNP, for example:
1160:
95:
88:
7301:
6197:
5459:
4855:
7809:
7607:
7495:
7018:
6918:
5605:
1355:(10.71% of 28 people), as well as near the Southwestern extreme facing Africa at
6309:
49:
7922:
7883:
7794:"Rare deep-rooting Y chromosome lineages in humans: lessons for phylogeography"
6396:"Geography, selected Afro-Asiatic families, and Y chromosome lineage variation"
4978:
4462:
4445:
4392:
4375:
1250:
found none in their study of 169 Israelis and Palestinians of various ancestry.
7969:
7543:"Evidence for an apartheid-like social structure in early Anglo-Saxon England"
7084:
6608:
6500:
6002:
5556:
5480:
5181:
4579:
4273:
3276:
1638:
1192:
98:
4472:
4402:
4149:
1504:
7222:
7205:
7133:
6987:
6970:
6770:
6731:
6566:
6278:
6070:
5970:
5953:
5927:
5778:
5761:
5398:
4764:
4747:
4718:
Jobling, Mark A.; Tyler-Smith, Chris (2000), "New uses for new haplotypes",
4521:
3296:
3281:
1474:
found 3 in their 168-person study of Crete, 2 in Heraklion and 1 in Lasithi.
1465:
1407:
reported E-M34 in several parts of Iberia, but most strikingly about 10% in
152:
148:
7987:
7940:
7901:
7862:
7827:
7784:
7749:
7693:
7651:
7626:
7576:
7558:
7514:
7456:
7419:
7390:
7352:
7309:
7271:
7231:
7196:
7152:
7092:
7055:
6996:
6960:
6897:
6858:
6829:
6789:
6740:
6700:
6660:
6585:
6536:
6471:
6382:
6340:
6297:
6244:
6215:
6166:
6118:
6089:
6040:
6011:
5979:
5945:
5905:
5854:
5826:
5787:
5744:
5712:
5668:
5624:
5564:
5535:
5488:
5451:
5407:
5368:
5339:
5227:
5189:
5159:
5116:
5077:
5028:
4996:
4934:
4905:
4863:
4826:
4773:
4739:
4710:
4675:
4597:
4539:
4481:
4411:
4291:
4020:
1418:
6879:
6433:
6110:
5808:
5517:
5098:
6942:
6157:
3301:
1626:
1622:
1588:
1584:
1546:
1439:
1429:
1300:
1223:
1152:
106:
4588:
4056:
Haplogroup A0-T is also known as A-L1085 (and previously as A0'1'2'3'4).
7448:
7411:
6528:
6364:
6236:
6032:
5298:
5019:
4012:
1609:
1570:
has been found in 75% of the 'Ain Ghazal population, along with 60% of
1461:
1347:(28 people) along the south coast; and then high levels in the east in
1340:
1324:
1317:
tested for E-M34 in Oman and found 7.7% to be E-M34+, with no E-M123*.)
1296:
1274:
1182:
1163:
out of 1062 people tested, including 184 from Central Asia and Siberia.
452:
7844:
7047:
6417:
5833:
Ehret, C.; Keita, SO; Newman, P (2004), "The Origins of Afroasiatic",
5735:
4142:
4096:
was known as "Haplogroup K2". That name has since been re-assigned to
1144:
Such cases are relatively rare, but the following have been reported.
7541:
Thomas, Mark G; Stumpf, Michael P.H; Härke, Heinrich (18 July 2006).
7524:
Blood of the Isles: Exploring the Genetic Roots of Our Tribal History
4336:
4243:
4230:
4153:, 2017, "Details of the Y-SNP markers included in the minimal Y tree"
1630:
1592:
1555:
1551:
1487:
1391:
1373:
1348:
1328:
1314:
1266:
1167:
1148:
131:
118:
114:
4359:
7372:
7334:
7178:
6811:
6642:
5694:
5650:
5321:
5256:
5141:
5059:
4887:
4807:
4656:
4322:
4109:
Haplogroup K2b (M1221/P331/PF5911) is also known as Haplogroup MPS.
1295:
reports relatively high levels of 13% in the Albanian community of
7643:
7280:
sapiens sapiens in Extant Europeans: A Y Chromosome Perspective".
6305:
4926:
1596:
1471:
1447:
1443:
1433:
1286:
1241:
1235:
1231:
1196:
1531:
1634:
1571:
1432:
area of Jordan to be M34 positive (31.1%), while in the capital
1304:
1262:
155:
110:
3433:
4685:"Hierarchical Patterns of Global Human Y-Chromosome Diversity"
4506:"The formation of human populations in South and Central Asia"
3470:
1477:
1451:
1425:
1404:
1386:
And E-M34 has sometimes been tested without testing for M123:
1227:
1188:
1171:
7713:. Cambridge: McDonald Institute for Archaeological Research.
1398:
1362:
1217:
4556:"137 ancient human genomes from across the Eurasian steppes"
1457:
1320:
1202:
1483:
6907:
Forensic Science International: Genetics Supplement Series
4042:
International Society of Genetic Genealogy (ISOGG; 2015),
1414:
1208:
Studies which tested for E-M123* but found none include...
1178:
5760:
Coppa, A.; Novelletto, A.; Scozzari, R. (10 March 2007).
4083:
Haplogroup LT (L298/P326) is also known as Haplogroup K1.
1369:
1446:
were positive for M34, compared to much lower levels in
1335:(16 people) along the west of the north coast; 3.23% in
1310:
7547:
Proceedings of the Royal Society B: Biological Sciences
6314:, International Society of Genetic Genealogists "ISOGG"
3130:
7759:"Y Chromosome Evidence for Anglo-Saxon Mass Migration"
3317:
Y-DNA haplogroups in populations of Sub-Saharan Africa
1303:. A notably high regional frequency for E-M123 was in
1280:
1307:, where it is apparently the dominant clade of E-M35.
1292:
159:
7951:, Kluwer Academic/Plenum Publishers, pp. 91–101
3307:
Y-chromosome haplogroups in populations of the world
1517:
1379:
1247:
3512:
1641:, one male individual who is E-M84 (E1b1b1b2a1a1~).
1156:
73:
65:
55:
45:
37:
32:
5677:Novelletto, Andrea; Scozzari, Rosaria (May 2004).
1195:, this being one of the 101 individuals tested in
7367:, vol. 74, no. 5, pp. 1023–1034,
7073:Journal International Journal of Legal Medicine
6258:Proceedings of the National Academy of Sciences
7839:, vol. 12, no. 2, pp. 339–348,
7638:, vol. 26, no. 3, pp. 358–361,
4155:(Access date of these pages: 9 December 2017)
1545:According to the genetic analyses done on six
1244:found none in their 168 person study of Crete.
7361:Migratory Events in the Mediterranean Area",
3445:
1668:Conversion table for Y chromosome haplogroups
1140:E-M123* (tested and definitely without E-M34)
158:, accounting for over 10% of all male lines (
8:
5417:"A Y Chromosome Census of the British Isles"
6311:Y-DNA Haplogroup E and its Subclades - 2011
4127:Haplogroup P (P295) is also klnown as K2b2.
3452:
3438:
3430:
3134:
164:
7977:
7930:
7891:
7852:
7817:
7774:
7683:
7616:
7606:
7566:
7504:
7494:
7380:
7342:
7253:
7221:
7186:
7142:
7132:
7017:
6986:
6950:
6887:
6819:
6779:
6769:
6730:
6682:
6650:
6575:
6565:
6517:American Journal of Physical Anthropology
6416:Keita, S.O.Y.; Boyce, A.J. (9 May 2014).
6372:
6287:
6277:
6225:American Journal of Physical Anthropology
6205:
6156:
6079:
6069:
6021:American Journal of Physical Anthropology
6001:
5969:
5935:
5895:
5816:
5777:
5734:
5702:
5658:
5614:
5604:
5525:
5397:
5329:
5297:
5287:
5149:
5106:
5067:
5018:
4986:
4895:
4816:
4806:
4763:
4700:
4665:
4655:
4587:
4529:
4471:
4461:
4401:
4391:
4321:
4281:
3142:
1574:populations circa 8,800–6,500 BC. (See:
1339:(31 people) inland in the west; 3.57% in
4065:Haplogroup A1 is also known as A1'2'3'4.
1675:
7398:mitochondrial DNA sequence Variation".
5469:International Journal of Legal Medicine
4222:
4198:
3991:
3182:
3158:
182:Natufians (Northern Israel, 10,000 ybp)
7911:The American Journal of Human Genetics
7742:10.1146/annurev.genet.41.110306.130407
7323:The American Journal of Human Genetics
7167:The American Journal of Human Genetics
6969:Richard; Rudan, Pavao (October 2005).
6631:The American Journal of Human Genetics
5683:The American Journal of Human Genetics
5639:The American Journal of Human Genetics
5130:The American Journal of Human Genetics
4967:The American Journal of Human Genetics
4876:The American Journal of Human Genetics
4795:The American Journal of Human Genetics
4644:The American Journal of Human Genetics
3166:
3150:
29:
7776:10.1093/oxfordjournals.molbev.a004160
4702:10.1093/oxfordjournals.molbev.a003906
4304:
1428:found 14 out of 45 men tested in the
7:
4205:As of 11 November 2008 for example,
3978:Y-DNA haplogroups of historic people
3953:
3948:
3939:
3937:
3932:
3930:
3925:
3920:
3915:
3909:
3902:
3892:
3890:
3880:
3872:
3860:
3858:
3849:
3841:
3799:
3792:
3786:
3784:
3778:
3772:
3770:
3764:
3758:
3746:
3736:
3730:
3724:
3715:
3710:
3701:
3696:
3687:
3682:
3646:
3632:
3515:
3620:
3560:
3543:
1220:, with 523 individuals in Anatolia;
1177:In a 553-person study of Portugal,
7872:American Journal of Human Genetics
7685:10.1046/j.1469-1809.2001.6510043.x
7364:American Journal of Human Genetics
7210:European Journal of Human Genetics
6931:European Journal of Human Genetics
6799:American Journal of Human Genetics
6719:European Journal of Human Genetics
5958:European Journal of Human Genetics
5916:European Journal of Human Genetics
5797:European Journal of Human Genetics
5506:European Journal of Human Genetics
5386:European Journal of Human Genetics
5087:European Journal of Human Genetics
5047:American Journal of Human Genetics
4752:European Journal of Human Genetics
3174:
1621:A study on population genomics of
1583:A study on population genomics of
147:communities, including among both
25:
7437:American Journal of Human Biology
5007:American Journal of Human Biology
4047:. (Access date: 1 February 2015.)
