35:
1774:
318:
Wang, Yan; White, Merlin M.; Moncalvo, Jean-Marc (October 2019). "Diversification of the gut fungi
Smittium and allies (Harpellales) co-occurred with the origin of complete metamorphosis of their symbiotic insect hosts (lower Diptera)".
198:
publication. Harpellales has served as a model to study and understand the evolution, growth, and biodiversity of other such fungi found in the gut as species are plentiful around the world.
1720:
1576:
407:
1733:
1628:
1485:
1815:
1550:
1589:
279:
354:
Valle LG, White MM, Cafaro MJ (2008). "Harpellales in the digestive tracts of
Ephemeroptera and Plecoptera nymphs from Veracruz, Mexico".
400:
448:
443:
1594:
1808:
1615:
527:
517:
500:
393:
1498:
34:
1738:
1633:
1844:
1801:
215:
1100:
1307:
957:
915:
874:
532:
221:
203:
1839:
1438:
1173:
907:
691:
209:
1105:
1075:
967:
733:
1503:
1052:
962:
714:
701:
683:
495:
424:
385:
1193:
744:
1080:
1711:
1643:
1340:
1287:
846:
709:
1542:
563:
296:
1834:
1317:
1292:
1090:
1042:
1032:
833:
772:
767:
91:
Tedersoo, Sanchez-Ramirez, Kõljalg, Bahram, M. Döring, Schigel, T.W. May, M. Ryberg & Abarenkov
1231:
739:
1393:
1345:
1332:
1312:
1297:
1277:
1226:
1085:
897:
869:
823:
759:
512:
490:
478:
453:
29:
1725:
271:
1746:
1651:
1511:
1360:
1355:
1350:
1302:
1153:
1003:
889:
815:
522:
485:
473:
463:
371:
336:
275:
177:
164:
are biconical. Species in the order are found attached to the gut lining of aquatic larvae of
153:
130:
76:
1785:
1751:
1476:
105:
1656:
1581:
1422:
1269:
1216:
1158:
1067:
1024:
995:
879:
841:
749:
437:
363:
328:
263:
109:
1398:
1388:
1370:
1322:
1236:
1188:
1178:
1168:
1095:
1057:
933:
856:
724:
640:
610:
605:
145:
66:
1537:
1383:
1378:
1203:
1183:
1163:
1145:
1130:
1115:
972:
864:
805:
468:
1828:
1490:
1254:
1221:
1211:
1135:
948:
938:
797:
573:
264:
1282:
1047:
925:
555:
173:
125:
1696:
1516:
1705:
1620:
1563:
1470:
615:
416:
367:
332:
1461:
1781:
1013:
786:
1669:
1524:
194:
161:
157:
46:
375:
340:
1690:
1664:
1602:
1455:
1037:
671:
662:
648:
507:
1773:
1259:
986:
189:
169:
165:
1568:
420:
185:
56:
1607:
1432:
1555:
1529:
242:
149:
1436:
389:
1789:
156:. Thalli are either unbranched or branched, producing
1680:
1445:
1369:
1331:
1268:
1253:
1202:
1144:
1123:
1114:
1066:
1023:
1012:
994:
985:
947:
924:
906:
888:
855:
832:
814:
796:
785:
758:
723:
700:
682:
670:
661:
639:
632:
598:
572:
554:
547:
262:Kirk PM, Cannon PF, Minter DW, Stalpers JA (2008).
172:. Harpellales are divided into two families, the
297:"Validation of the Harpellales and Asellariales"
270:(10th ed.). Wallingford, UK: CABI. p.
1809:
401:
8:
184:(10th edition, 2008), the order contains 38
192:. The order was formally described in 1978
1816:
1802:
1433:
1265:
1120:
1020:
991:
793:
679:
667:
636:
578:
551:
408:
394:
386:
20:
234:
321:Molecular Phylogenetics and Evolution
7:
1770:
1768:
1644:e4168515-d96a-4c96-8710-c70fbceb7d91
1788:. You can help Knowledge (XXG) by
295:Lichtwardt RW, Manier J-F (1978).
14:
1772:
33:
152:classified in the subdivision
1:
368:10.3852/mycologia.100.1.149
333:10.1016/j.ympev.2019.106550
1861:
1767:
1419:
581:
431:
122:
117:
30:Scientific classification
28:
23:
216:Gauthieromyces viviparus
160:series of trichospores.