3312:Y-DNA haplogroups by ethnic group
3287:Human Y-chromosome DNA haplogroup
6693:10.1046/j.1469-1809.2003.00045.x
6464:10.1111/j.1469-1809.2007.00414.x
6149:10.1111/j.1529-8817.2005.00161.x
5888:10.1111/j.1469-1809.2009.00538.x
5361:10.1111/j.1469-1809.2005.00251.x
5220:10.1111/j.1529-8817.2005.00221.x
3475:
1516:, defined by SNP mutation M290.
1283:also found examples in Portugal.
1191:found one individual out of 146
7763:Molecular Biology and Evolution
6975:Molecular Biology and Evolution
6851:10.1016/j.forsciint.2005.03.014
5766:Molecular Biology and Evolution
5289:10.1590/S1415-47572004000100001
4689:Molecular Biology and Evolution
4100:, the sibling of Haplogroup LT.
1530:, defined by SNP mutation V23.
1442:found 8.1% of 62 men tested in
1185:out of 129 people tested there.
7006:Forensic Science International
6839:Forensic Science International
5847:10.1126/science.306.5702.1680c
5276:Genetics and Molecular Biology
103:single nucleotide polymorphism
1:
7302:10.1126/science.290.5494.1155
7264:10.1016/S0198-8859(01)00286-5
6333:10.1016/S0168-9525(00)02057-6
5444:10.1016/s0960-9822(03)00373-7
4856:10.1126/science.290.5494.1155
4732:10.1016/S0168-9525(00)02057-6
4631:Sources for conversion tables
1633:found in the ancient city of
1436:there were only 4 out of 101.
378:Mazara del Vallo Sicily Italy
27:Human Y-chromosome haplogroup
7608:10.1371/journal.pone.0016073
7496:10.1371/journal.pone.0002105
7019:10.1016/j.fsigss.2009.09.026
6919:10.1016/j.fsigss.2007.10.025
6601:Journal of Genetic Genealogy
5606:10.1371/journal.pone.0001430
5249:Journal of Genetic Genealogy
3462:Y-chromosome DNA haplogroups
3131:Jobling and Tyler-Smith 2000
1768:
1763:
1758:
1753:
1748:
1743:
1738:
1733:
1728:
1723:
1718:
1713:
1708:
1703:
1698:
1693:
1688:
1683:
1678:
1289:found one sample in Somalia.
392:Piazza Armerina Sicily Italy
6198:10.1093/genetics/164.4.1495
5347:for the isolated Aromuns".
4338:Y-DNA E-M123; A Closer Look
4207:the E-M35 phylogeny project
3460:Phylogenetic tree of human
3257:African admixture in Europe
8022:
7923:10.1016/j.ajhg.2008.10.012
7884:10.1016/j.ajhg.2008.01.020
7810:10.1093/genetics/165.1.229
6592:Lancaster, Andrew (2009),
4979:10.1016/j.ajhg.2008.11.007
4463:10.1016/j.cell.2020.04.044
4433:(doi:10.1038/nature14507).
4393:10.1016/j.cell.2020.04.044
4044:Y-DNA Haplogroup Tree 2015
3946:
3913:
3870:
3839:
3830:
3822:
3815:
3808:
3801:
3756:
3722:
3708:
3694:
3680:
3664:
3648:
3517:
1665:
1159:located one individual in
1151:located one individual in
464:Di Cristofaro et al. 2013
7970:10.1080/03014460802558522
7730:Annual Review of Genetics
7085:10.1007/s00414-007-0203-5
6501:10.1017/S0003598X00091158
6003:10.1007/s10038-005-0274-4
5557:10.1007/s00439-003-1031-4
5481:10.1007/s00414-009-0350-y
5182:10.1007/s00439-003-1073-7
4580:10.1038/s41586-018-0094-2
4274:10.1007/s00439-009-0727-5
3967:
3745:
3738:
3672:
3655:
3639:
3634:
3627:
3622:
3613:
3608:
3606:
3599:
3594:
3592:
3585:
3578:
3576:
3569:
3562:
3552:
3545:
3535:
3528:
3526:
3510:
3484:This article needs to be
3468:
1680:YCC 2002/2008 (Shorthand)
1486:found 26 E-M123 cases in
1265:found E-M123 examples in
1166:In a 568-person study in
18:Haplogroup E-M123 (Y-DNA)
7672:Annals of Human Genetics
7113:BMC Evolutionary Biology
6671:Annals of Human Genetics
6452:Annals of Human Genetics
6137:Annals of Human Genetics
6050:BMC Evolutionary Biology
5876:Annals of Human Genetics
5349:Annals of Human Genetics
5208:Annals of Human Genetics
1321:Di Gaetano et al. (2009)
188:40-100 (incomplete data)
77:M123, L798.1, L799, L857
46:Possible place of origin
8006:Human Y-DNA haplogroups
7958:Annals of Human Biology
7223:10.1038/sj.ejhg.5201390
7134:10.1186/1471-2148-7-124
6771:10.1186/1471-2148-8-213
6732:10.1038/sj.ejhg.5201769
6567:10.1073/pnas.1113061108
6279:10.1073/pnas.0801184105
6071:10.1186/1471-2148-9-181
5971:10.1038/sj.ejhg.5201225
5928:10.1038/sj.ejhg.5201726
5399:10.1038/sj.ejhg.5201934
4765:10.1038/sj.ejhg.5200597
4522:10.1126/science.aat7487
4092:Between 2002 and 2008,
3292:Molecular phylogenetics
1532:Trombetta et al. (2011)
1505:E-M35 Phylogeny Project
1415:Gonçalves et al. (2005)
1399:Cinnioğlu et al. (2004)
1218:Cinnioğlu et al. (2004)
1179:Gonçalves et al. (2005)
1157:Underhill et al. (2000)
1114:Karachanak et al. 2013
937:Pasiegos from Cantabria
904:Yunusbayev et al. 2011
502:non-Levite, non-Cohanim
415:Di Gaetano et al. 2009
401:Di Gaetano et al. 2009
387:Di Gaetano et al. 2009
38:Possible time of origin
7559:10.1098/rspb.2006.3627
4244:Cruciani et al. (2004)
1478:Regueiro et al. (2006)
1472:Martinez et al. (2007)
1392:Cruciani et al. (2004)
1359:(11.11% of 18 people).
1315:Cruciani et al. (2004)
1242:Martinez et al. (2007)
1149:Cruciani et al. (2004)
443:Trombetta et al. 2015
345:Abu-amero et al. 2009
191:Lazaridis et al. 2016
168:Region and Population
7521:Sykes, Bryan (2006),
6988:10.1093/molbev/msi185
6880:10.1136/jmg.40.11.815
6434:10.1353/hia.2005.0013
6111:10.1353/hub.2006.0045
5809:10.1038/ejhg.2008.120
5779:10.1093/molbev/msm049
5518:10.1038/ejhg.2008.258
5243:Bird, Steven (2007),
5099:10.1038/ejhg.2008.249
4341:, yfull, 3 March 2019
3272:Haplogroup DE (Y-DNA)
3118:Research publications
1484:Zalloua et al. (2008)
1440:Cadenas et al. (2007)
1287:Sanchez et al. (2005)
1271:Republic of Macedonia
1224:Cadenas et al. (2007)
1135:Subclade distribution
1128:Cruciani et al. 2004
1100:Cruciani et al. 2004
1086:Cruciani et al. 2004
1072:Cruciani et al. 2004
1058:Cruciani et al. 2004
1044:Cruciani et al. 2004
1030:Cruciani et al. 2004
1016:Cruciani et al. 2004
1002:Cruciani et al. 2004
988:Cruciani et al. 2004
974:Cruciani et al. 2004
960:Cruciani et al. 2004
946:Cruciani et al. 2004
932:Cruciani et al. 2004
918:Cruciani et al. 2004
890:Cruciani et al. 2004
876:Cruciani et al. 2004
862:Cruciani et al. 2004
848:Cruciani et al. 2004
834:Cruciani et al. 2004
820:Cruciani et al. 2004
806:Cruciani et al. 2004
792:Cruciani et al. 2004
778:Cruciani et al. 2004
764:Cruciani et al. 2004
750:Cruciani et al. 2004
736:Cruciani et al. 2004
722:Cruciani et al. 2004
694:Cruciani et al. 2004
680:Cruciani et al. 2004
666:Cruciani et al. 2004
652:Cruciani et al. 2004
638:Cruciani et al. 2004
624:Cruciani et al. 2004
610:Cruciani et al. 2004
596:Cruciani et al. 2004
582:Cruciani et al. 2004
554:Cruciani et al. 2004
275:Cruciani et al. 2004
219:Cruciani et al. 2004
196:Jordanians (Dead sea)
101:, and defined by the
6943:10.1038/ejhg.2010.21
4456:(5): 1158–1175.e28.
4386:(5): 1158–1175.e28.
3267:Haplogroup D (Y-DNA)
1735:YCC 2010r (Longhand)
1662:Phylogenetic history
1458:Arredi et al. (2004)
1426:Flores et al. (2005)
1405:Flores et al. (2004)
1351:(10% of 30 people),
1293:Semino et al. (2004)
1281:Beleza et al. (2006)
1203:Arredi et al. (2004)
1189:Flores et al. (2005)
1172:Flores et al. (2005)
797:Southwestern Turkish
708:Herrera et al. 2011
685:Southeastern Turkish
657:United Arab Emirates
568:Cadenas et al. 2007
526:Zalloua et al. 2008
364:Assyrian Tehran Iran
132:Cruciani et al. 2004
7599:2011PLoSO...616073T
7553:(1601): 2651–2657.
7487:2008PLoSO...3.2105S
7294:2000Sci...290.1155S
7288:(5494): 1155–1159.
7125:2007BMCEE...7..124R
6762:2008BMCEE...8..213M
6607:(1), archived from
6558:2011PNAS..10818255L
6270:2008PNAS..10510693H
6264:(31): 10693–10698.
6062:2009BMCEE...9..181F
5597:2008PLoSO...3.1430C
5436:2003CBio...13..979C
5255:(2), archived from
4848:2000Sci...290.1155S
4572:2018Natur.557..369D
3973:Y-DNA by population
3888:
3426:Y-DNA backbone tree
1730:YCC 2008 (Longhand)
1725:YCC 2005 (Longhand)
1720:YCC 2002 (Longhand)
1612:, lived during the
1370:Contu et al. (2008)
1363:Adams et al. (2008)
1263:Bosch et al. (2006)
923:Southern Portuguese
909:Northern Portuguese
540:Grugni et al. 2012
512:Hammer et al. 2009
492:Semino et al. 2004
429:Arredi et al. 2004
406:Troina Sicily Italy
373:Grugni et al. 2012
359:Bekada et al. 2013
331:Beleza et al. 2006
317:Floris et al. 2005
303:Semino et al. 2004
289:Grugni et al. 2012
205:Flores et al. 2005
7449:10.1002/ajhb.20567
7412:10.1002/humu.20077
6529:10.1002/ajpa.21078
6402:, John Benjamins,
6390:John D. Bengtson;
6365:10.1101/gr.7172008
6237:10.1002/ajpa.20876
6033:10.1002/ajpa.10265
5020:10.1002/ajhb.20888
4952:Additional sources
4720:Trends in Genetics
4516:(6457): eaat7487.
4013:10.1002/humu.22468
3739:
3520:Y-chromosomal Adam
3190:Phylogenetic trees
1518:Shen et al. (2004)
1495:Subclades of E-M34
1380:Shen et al. (2004)
1311:Luis et al. (2004)
1248:Shen et al. (2004)
951:Southern Spaniards
671:Northern Egyptians
434:Northern Egyptians
160:Semino et al. 2004
74:Defining mutations
7845:10.1101/gr.217602
7720:978-1-902937-20-5
7534:978-0-593-05652-3
7048:10.1159/000093774
6981:(10): 1964–1975.
6677:(Pt 4): 329–339,
6422:History in Africa
6409:978-90-272-3252-6
5736:10.1002/humu.9445
4566:(7705): 369–374.
4473:20.500.12154/1254
4403:20.500.12154/1254
4184:
4183:
4146:, 2017, "K-M2335"
4144:YFull YTree v5.08
4094:Haplogroup T-M184
3962:
3961:
3907:
3897:
3887:
3878:
3866:
3855:
3847:
3835:
3827:
3820:
3813:
3806:
3751:
3670:
3662:
3653:
3583:
3567:
3550:
3533:
3505:
3504:
3419:Haplogroup E-Z827
3414:Haplogroup E-Z820
3374:Haplogroup E-P177
3369:Haplogroup E-P147
3354:Haplogroup E-M521
3344:Haplogroup E-M215
3339:Haplogroup E-M180
3335:Haplogroup E-M123
3331:Haplogroup E-L485
3324:Y-DNA E subclades
3262:Genetic genealogy
3115:
3114:
1614:Middle Bronze Age
1331:(24 people), and
1132:
1131:
853:Northern Italians
811:Southern Italians
769:Central Anatolian
741:Marrakesh Berbers
545:Ethiopian Wolayta
503:
478:Shen et al. 2004
350:Sahara/Mauritania
247:Shen et al. 2004
81:
80:
41:15,000-20,000 ybp
33:Haplogroup E-M123
16:(Redirected from
8013:
7990:
7981:
7952:
7943:
7934:
7904:
7895:
7865:
7856:
7831:
7821:
7788:
7778:
7769:(7): 1008–1021.
7753:
7724:
7705:
7687:
7662:
7629:
7620:
7610:
7580:
7570:
7537:
7517:
7508:
7498:
7468:
7431:
7393:
7384:
7356:
7346:
7313:
7275:
7257:
7242:Human Immunology
7235:
7225:
7200:
7190:
7173:(6): 1526–1543.
7156:
7146:
7136:
7103:
7066:
7030:
7021:
7000:
6990:
6964:
6954:
6921:
6900:
6891:
6862:
6832:
6823:
6806:(5): 1095–1112,
6792:
6783:
6773:
6743:
6734:
6712:
6686:
6664:
6654:
6621:
6620:
6619:
6613:
6598:
6588:
6579:
6569:
6539:
6511:
6483:
6445:
6412:
6386:
6376:
6343:
6315:
6301:
6291:
6281:
6248:
6218:
6209:
6192:(4): 1495–1509,
6178:
6160:
6130:
6092:
6083:
6073:
6043:
6014:
6005:
5983:
5973:
5948:
5939:
5909:
5899:
5865:
5829:
5820:
5791:
5781:
5772:(6): 1300–1311.
5755:
5738:
5716:
5706:
5689:(5): 1014–1022.
5672:
5662:
5645:(5): 1197–1214.
5628:
5618:
5608:
5575:
5538:
5529:
5499:
5463:
5421:
5410:
5401:
5380:
5342:
5333:
5316:(4): 1019–1029,
5303:
5301:
5291:
5266:
5265:
5264:
5239:
5201:
5163:
5153:
5119:
5110:
5080:
5071:
5039:
5022:
5000:
4990:
4946:
4909:
4899:
4882:(6): 1718–1724.
4866:
4842:(5494): 1155–9,
4830:
4820:
4810:
4785:
4767:
4742:
4714:
4704:
4695:(7): 1189–1203.
4679:
4669:
4659:
4624:
4623:
4616:
4610:
4609:
4591:
4550:
4544:
4543:
4533:
4500:
4494:
4493:
4475:
4465:
4440:
4434:
4430:
4424:
4423:
4405:
4395:
4370:
4364:
4363:
4356:
4350:
4349:
4348:
4346:
4333:
4327:
4326:
4325:
4313:
4307:
4302:
4296:
4295:
4285:
4253:
4247:
4241:
4235:
4234:
4227:
4210:
4203:
4174:
4171:
4165:
4162:
4156:
4134:
4128:
4125:
4119:
4116:
4110:
4107:
4101:
4090:
4084:
4081:
4075:
4072:
4066:
4063:
4057:
4054:
4048:
4039:
4033:
4032:
3996:
3903:
3893:
3881:
3873:
3861:
3851:
3842:
3831:
3823:
3816:
3809:
3802:
3747:
3666:
3657:
3649:
3579:
3563:
3546:
3529:
3513:
3500:
3497:
3491:
3479:
3478:
3471:
3454:
3447:
3440:
3431:
3409:Haplogroup E-V68
3404:Haplogroup E-V65
3394:Haplogroup E-V22
3389:Haplogroup E-V13
3384:Haplogroup E-V12
3364:Haplogroup E-M96
3359:Haplogroup E-M75
3349:Haplogroup E-M33
1676:
1343:(28 people) and
1035:Central Italians
1007:Orkney Islanders
825:Turkish Cypriots
783:Istanbul Turkish
643:Sephardi Turkish
501:
483:Sephardim Jewish
420:Algerian Kabyles
294:Italian Calabria
224:Maronite Cypriot
210:Ethiopian Amhara
165:
145:Semitic language
30:
21:
8021:
8020:
8016:
8015:
8014:
8012:
8011:
8010:
7996:
7995:
7994:
7993:
7955:
7947:Zerjal (1999),
7946:
7907:
7868:
7837:Genome Research
7834:
7791:
7756:
7727:
7721:
7708:
7665:
7632:
7583:
7540:
7535:
7520:
7471:
7434:
7396:
7359:
7316:
7278:
7255:10.1.1.408.4857
7238:
7203:
7159:
7106:
7069:
7033:
7003:
6967:
6924:
6903:
6874:(11): 815–819,
6865:
6835:
6795:
6750:BMC Evol. Biol.
6746:
6715:
6684:10.1.1.584.4253
6667:
6624:
6617:
6615:
6611:
6596:
6591:
6552:(45): 18255–9,
6542:
6514:
6495:(293): 707–14,
6486:
6448:
6415:
6410:
6392:Keita, Shomarka
6389:
6353:Genome Research
6346:
6318:
6304:
6251:
6221:
6181:
6133:
6095:
6046:
6017:
5986:
5964:(10): 855–863.
5951:
5912:
5868:
5832:
5794:
5758:
5719:
5675:
5631:
5578:
5541:
5502:
5466:
5430:(11): 979–984.
5424:Current Biology
5419:
5413:
5383:
5345:
5306:
5269:
5262:
5260:
5242:
5204:
5166:
5122:
5083:
5042:
5003:
4959:
4954:
4949:
4915:Nature Genetics
4912:
4869:
4833:
4788:
4745:
4717:
4682:
4637:
4633:
4628:
4627:
4618:
4617:
4613:
4552:
4551:
4547:
4502:
4501:
4497:
4442:
4441:
4437:
4431:
4427:
4372:
4371:
4367:
4358:
4357:
4353:
4344:
4342:
4335:
4334:
4330:
4315:
4314:
4310:
4303:
4299:
4255:
4254:
4250:
4242:
4238:
4229:
4228:
4224:
4219:
4214:
4213:
4204:
4200:
4195:
4190:
4185:
4180:
4179:
4178:
4177:
4172:
4168:
4163:
4159:
4135:
4131:
4126:
4122:
4117:
4113:
4108:
4104:
4091:
4087:
4082:
4078:
4073:
4069:
4064:
4060:
4055:
4051:
4040:
4036:
3998:
3997:
3993:
3988:
3982:
3963:
3506:
3501:
3495:
3492:
3489:
3480:
3476:
3464:
3458:
3428:
3423:
3399:Haplogroup E-M2
3379:Haplogroup E-P2
3326:
3321:
3252:
3247:
3192:
3187:
3135:Kaladjieva 2001
3120:
1670:
1664:
1659:
1606:Nerkin Getashen
1542:
1497:
1450:(1.4%) and the
1357:Mazaro de Vallo
1353:Piazza Armerina
1142:
1137:
965:Spanish Basques
587:Erzurum Turkish
573:Ethiopian Oromo
500:
140:
127:
28:
23:
22:
15:
12:
11:
5:
8019:
8017:
8009:
8008:
7998:
7997:
7992:
7991:
7953:
7944:
7917:(5): 633–642,
7905:
7878:(4): 873–882,
7866:
7832:
7804:(1): 229–234.
7789:
7754:
7736:(1): 539–564.
7725:
7719:
7706:
7663:
7630:
7581:
7538:
7533:
7518:
7469:
7443:(6): 829–837.
7432:
7406:(3): 248–260.
7400:Human Mutation
7394:
7373:10.1086/386295
7357:
7335:10.1086/338306
7329:(1): 265–268.
7314:
7276:
7248:(9): 871–884.
7236:
7216:(7): 856–866.
7201:
7179:10.1086/316890
7157:
7104:
7067:
7042:(3): 132–143,
7031:
7001:
6965:
6937:(8): 915–923.
6922:
6913:(1): 242–243,
6901:
6863:
6833:
6812:10.1086/324070
6793:
6744:
6713:
6665:
6643:10.1086/382286
6637:(3): 532–544.
6622:
6589:
6540:
6523:(2): 336–346,
6512:
6484:
6458:(2): 205–214.
6446:
6413:
6408:
6387:
6359:(5): 830–838.
6344:
6327:(8): 356–362,
6316:
6302:
6249:
6231:(3): 316–323.
6219:
6179:
6143:(4): 443–454.
6131:
6105:(3): 307–316.
6093:
6044:
6027:(3): 270–279,
6015:
5996:(9): 435–441,
5984:
5949:
5922:(1): 121–126,
5910:
5882:(6): 568–581.
5866:
5841:(5702): 1680,
5830:
5792:
5756:
5723:Human Mutation
5717:
5695:10.1086/386294
5673:
5651:10.1086/340257
5629:
5576:
5539:
5512:(6): 848–852,
5500:
5475:(4): 357–360,
5464:
5411:
5381:
5355:(4): 459–487.
5343:
5322:10.1086/319521
5310:Am J Hum Genet
5304:
5267:
5240:
5214:(2): 181–194.
5202:
5176:(4): 354–365.
5170:Human Genetics
5168:populations".
5164:
5142:10.1086/378506
5136:(4): 768–779.
5120:
5093:(6): 820–830,
5081:
5060:10.1086/423147
5054:(2): 338–345,
5040:
5013:(3): 407–409,
5001:
4973:(6): 725–736.
4956:
4955:
4953:
4950:
4948:
4947:
4921:(3): 358–361.
4910:
4888:10.1086/302680
4867:
4831:
4808:10.1086/323299
4801:(3): 615–628.
4786:
4743:
4715:
4680:
4657:10.1086/318205
4650:(2): 432–443.
4634:
4632:
4629:
4626:
4625:
4611:
4545:
4495:
4435:
4425:
4365:
4351:
4328:
4323:10.1101/059311
4308:
4297:
4268:(5): 707–717.
4262:Human Genetics
4248:
4236:
4231:"E-M123 YTree"
4221:
4220:
4218:
4215:
4212:
4211:
4197:
4196:
4194:
4191:
4189:
4186:
4182:
4181:
4176:
4175:
4166:
4157:
4129:
4120:
4111:
4102:
4085:
4076:
4067:
4058:
4049:
4034:
4001:Human Mutation
3990:
3989:
3984:
3983:
3981:
3980:
3975:
3969:
3968:
3965:
3964:
3960:
3959:
3957:
3952:
3947:
3944:
3943:
3938:
3936:
3931:
3929:
3924:
3919:
3914:
3911:
3910:
3908:
3901:
3899:
3891:
3889:
3879:
3871:
3868:
3867:
3859:
3857:
3848:
3840:
3837:
3836:
3829:
3821:
3814:
3807:
3800:
3798:
3791:
3785:
3783:
3777:
3771:
3769:
3763:
3757:
3754:
3753:
3744:
3737:
3735:
3729:
3723:
3720:
3719:
3714:
3709:
3706:
3705:
3700:
3695:
3692:
3691:
3686:
3681:
3678:
3677:
3671:
3663:
3654:
3647:
3644:
3643:
3638:
3633:
3631:
3626:
3621:
3618:
3617:
3612:
3607:
3604:
3603:
3598:
3593:
3590:
3589:
3584:
3577:
3574:
3573:
3568:
3561:
3558:
3557:
3551:
3544:
3541:
3540:
3534:
3527:
3524:
3523:
3516:
3511:
3508:
3507:
3503:
3502:
3483:
3481:
3474:
3469:
3466:
3465:
3459:
3457:
3456:
3449:
3442:
3434:
3427:
3424:
3422:
3421:
3416:
3411:
3406:
3401:
3396:
3391:
3386:
3381:
3376:
3371:
3366:
3361:
3356:
3351:
3346:
3341:
3336:
3333:
3327:
3325:
3322:
3320:
3319:
3314:
3309:
3304:
3299:
3294:
3289:
3284:
3279:
3274:
3269:
3264:
3259:
3253:
3251:
3248:
3246:
3243:
3242:
3241:
3240:
3239:
3238:
3237:
3231:
3225:
3219:
3218:
3217:
3191:
3188:
3186:
3185:
3177:
3169:
3161:
3153:
3145:
3143:Underhill 2000
3137:
3124:
3119:
3116:
3113:
3112:
3109:
3106:
3103:
3100:
3097:
3094:
3091:
3088:
3085:
3082:
3079:
3076:
3073:
3070:
3067:
3064:
3061:
3058:
3052:
3051:
3048:
3045:
3042:
3039:
3036:
3033:
3030:
3027:
3024:
3021:
3018:
3015:
3012:
3009:
3006:
3003:
3000:
2997:
2991:
2990:
2987:
2984:
2981:
2978:
2975:
2972:
2969:
2966:
2963:
2960:
2957:
2954:
2951:
2948:
2945:
2942:
2939:
2936:
2932:
2931:
2928:
2925:
2922:
2919:
2916:
2913:
2910:
2907:
2904:
2901:
2898:
2895:
2892:
2889:
2886:
2883:
2880:
2877:
2871:
2870:
2867:
2864:
2861:
2858:
2855:
2852:
2849:
2846:
2843:
2840:
2837:
2834:
2831:
2828:
2825:
2822:
2819:
2816:
2810:
2809:
2806:
2803:
2800:
2797:
2794:
2791:
2788:
2785:
2782:
2779:
2776:
2773:
2770:
2767:
2764:
2761:
2758:
2755:
2749:
2748:
2745:
2742:
2739:
2736:
2733:
2730:
2727:
2724:
2721:
2718:
2715:
2712:
2709:
2706:
2703:
2700:
2697:
2694:
2688:
2687:
2684:
2681:
2678:
2675:
2672:
2669:
2666:
2663:
2660:
2657:
2654:
2651:
2648:
2645:
2642:
2639:
2636:
2633:
2627:
2626:
2623:
2620:
2617:
2614:
2611:
2608:
2605:
2602:
2599:
2596:
2593:
2590:
2587:
2584:
2581:
2578:
2575:
2572:
2566:
2565:
2562:
2559:
2556:
2553:
2550:
2547:
2544:
2541:
2538:
2535:
2532:
2529:
2526:
2523:
2520:
2517:
2514:
2511:
2505:
2504:
2501:
2498:
2495:
2492:
2489:
2486:
2483:
2480:
2477:
2474:
2471:
2468:
2465:
2462:
2459:
2456:
2453:
2450:
2444:
2443:
2440:
2437:
2434:
2431:
2428:
2425:
2422:
2419:
2416:
2413:
2410:
2407:
2404:
2401:
2398:
2395:
2392:
2389:
2383:
2382:
2379:
2376:
2373:
2370:
2367:
2364:
2361:
2358:
2355:
2352:
2349:
2346:
2343:
2340:
2337:
2334:
2331:
2328:
2322:
2321:
2318:
2315:
2312:
2309:
2306:
2303:
2300:
2297:
2294:
2291:
2288:
2285:
2282:
2279:
2276:
2273:
2270:
2267:
2261:
2260:
2257:
2254:
2251:
2248:
2245:
2242:
2239:
2236:
2233:
2230:
2227:
2224:
2221:
2218:
2215:
2212:
2209:
2206:
2200:
2199:
2196:
2193:
2190:
2187:
2184:
2181:
2178:
2175:
2172:
2169:
2166:
2163:
2160:
2157:
2154:
2151:
2148:
2145:
2139:
2138:
2135:
2132:
2129:
2126:
2123:
2120:
2117:
2114:
2111:
2108:
2105:
2102:
2099:
2096:
2093:
2090:
2087:
2084:
2078:
2077:
2074:
2071:
2068:
2065:
2062:
2059:
2056:
2053:
2050:
2047:
2044:
2041:
2038:
2035:
2032:
2029:
2026:
2023:
2017:
2016:
2013:
2010:
2007:
2004:
2001:
1998:
1995:
1992:
1989:
1986:
1983:
1980:
1977:
1974:
1971:
1968:
1965:
1962:
1956:
1955:
1952:
1949:
1946:
1943:
1940:
1937:
1934:
1931:
1928:
1925:
1922:
1919:
1916:
1913:
1910:
1907:
1904:
1901:
1895:
1894:
1891:
1888:
1885:
1882:
1879:
1876:
1873:
1870:
1867:
1864:
1861:
1858:
1855:
1852:
1849:
1846:
1843:
1840:
1834:
1833:
1830:
1827:
1824:
1821:
1818:
1815:
1812:
1809:
1806:
1803:
1800:
1797:
1794:
1791:
1788:
1785:
1782:
1779:
1773:
1772:
1767:
1762:
1757:
1752:
1747:
1742:
1737:
1732:
1727:
1722:
1717:
1712:
1707:
1702:
1697:
1692:
1687:
1682:
1666:Main article:
1663:
1660:
1658:
1655:
1654:
1653:
1652:under E-M123*.
1648:
1647:
1643:
1642:
1618:
1617:
1601:
1600:
1580:
1579:
1561:
1560:
1541:
1538:
1537:
1536:
1525:
1511:
1496:
1493:
1492:
1491:
1481:
1475:
1469:
1455:
1437:
1422:
1412:
1402:
1395:
1384:
1383:
1377:
1367:
1360:
1318:
1308:
1290:
1284:
1278:
1256:
1255:
1254:
1253:
1252:
1251:
1245:
1239:
1221:
1210:
1209:
1206:
1200:
1186:
1175:
1164:
1141:
1138:
1136:
1133:
1130:
1129:
1126:
1123:
1120:
1116:
1115:
1112:
1109:
1106:
1102:
1101:
1098:
1095:
1092:
1088:
1087:
1084:
1081:
1078:
1074:
1073:
1070:
1067:
1064:
1060:
1059:
1056:
1053:
1050:
1046:
1045:
1042:
1039:
1036:
1032:
1031:
1028:
1025:
1022:
1018:
1017:
1014:
1011:
1008:
1004:
1003:
1000:
997:
994:
993:French Basques
990:
989:
986:
983:
980:
976:
975:
972:
969:
966:
962:
961:
958:
955:
952:
948:
947:
944:
941:
938:
934:
933:
930:
927:
924:
920:
919:
916:
913:
910:
906:
905:
902:
899:
896:
892:
891:
888:
885:
882:
878:
877:
874:
871:
868:
864:
863:
860:
857:
854:
850:
849:
846:
843:
840:
836:
835:
832:
829:
826:
822:
821:
818:
815:
812:
808:
807:
804:
801:
798:
794:
793:
790:
787:
784:
780:
779:
776:
773:
770:
766:
765:
762:
759:
756:
752:
751:
748:
745:
742:
738:
737:
734:
731:
728:
724:
723:
720:
717:
714:
710:
709:
706:
703:
700:
696:
695:
692:
689:
686:
682:
681:
678:
675:
672:
668:
667:
664:
661:
658:
654:
653:
650:
647:
644:
640:
639:
636:
633:
630:
626:
625:
622:
619:
616:
612:
611:
608:
605:
602:
598:
597:
594:
591:
588:
584:
583:
580:
577:
574:
570:
569:
566:
563:
560:
556:
555:
552:
549:
546:
542:
541:
538:
535:
532:
528:
527:
524:
521:
518:
514:
513:
510:
507:
504:
498:Ashkenazi Jews
494:
493:
490:
487:
484:
480:
479:
476:
473:
470:
466:
465:
462:
459:
456:
445:
444:
441:
438:
435:
431:
430:
427:
424:
421:
417:
416:
413:
410:
407:
403:
402:
399:
396:
393:
389:
388:
385:
382:
379:
375:
374:
371:
368:
365:
361:
360:
357:
354:
351:
347:
346:
343:
340:
337:
333:
332:
329:
326:
323:
319:
318:
315:
312:
309:
305:
304:
301:
298:
295:
291:
290:
287:
284:
281:
277:
276:
273:
270:
267:
266:Ethiopian Jews
263:
262:
259:
256:
253:
249:
248:
245:
242:
239:
235:
234:
231:
228:
225:
221:
220:
217:
214:
211:
207:
206:
203:
200:
197:
193:
192:
189:
186:
183:
179:
178:
175:
172:
169:
139:
136:
126:
123:
85:human genetics
79:
78:
75:
71:
70:
67:
63:
62:
57:
53:
52:
47:
43:
42:
39:
35:
34:
26:
24:
14:
13:
10:
9:
6:
4:
3:
2:
8018:
8007:
8004:
8003:
8001:
7989:
7985:
7980:
7975:
7971:
7967:
7963:
7959:
7954:
7950:
7945:
7942:
7938:
7933:
7928:
7924:
7920:
7916:
7912:
7906:
7903:
7899:
7894:
7889:
7885:
7881:
7877:
7873:
7867:
7864:
7860:
7855:
7850:
7846:
7842:
7838:
7833:
7829:
7825:
7820:
7815:
7811:
7807:
7803:
7799:
7795:
7790:
7786:
7782:
7777:
7772:
7768:
7764:
7760:
7755:
7751:
7747:
7743:
7739:
7735:
7731:
7726:
7722:
7716:
7712:
7707:
7703:
7699:
7695:
7691:
7686:
7681:
7677:
7673:
7669:
7664:
7661:
7657:
7653:
7649:
7645:
7644:10.1038/81685
7641:
7637:
7631:
7628:
7624:
7619:
7614:
7609:
7604:
7600:
7596:
7593:(1): e16073,
7592:
7588:
7582:
7578:
7574:
7569:
7564:
7560:
7556:
7552:
7548:
7544:
7539:
7536:
7530:
7526:
7525:
7519:
7516:
7512:
7507:
7502:
7497:
7492:
7488:
7484:
7480:
7476:
7470:
7466:
7462:
7458:
7454:
7450:
7446:
7442:
7438:
7433:
7429:
7425:
7421:
7417:
7413:
7409:
7405:
7401:
7395:
7392:
7388:
7383:
7378:
7374:
7370:
7366:
7365:
7358:
7354:
7350:
7345:
7340:
7336:
7332:
7328:
7324:
7320:
7315:
7311:
7307:
7303:
7299:
7295:
7291:
7287:
7283:
7277:
7273:
7269:
7265:
7261:
7256:
7251:
7247:
7243:
7237:
7233:
7229:
7224:
7219:
7215:
7211:
7207:
7202:
7198:
7194:
7189:
7184:
7180:
7176:
7172:
7168:
7164:
7158:
7154:
7150:
7145:
7140:
7135:
7130:
7126:
7122:
7118:
7114:
7110:
7105:
7102:
7098:
7094:
7090:
7086:
7082:
7078:
7074:
7068:
7065:
7061:
7057:
7053:
7049:
7045:
7041:
7037:
7032:
7029:
7025:
7020:
7015:
7011:
7007:
7002:
6998:
6994:
6989:
6984:
6980:
6976:
6972:
6966:
6962:
6958:
6953:
6948:
6944:
6940:
6936:
6932:
6928:
6923:
6920:
6916:
6912:
6908:
6902:
6899:
6895:
6890:
6885:
6881:
6877:
6873:
6869:
6864:
6860:
6856:
6852:
6848:
6844:
6840:
6834:
6831:
6827:
6822:
6817:
6813:
6809:
6805:
6801:
6800:
6794:
6791:
6787:
6782:
6777:
6772:
6767:
6763:
6759:
6755:
6751:
6745:
6742:
6738:
6733:
6728:
6725:(4): 485–93,
6724:
6720:
6714:
6710:
6706:
6702:
6698:
6694:
6690:
6685:
6680:
6676:
6672:
6666:
6662:
6658:
6653:
6648:
6644:
6640:
6636:
6632:
6628:
6623:
6614:on 2016-05-06
6610:
6606:
6602:
6595:
6590:
6587:
6583:
6578:
6573:
6568:
6563:
6559:
6555:
6551:
6547:
6541:
6538:
6534:
6530:
6526:
6522:
6518:
6513:
6510:
6506:
6502:
6498:
6494:
6490:
6485:
6481:
6477:
6473:
6469:
6465:
6461:
6457:
6453:
6447:
6443:
6439:
6435:
6431:
6427:
6423:
6419:
6414:
6411:
6405:
6401:
6397:
6393:
6388:
6384:
6380:
6375:
6370:
6366:
6362:
6358:
6354:
6350:
6345:
6342:
6338:
6334:
6330:
6326:
6322:
6321:Trends Genet.
6317:
6313:
6312:
6307:
6303:
6299:
6295:
6290:
6285:
6280:
6275:
6271:
6267:
6263:
6259:
6255:
6250:
6246:
6242:
6238:
6234:
6230:
6226:
6220:
6217:
6213:
6208:
6203:
6199:
6195:
6191:
6187:
6186:
6180:
6176:
6172:
6168:
6164:
6159:
6158:10400.13/3018
6154:
6150:
6146:
6142:
6138:
6132:
6128:
6124:
6120:
6116:
6112:
6108:
6104:
6100:
6099:Human Biology
6094:
6091:
6087:
6082:
6077:
6072:
6067:
6063:
6059:
6055:
6051:
6045:
6042:
6038:
6034:
6030:
6026:
6022:
6016:
6013:
6009:
6004:
5999:
5995:
5991:
5985:
5981:
5977:
5972:
5967:
5963:
5959:
5955:
5950:
5947:
5943:
5938:
5933:
5929:
5925:
5921:
5917:
5911:
5907:
5903:
5898:
5893:
5889:
5885:
5881:
5877:
5873:
5867:
5864:
5860:
5856:
5852:
5848:
5844:
5840:
5836:
5831:
5828:
5824:
5819:
5814:
5810:
5806:
5802:
5798:
5793:
5789:
5785:
5780:
5775:
5771:
5767:
5763:
5757:
5754:
5750:
5746:
5742:
5737:
5732:
5728:
5724:
5718:
5714:
5710:
5705:
5700:
5696:
5692:
5688:
5684:
5680:
5674:
5670:
5666:
5661:
5656:
5652:
5648:
5644:
5640:
5636:
5630:
5626:
5622:
5617:
5612:
5607:
5602:
5598:
5594:
5590:
5586:
5582:
5577:
5574:
5570:
5566:
5562:
5558:
5554:
5551:(2): 127–48,
5550:
5546:
5540:
5537:
5533:
5528:
5523:
5519:
5515:
5511:
5507:
5501:
5498:
5494:
5490:
5486:
5482:
5478:
5474:
5470:
5465:
5461:
5457:
5453:
5449:
5445:
5441:
5437:
5433:
5429:
5425:
5418:
5412:
5409:
5405:
5400:
5395:
5391:
5387:
5382:
5378:
5374:
5370:
5366:
5362:
5358:
5354:
5350:
5344:
5341:
5337:
5332:
5327:
5323:
5319:
5315:
5311:
5305:
5300:
5295:
5290:
5285:
5281:
5277:
5273:
5268:
5259:on 2016-04-22
5258:
5254:
5250:
5246:
5241:
5237:
5233:
5229:
5225:
5221:
5217:
5213:
5209:
5203:
5199:
5195:
5191:
5187:
5183:
5179:
5175:
5171:
5165:
5161:
5157:
5152:
5147:
5143:
5139:
5135:
5131:
5127:
5121:
5118:
5114:
5109:
5104:
5100:
5096:
5092:
5088:
5082:
5079:
5075:
5070:
5065:
5061:
5057:
5053:
5049:
5048:
5041:
5038:
5034:
5030:
5026:
5021:
5016:
5012:
5008:
5002:
4998:
4994:
4989:
4984:
4980:
4976:
4972:
4968:
4964:
4958:
4957:
4951:
4944:
4940:
4936:
4932:
4928:
4927:10.1038/81685
4924:
4920:
4916:
4911:
4907:
4903:
4898:
4893:
4889:
4885:
4881:
4877:
4873:
4868:
4865:
4861:
4857:
4853:
4849:
4845:
4841:
4837:
4832:
4828:
4824:
4819:
4814:
4809:
4804:
4800:
4796:
4792:
4787:
4783:
4779:
4775:
4771:
4766:
4761:
4758:(2): 97–104.
4757:
4753:
4749:
4744:
4741:
4737:
4733:
4729:
4726:(8): 356–62,
4725:
4721:
4716:
4712:
4708:
4703:
4698:
4694:
4690:
4686:
4681:
4677:
4673:
4668:
4663:
4658:
4653:
4649:
4645:
4641:
4636:
4635:
4630:
4621:
4615:
4612:
4607:
4603:
4599:
4595:
4590:
4585:
4581:
4577:
4573:
4569:
4565:
4561:
4557:
4549:
4546:
4541:
4537:
4532:
4527:
4523:
4519:
4515:
4511:
4507:
4499:
4496:
4491:
4487:
4483:
4479:
4474:
4469:
4464:
4459:
4455:
4451:
4447:
4439:
4436:
4429:
4426:
4421:
4417:
4413:
4409:
4404:
4399:
4394:
4389:
4385:
4381:
4377:
4369:
4366:
4361:
4360:"E-M84 YTree"
4355:
4352:
4340:
4339:
4332:
4329:
4324:
4319:
4312:
4309:
4306:
4301:
4298:
4293:
4289:
4284:
4279:
4275:
4271:
4267:
4263:
4259:
4252:
4249:
4245:
4240:
4237:
4232:
4226:
4223:
4216:
4208:
4202:
4199:
4192:
4187:
4170:
4167:
4161:
4158:
4154:
4152:
4147:
4145:
4140:
4133:
4130:
4124:
4121:
4115:
4112:
4106:
4103:
4099:
4095:
4089:
4086:
4080:
4077:
4071:
4068:
4062:
4059:
4053:
4050:
4046:
4045:
4038:
4035:
4030:
4026:
4022:
4018:
4014:
4010:
4007:(2): 187–91.
4006:
4002:
3995:
3992:
3987:
3979:
3976:
3974:
3971:
3970:
3966:
3958:
3956:
3951:
3945:
3942:
3935:
3928:
3923:
3918:
3912:
3906:
3900:
3898:
3896:
3885:
3876:
3869:
3864:
3854:
3845:
3838:
3834:
3828:
3826:
3819:
3812:
3805:
3796:
3790:
3789:
3781:
3776:
3775:
3767:
3762:
3761:
3755:
3750:
3742:
3733:
3728:
3727:
3721:
3718:
3713:
3707:
3704:
3699:
3693:
3690:
3685:
3679:
3676:
3669:
3660:
3652:
3645:
3642:
3637:
3630:
3625:
3619:
3616:
3611:
3605:
3602:
3597:
3591:
3588:
3582:
3575:
3572:
3566:
3559:
3555:
3549:
3542:
3538:
3532:
3525:
3521:
3514:
3509:
3499:
3496:February 2021
3487:
3482:
3473:
3472:
3467:
3463:
3455:
3450:
3448:
3443:
3441:
3436:
3435:
3432:
3425:
3420:
3417:
3415:
3412:
3410:
3407:
3405:
3402:
3400:
3397:
3395:
3392:
3390:
3387:
3385:
3382:
3380:
3377:
3375:
3372:
3370:
3367:
3365:
3362:
3360:
3357:
3355:
3352:
3350:
3347:
3345:
3342:
3340:
3337:
3334:
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3329:
3328:
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3318:
3315:
3313:
3310:
3308:
3305:
3303:
3300:
3298:
3295:
3293:
3290:
3288:
3285:
3283:
3280:
3278:
3275:
3273:
3270:
3268:
3265:
3263:
3260:
3258:
3255:
3254:
3249:
3244:
3235:
3232:
3229:
3226:
3223:
3220:
3215:
3212:
3211:
3209:
3206:
3205:
3203:
3200:
3199:
3197:
3194:
3193:
3189:
3184:
3181:
3178:
3176:
3173:
3170:
3168:
3165:
3162:
3160:
3157:
3154:
3152:
3149:
3146:
3144:
3141:
3138:
3136:
3132:
3129:
3126:
3125:
3123:
3117:
3111:E1b1b1b2a1a1
3110:
3107:
3104:
3101:
3098:
3095:
3092:
3089:
3086:
3083:
3080:
3077:
3074:
3071:
3068:
3065:
3062:
3059:
3057:
3054:
3053:
3049:
3046:
3043:
3040:
3037:
3034:
3031:
3028:
3025:
3022:
3019:
3016:
3013:
3010:
3007:
3004:
3001:
2998:
2996:
2993:
2992:
2988:
2985:
2982:
2979:
2976:
2973:
2970:
2967:
2964:
2961:
2958:
2955:
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2949:
2946:
2943:
2940:
2937:
2934:
2933:
2929:
2926:
2923:
2920:
2917:
2914:
2911:
2908:
2905:
2902:
2899:
2896:
2893:
2890:
2887:
2884:
2881:
2878:
2876:
2873:
2872:
2868:
2865:
2862:
2859:
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2853:
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2847:
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2835:
2832:
2829:
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2820:
2817:
2815:
2812:
2811:
2807:
2804:
2801:
2798:
2795:
2792:
2789:
2786:
2783:
2780:
2777:
2774:
2771:
2768:
2765:
2762:
2759:
2756:
2754:
2751:
2750:
2746:
2743:
2740:
2737:
2734:
2731:
2728:
2725:
2722:
2719:
2716:
2713:
2710:
2707:
2704:
2701:
2698:
2695:
2693:
2690:
2689:
2685:
2682:
2679:
2676:
2673:
2670:
2667:
2664:
2661:
2658:
2655:
2652:
2649:
2646:
2643:
2640:
2637:
2634:
2632:
2629:
2628:
2624:
2621:
2618:
2615:
2612:
2609:
2606:
2603:
2600:
2597:
2594:
2591:
2588:
2585:
2582:
2579:
2576:
2573:
2571:
2568:
2567:
2563:
2560:
2557:
2554:
2551:
2548:
2545:
2542:
2539:
2536:
2533:
2530:
2527:
2524:
2521:
2518:
2515:
2512:
2510:
2507:
2506:
2502:
2499:
2496:
2493:
2490:
2487:
2484:
2481:
2478:
2475:
2472:
2469:
2466:
2463:
2460:
2457:
2454:
2451:
2449:
2446:
2445:
2441:
2438:
2435:
2432:
2429:
2426:
2423:
2420:
2417:
2414:
2411:
2408:
2405:
2402:
2399:
2396:
2393:
2390:
2388:
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2384:
2380:
2377:
2374:
2371:
2368:
2365:
2362:
2359:
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2344:
2341:
2338:
2335:
2332:
2329:
2327:
2324:
2323:
2319:
2316:
2313:
2310:
2307:
2304:
2301:
2298:
2295:
2292:
2289:
2286:
2283:
2280:
2277:
2274:
2271:
2268:
2266:
2263:
2262:
2258:
2255:
2252:
2249:
2246:
2243:
2240:
2237:
2234:
2231:
2228:
2225:
2222:
2219:
2216:
2213:
2210:
2207:
2205:
2202:
2201:
2197:
2194:
2191:
2188:
2185:
2182:
2179:
2176:
2173:
2170:
2167:
2164:
2161:
2158:
2155:
2152:
2149:
2146:
2144:
2141:
2140:
2136:
2133:
2130:
2127:
2124:
2121:
2118:
2115:
2112:
2109:
2106:
2103:
2100:
2097:
2094:
2091:
2088:
2085:
2083:
2080:
2079:
2075:
2072:
2069:
2066:
2063:
2060:
2057:
2054:
2051:
2048:
2045:
2042:
2039:
2036:
2033:
2030:
2027:
2024:
2022:
2019:
2018:
2014:
2011:
2008:
2005:
2002:
1999:
1996:
1993:
1990:
1987:
1984:
1981:
1978:
1975:
1972:
1969:
1966:
1963:
1961:
1958:
1957:
1953:
1950:
1947:
1944:
1941:
1938:
1935:
1932:
1929:
1926:
1923:
1920:
1917:
1914:
1911:
1908:
1905:
1902:
1900:
1897:
1896:
1892:
1889:
1886:
1883:
1880:
1877:
1874:
1871:
1868:
1865:
1862:
1859:
1856:
1853:
1850:
1847:
1844:
1841:
1839:
1836:
1835:
1831:
1828:
1825:
1822:
1819:
1816:
1813:
1810:
1807:
1804:
1801:
1798:
1795:
1792:
1789:
1786:
1783:
1780:
1778:
1775:
1774:
1771:
1766:
1761:
1756:
1751:
1746:
1741:
1736:
1731:
1726:
1721:
1716:
1711:
1706:
1701:
1696:
1691:
1686:
1681:
1677:
1674:
1669:
1661:
1657:Phylogenetics
1656:
1650:
1649:
1645:
1644:
1640:
1636:
1632:
1628:
1624:
1620:
1619:
1615:
1611:
1607:
1603:
1602:
1598:
1594:
1590:
1586:
1582:
1581:
1577:
1573:
1569:
1566:
1563:
1562:
1557:
1553:
1548:
1544:
1543:
1539:
1533:
1529:
1526:
1523:
1519:
1515:
1512:
1509:
1506:
1502:
1499:
1498:
1494:
1489:
1485:
1482:
1479:
1476:
1473:
1470:
1467:
1463:
1459:
1456:
1453:
1449:
1445:
1441:
1438:
1435:
1431:
1427:
1423:
1420:
1416:
1413:
1410:
1406:
1403:
1400:
1396:
1393:
1390:According to
1389:
1388:
1387:
1381:
1378:
1375:
1371:
1368:
1364:
1361:
1358:
1354:
1350:
1346:
1342:
1338:
1334:
1330:
1327:(33 people),
1326:
1322:
1319:
1316:
1312:
1309:
1306:
1302:
1298:
1294:
1291:
1288:
1285:
1282:
1279:
1276:
1272:
1268:
1264:
1261:
1260:
1259:
1249:
1246:
1243:
1240:
1237:
1233:
1229:
1225:
1222:
1219:
1216:
1215:
1214:
1213:
1212:
1211:
1207:
1204:
1201:
1198:
1194:
1190:
1187:
1184:
1180:
1176:
1173:
1169:
1165:
1162:
1158:
1154:
1150:
1147:
1146:
1145:
1139:
1134:
1127:
1124:
1121:
1118:
1117:
1113:
1110:
1107:
1104:
1103:
1099:
1096:
1093:
1090:
1089:
1085:
1082:
1079:
1076:
1075:
1071:
1068:
1065:
1062:
1061:
1057:
1054:
1051:
1048:
1047:
1043:
1040:
1037:
1034:
1033:
1029:
1026:
1023:
1020:
1019:
1015:
1012:
1009:
1006:
1005:
1001:
998:
995:
992:
991:
987:
984:
981:
978:
977:
973:
970:
967:
964:
963:
959:
956:
953:
950:
949:
945:
942:
939:
936:
935:
931:
928:
925:
922:
921:
917:
914:
911:
908:
907:
903:
900:
897:
894:
893:
889:
886:
883:
880:
879:
875:
872:
869:
866:
865:
861:
858:
855:
852:
851:
847:
844:
841:
838:
837:
833:
830:
827:
824:
823:
819:
816:
813:
810:
809:
805:
802:
799:
796:
795:
791:
788:
785:
782:
781:
777:
774:
771:
768:
767:
763:
760:
757:
754:
753:
749:
746:
743:
740:
739:
735:
732:
729:
726:
725:
721:
718:
715:
712:
711:
707:
704:
701:
698:
697:
693:
690:
687:
684:
683:
679:
676:
673:
670:
669:
665:
662:
659:
656:
655:
651:
648:
645:
642:
641:
637:
634:
631:
628:
627:
623:
620:
617:
614:
613:
609:
606:
603:
600:
599:
595:
592:
589:
586:
585:
581:
578:
575:
572:
571:
567:
564:
561:
558:
557:
553:
550:
547:
544:
543:
539:
536:
533:
530:
529:
525:
522:
519:
516:
515:
511:
508:
505:
499:
496:
495:
491:
488:
485:
482:
481:
477:
474:
471:
468:
467:
463:
460:
457:
454:
450:
447:
446:
442:
439:
436:
433:
432:
428:
425:
422:
419:
418:
414:
411:
408:
405:
404:
400:
397:
394:
391:
390:
386:
383:
380:
377:
376:
372:
369:
366:
363:
362:
358:
355:
352:
349:
348:
344:
341:
338:
335:
334:
330:
327:
324:
322:Beja Portugal
321:
320:
316:
313:
310:
307:
306:
302:
299:
296:
293:
292:
288:
285:
282:
279:
278:
274:
271:
268:
265:
264:
260:
257:
254:
252:Greek Cypriot
251:
250:
246:
243:
240:
237:
236:
232:
229:
226:
223:
222:
218:
215:
212:
209:
208:
204:
201:
198:
195:
194:
190:
187:
184:
181:
180:
176:
173:
170:
167:
166:
163:
161:
157:
154:
150:
146:
137:
135:
133:
124:
122:
120:
116:
112:
108:
104:
100:
97:
93:
90:
86:
76:
72:
68:
64:
61:
58:
54:
51:
48:
44:
40:
36:
31:
19:
7961:
7957:
7948:
7914:
7910:
7875:
7871:
7836:
7801:
7797:
7766:
7762:
7733:
7729:
7710:
7678:(1): 43–62.
7675:
7671:
7635:
7590:
7586:
7550:
7546:
7523:
7481:(5): e2105,
7478:
7474:
7440:
7436:
7403:
7399:
7362:
7326:
7322:
7285:
7281:
7245:
7241:
7213:
7209:
7170:
7166:
7116:
7112:
7079:(3): 251–5,
7076:
7072:
7039:
7035:
7009:
7005:
6978:
6974:
6934:
6930:
6910:
6906:
6871:
6867:
6845:(1): 23–35.
6842:
6838:
6803:
6797:
6753:
6749:
6722:
6718:
6674:
6670:
6634:
6630:
6616:, retrieved
6609:the original
6604:
6600:
6549:
6545:
6520:
6516:
6492:
6488:
6455:
6451:
6425:
6421:
6399:
6356:
6352:
6324:
6320:
6310:
6261:
6257:
6228:
6224:
6189:
6183:
6140:
6136:
6102:
6098:
6053:
6049:
6024:
6020:
5993:
5989:
5961:
5957:
5919:
5915:
5879:
5875:
5838:
5834:
5803:(1): 91–99,
5800:
5796:
5769:
5765:
5729:(8): 831–2,
5726:
5722:
5686:
5682:
5642:
5638:
5633:(May 2002).
5591:(1): e1430.
5588:
5584:
5548:
5544:
5509:
5505:
5472:
5468:
5427:
5423:
5389:
5385:
5352:
5348:
5313:
5309:
5282:(1): 01–08.
5279:
5275:
5261:, retrieved
5257:the original
5252:
5248:
5211:
5207:
5173:
5169:
5133:
5129:
5090:
5086:
5051:
5045:
5010:
5006:
4970:
4966:
4918:
4914:
4879:
4875:
4839:
4835:
4798:
4794:
4755:
4751:
4723:
4719:
4692:
4688:
4647:
4643:
4614:
4589:1887/3202709
4563:
4559:
4548:
4513:
4509:
4498:
4453:
4449:
4438:
4428:
4383:
4379:
4368:
4354:
4343:, retrieved
4337:
4331:
4311:
4305:ISOGG (2011)
4300:
4265:
4261:
4251:
4239:
4225:
4201:
4169:
4160:
4150:
4143:
4138:
4132:
4123:
4114:
4105:
4088:
4079:
4070:
4061:
4052:
4043:
4037:
4004:
4000:
3994:
3985:
3850:
3793:
3493:
3485:
3233:
3227:
3221:
3213:
3207:
3201:
3195:
3183:Capelli 2001
3179:
3171:
3163:
3159:Karafet 2001
3155:
3147:
3139:
3127:
3121:
2930:E1b1b1b1a2a
2442:E1b1a1a1g1c
1769:
1764:
1759:
1754:
1749:
1744:
1739:
1734:
1729:
1724:
1719:
1714:
1709:
1704:
1699:
1694:
1689:
1684:
1679:
1671:
1616:, two E-M84.
1527:
1513:
1500:
1424:Strikingly,
1385:
1257:
1161:Central Asia
1143:
755:Palestinians
517:Palestinians
141:
138:Distribution
128:
96:Y-chromosome
91:
89:Y Haplogroup
82:
7964:(1): 1–14,
7012:: 439–441,
6868:J Med Genet
6428:: 221–246.
5990:J Hum Genet
5392:(3): 1–13,
5299:10183/23369
4217:Works cited
3236:(L791,L792)
3167:Semino 2000
3151:Hammer 2001
3050:E1b1b1b2a1
2869:E1b1b1b1a1
2747:E1b1b1a1c1
2439:E1b1a1a1g1c
1637:during the
1576:'Ain Ghazal
1540:Ancient DNA
1522:Palestinian
1397:In Turkey,
713:Druze Arabs
469:Yemeni Jews
238:Libyan Jews
66:Descendants
50:Middle East
7527:, Bantam,
7119:(1): 124.
6756:(1): 213,
6618:2012-05-11
6056:(1): 181,
5263:2012-05-11
4188:References
3277:Haplogroup
3108:E1b1b1c1a1
3105:E1b1b1c1a1
3102:E1b1b1c1a1
3099:E1b1b1c1a1
3090:E1b1b1c1a1
2989:E1b1b1b2a
2909:E1b1b1b1b1
2808:E1b1b1b1a
2744:E1b1b1a1c1
2726:E1b1b1a1c1
2564:E1b1a1a1e
2503:E1b1a1a1d
2381:E1b1a1a1c
2259:E1b1a1a1a
1770:ISOGG 2012
1765:ISOGG 2011
1760:ISOGG 2010
1755:ISOGG 2009
1750:ISOGG 2008
1745:ISOGG 2007
1740:ISOGG 2006
1639:Bronze Age
1597:Arslantepe
1565:Haplogroup
1535:tentative.
1193:Jordanians
1105:Bulgarians
727:Sardinians
531:Azeri Iran
308:Jordanians
99:haplogroup
7636:Nat Genet
7250:CiteSeerX
7036:Hum Hered
6679:CiteSeerX
6509:160359661
6489:Antiquity
6442:163020672
5545:Hum Genet
4490:219105572
4420:219105572
4151:PhyloTree
3986:Footnotes
3297:Paragroup
3282:Haplotype
3087:E1b1b1c1a
2927:E1b1b1b2a
2924:E1b1b1b2a
2921:E1b1b1b2a
2918:E1b1b1b2a
2866:E1b1b1b1a
2848:E1b1b1b1a
2741:E1b1b1a3a
2738:E1b1b1a3a
2735:E1b1b1a3a
2723:E1b1b1a3a
2686:E1b1b1a1
2561:E1b1a1a1e
2500:E1b1a1a1d
2378:E1b1a1a1c
2256:E1b1a1a1a
1595:found in
1524:exemplar.
1508:estimates
1466:Egyptians
1337:San Ninfa
1119:Albanians
1091:Romanians
1063:Estonians
881:Asturians
867:Corsicans
699:Armenians
629:Sicilians
153:Sephardic
149:Ashkenazi
8000:Category
7988:19058044
7941:18976729
7902:18374297
7863:11827954
7828:14504230
7798:Genetics
7785:12082121
7750:18076332
7694:11415522
7660:12893406
7652:11062480
7627:21253605
7587:PLOS ONE
7577:17002951
7515:18461126
7475:PLOS ONE
7465:23778828
7457:17039481
7420:15300852
7391:15069642
7353:11719903
7310:11073453
7272:11543889
7232:15756297
7197:11078479
7153:17662131
7101:11556974
7093:17909833
7056:16770078
7028:85134429
6997:15944443
6961:20234393
6898:14627670
6859:15896936
6830:11573163
6790:18644108
6741:17264870
6709:40355653
6701:12914567
6661:14973781
6586:22042855
6537:19425100
6480:22406638
6472:18269686
6394:(2008),
6383:18385274
6341:10904265
6308:(2011),
6298:18678889
6245:18618658
6216:12930755
6185:Genetics
6167:15996172
6127:13347549
6119:17216803
6090:19650893
6041:12772214
6012:16142507
5980:15280900
5946:17047675
5906:19686289
5855:15576591
5827:18685561
5788:17351267
5753:26886757
5745:16835895
5713:15042509
5669:11910562
5625:18183308
5585:PLOS ONE
5573:10763736
5565:14586639
5536:19156170
5489:19430804
5452:12781138
5408:17928816
5377:23156886
5369:16759179
5340:11254456
5228:16626329
5198:10310338
5190:14740294
5160:13680527
5117:19107149
5078:15202071
5029:19213004
4997:19061982
4943:12893406
4935:11062480
4906:10577926
4864:11073453
4827:11481588
4782:21432405
4774:11313742
4740:10904265
4711:11420360
4676:11170891
4606:13670282
4598:29743675
4540:31488661
4482:32470401
4412:32470401
4292:19669163
4148:, and;
4139:op. cit.
4029:23291764
4021:24166809
3659:F-Y27277
3302:Subclade
3250:Genetics
3245:See also
3047:E1b1b1c1
3044:E1b1b1c1
3041:E1b1b1c1
3038:E1b1b1c1
3029:E1b1b1c1
3026:E1b1b1c1
2906:E1b1b1b2
2863:E1b1b1b1
2860:E1b1b1b1
2857:E1b1b1b1
2845:E1b1b1b1
2805:E1b1b1b1
2787:E1b1b1b1
2683:E1b1b1a1
2665:E1b1b1a1
2625:removed
2320:removed
2265:E-M116.2
1629:and the
1627:Caucasus
1623:Anatolia
1591:and the
1589:Caucasus
1585:Anatolia
1568:E1b1b1b2
1547:Natufian
1520:found 1
1430:Dead Sea
1301:Calabria
1153:Bulgaria
1077:Russians
895:Caucasus
615:Bedouins
107:mutation
56:Ancestor
7979:2755252
7932:2668035
7893:2427286
7819:1462739
7702:9441236
7618:3017091
7595:Bibcode
7568:1635457
7506:2324201
7483:Bibcode
7428:1571356
7382:1181965
7290:Bibcode
7282:Science
7188:1287948
7144:1976131
7121:Bibcode
7064:7017701
6952:2987384
6889:1735314
6821:1274378
6781:2492877
6758:Bibcode
6652:1182266
6577:3215063
6554:Bibcode
6374:2336805
6289:2504844
6266:Bibcode
6207:1462677
6175:3229760
6081:2728732
6058:Bibcode
5937:2588664
5897:3312577
5863:8057990
5835:Science
5818:2985948
5704:1181964
5616:2174525
5593:Bibcode
5527:2947089
5497:5657112
5432:Bibcode
5331:1275654
5236:4652154
5151:1180600
5108:2947100
5069:1216069
5037:7041905
4988:2668061
4897:1288383
4844:Bibcode
4836:Science
4818:1235490
4667:1235276
4568:Bibcode
4531:6822619
4510:Science
4345:3 March
4318:bioRxiv
4283:2771134
3863:K-M2313
3856:
3797:
3782:
3768:
3752:
3743:
3734:
3556:
3539:
3486:updated
3198:(M123)
3175:Su 1999
3096:E3b1c1a
3093:E3b1c1a
2986:E1b1b1c
2983:E1b1b1c
2980:E1b1b1c
2977:E1b1b1c
2968:E1b1b1c
2965:E1b1b1c
2802:E1b1b1b
2799:E1b1b1b
2796:E1b1b1b
2784:E1b1b1b
2732:E3b1a3a
2729:E3b1a3a
2680:E1b1b1a
2677:E1b1b1a
2674:E1b1b1a
2662:E1b1b1a
2622:removed
2317:removed
2198:E1b1a1
1610:Armenia
1462:Kabylie
1454:(3.1%).
1409:Galicia
1341:Sciacca
1325:Trapani
1297:Cosenza
1275:Romania
1183:Madeira
601:Omanite
453:Bamiyan
7986:
7976:
7939:
7929:
7900:
7890:
7861:
7854:155271
7851:
7826:
7816:
7783:
7748:
7717:
7700:
7692:
7658:
7650:
7625:
7615:
7575:
7565:
7531:
7513:
7503:
7463:
7455:
7426:
7418:
7389:
7379:
7351:
7344:384897
7341:
7308:
7270:
7252:
7230:
7195:
7185:
7151:
7141:
7099:
7091:
7062:
7054:
7026:
6995:
6959:
6949:
6896:
6886:
6857:
6828:
6818:
6788:
6778:
6739:
6707:
6699:
6681:
6659:
6649:
6584:
6574:
6535:
6507:
6478:
6470:
6440:
6406:
6381:
6371:
6339:
6296:
6286:
6243:
6214:
6204:
6173:
6165:
6125:
6117:
6088:
6078:
6039:
6010:
5978:
5944:
5934:
5904:
5894:
5861:
5853:
5825:
5815:
5786:
5751:
5743:
5711:
5701:
5667:
5660:447595
5657:
5623:
5613:
5571:
5563:
5534:
5524:
5495:
5487:
5460:526263
5458:
5450:
5406:
5375:
5367:
5338:
5328:
5234:
5226:
5196:
5188:
5158:
5148:
5115:
5105:
5076:
5066:
5035:
5027:
4995:
4985:
4941:
4933:
4904:
4894:
4862:
4825:
4815:
4780:
4772:
4738:
4709:
4674:
4664:
4604:
4596:
4560:Nature
4538:
4528:
4488:
4480:
4418:
4410:
4320:
4290:
4280:
4098:K-M526
4027:
4019:
3886:
3882:
3877:
3865:
3846:
3673:
3665:
3661:
3656:
3234:E-L791
3224:(M290)
3222:E-M290
3216:(M136)
3214:E-M136
3210:(M84)
3204:(M34)
3196:E-M123
3084:E3b3a1
3056:E-M136
3035:E3b1c1
3032:E3b1c1
3020:E3b3a*
2935:E-M123
2915:E3b1b2
2912:E3b1b2
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3955:Q
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3788:L
3732:J
3726:I
3717:K
3703:H
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3641:F
3636:C
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3596:B
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3439:v
3180:η
3172:ζ
3164:ε
3156:δ
3148:γ
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3128:α
3078:B
3066:4
3017:B
3005:4
2956:B
2944:4
2897:B
2885:4
2836:B
2824:4
2775:B
2763:4
2714:B
2702:4
2653:B
2641:4
2592:B
2580:4
2531:B
2519:5
2513:8
2470:B
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2147:8
2104:B
2092:4
2073:-
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1982:B
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