1308:Cryptomycocolacomycetes
1101:Sporopachydermiomycetes
916:Calcarisporiellomycetes
266:Dictionary of the Fungi
182:Dictionary of the Fungi
1784:-related article is a
958:Palaeovesiarbusmycetes
908:Calcarisporiellomycota
875:Ramicandelaberomycetes
692:Neocallimastigomycetes
222:Graminella ophiuroidea
204:Allantomyces zopilotei
1425:(polyphyletic group).
1053:Schizosaccharomycetes
715:Monoblepharidomycetes
684:Neocallimastigomycota
1341:Peribolosporomycetes
1288:Agaricostilbomycetes
1174:Collemopsidiomycetes
1048:Pneumocystidomycetes
847:Entomophthoromycetes
710:Hyaloraphidiomycetes
591:Paramicrosporidiales
210:Bojamyces olmecensis
1318:Spiculogloeomycetes
1293:Cystobasidiomycetes
1106:Trigonopsidomycetes
1076:Alloascoideomycetes
1033:Archaeorhizomycetes
968:Archaeosporomycetes
834:Entomophthoromycota
773:Blastocladiomycetes
768:Physodermatomycetes
734:Caulochytriomycetes
574:Microsporidiomycota
180:. According to the
1394:Bartheletiomycetes
1346:Monilielliomycetes
1333:Ustilaginomycotina
1313:Atractiellomycetes
1298:Microbotryomycetes
1278:Tritirachiomycetes
1227:Laboulbeniomycetes
963:Paraglomeromycetes
898:Mortierellomycetes
870:Dimargaritomycetes
824:Basidiobolomycetes
760:Blastocladiomycota
702:Monoblepharomycota
1797:
1796:
1762:
1761:
1747:Open Tree of Life
1652:Open Tree of Life
1439:Taxon identifiers
1430:
1429:
1415:
1414:
1411:
1410:
1407:
1406:
1361:Exobasidiomycetes
1356:Ustilaginomycetes
1351:Malasseziomycetes
1303:Classiculomycetes
1249:
1248:
1245:
1244:
1194:Xylobotryomycetes
1154:Candelariomycetes
1004:Entorrhizomycetes
981:
980:
890:Mortierellomycota
816:Basidiobolomycota
781:
780:
745:Sanchytriomycetes
657:
656:
628:
627:
624:
623:
281:978-0-85199-826-8
178:Legeriomycetaceae
154:Kickxellomycotina
138:
137:
131:Legeriomycetaceae
113:
92:
77:Kickxellomycotina
1852:
1845:Zygomycota stubs
1818:
1811:
1804:
1776:
1769:
1755:
1754:
1742:
1741:
1729:
1728:
1716:
1715:
1714:
1701:
1700:
1699:
1673:
1672:
1660:
1659:
1647:
1646:
1637:
1636:
1624:
1623:
1621:NHMSYS0001483566
1611:
1610:
1598:
1597:
1585:
1584:
1572:
1571:
1559:
1558:
1546:
1545:
1533:
1532:
1520:
1519:
1507:
1506:
1494:
1493:
1481:
1480:
1479:
1466:
1465:
1464:
1434:
1423:fungi imperfecti
1270:Pucciniomycotina
1266:
1217:Geoglossomycetes
1159:Coniocybomycetes
1121:
1081:Dipodascomycetes
1068:Saccharomycotina
1025:Taphrinomycotina
1021:
996:Entorrhizomycota
992:
880:Kickxellomycetes
842:Neozygitomycetes
794:
750:Chytridiomycetes
680:
668:
637:
579:
552:
460:
440:
423:classification,
410:
403:
396:
387:
380:
379:
351:
345:
344:
315:
309:
308:
292:
286:
285:
269:
259:
253:
252:
250:
249:
239:
201:Species include
104:
90:
38:
37:
21:
1860:
1859:
1855:
1854:
1853:
1851:
1850:
1849:
1825:
1824:
1823:
1822:
1765:
1763:
1758:
1750:
1745:
1737:
1732:
1724:
1719:
1712:Harpellomycetes
1710:
1709:
1704:
1695:
1694:
1689:
1682:Harpellomycetes
1676:
1668:
1663:
1655:
1650:
1642:
1640:
1632:
1627:
1619:
1614:
1606:
1601:
1593:
1588:
1580:
1575:
1567:
1562:
1554:
1549:
1541:
1536:
1528:
1523:
1515:
1510:
1502:
1497:
1489:
1484:
1475:
1474:
1469:
1460:
1459:
1454:
1441:
1431:
1426:
1403:
1399:Tremellomycetes
1389:Wallemiomycetes
1371:Agaricomycotina
1365:
1327:
1323:Pucciniomycetes
1257:
1241:
1237:Sordariomycetes
1198:
1189:Lecanoromycetes
1179:Dothideomycetes
1169:Arthoniomycetes
1140:
1110:
1096:Saccharomycetes
1062:
1058:Taphrinomycetes
1038:Neolectomycetes
1016:
1008:
977:
943:
934:Endogonomycetes
920:
902:
884:
857:Kickxellomycota
851:
828:
810:
789:
777:
754:
725:Chytridiomycota
719:
696:
675:
653:
641:Aphelidiomycota
633:Aphelidiomyceta
620:
611:Metchnikovellea
606:Chytridiopsidea
594:
585:Morellosporales
568:
543:
542:
458:
436:
427:
414:
384:
383:
353:
352:
348:
317:
316:
312:
294:
293:
289:
282:
261:
260:
256:
247:
245:
241:
240:
236:
231:
103:
89:
87:Harpellomycetes
67:Kickxellomycota
32:
17:
12:
11:
5:
1858:
1856:
1848:
1847:
1842:
1837:
1827:
1826:
1821:
1820:
1813:
1806:
1798:
1795:
1794:
1777:
1760:
1759:
1757:
1756:
1743:
1730:
1717:
1702:
1686:
1684:
1678:
1677:
1675:
1674:
1661:
1648:
1638:
1625:
1612:
1599:
1586:
1573:
1560:
1547:
1534:
1521:
1508:
1495:
1482:
1467:
1451:
1449:
1443:
1442:
1437:
1428:
1427:
1420:
1417:
1416:
1413:
1412:
1409:
1408:
1405:
1404:
1402:
1401:
1396:
1391:
1386:
1384:Agaricomycetes
1381:
1379:Dacrymycetales
1375:
1373:
1367:
1366:
1364:
1363:
1358:
1353:
1348:
1343:
1337:
1335:
1329:
1328:
1326:
1325:
1320:
1315:
1310:
1305:
1300:
1295:
1290:
1285:
1280:
1274:
1272:
1263:
1251:
1250:
1247:
1246:
1243:
1242:
1240:
1239:
1234:
1229:
1224:
1219:
1214:
1208:
1206:
1204:Sordariomyceta
1200:
1199:
1197:
1196:
1191:
1186:
1184:Eurotiomycetes
1181:
1176:
1171:
1166:
1164:Lichinomycetes
1161:
1156:
1150:
1148:
1146:Dothideomyceta
1142:
1141:
1139:
1138:
1133:
1131:Orbiliomycetes
1127:
1125:
1118:
1116:Pezizomycotina
1112:
1111:
1109:
1108:
1103:
1098:
1093:
1088:
1083:
1078:
1072:
1070:
1064:
1063:
1061:
1060:
1055:
1050:
1045:
1040:
1035:
1029:
1027:
1018:
1010:
1009:
1007:
1006:
1000:
998:
989:
983:
982:
979:
978:
976:
975:
973:Glomeromycetes
970:
965:
960:
953:
951:
945:
944:
942:
941:
936:
930:
928:
922:
921:
919:
918:
912:
910:
904:
903:
901:
900:
894:
892:
886:
885:
883:
882:
877:
872:
867:
865:Zoopagomycetes
861:
859:
853:
852:
850:
849:
844:
838:
836:
830:
829:
827:
826:
820:
818:
812:
811:
809:
808:
806:Olpidiomycetes
802:
800:
791:
790:(paraphyletic)
783:
782:
779:
778:
776:
775:
770:
764:
762:
756:
755:
753:
752:
747:
742:
736:
729:
727:
721:
720:
718:
717:
712:
706:
704:
698:
697:
695:
694:
688:
686:
677:
676:(paraphyletic)
665:
659:
658:
655:
654:
652:
651:
645:
643:
634:
630:
629:
626:
625:
622:
621:
619:
618:
613:
608:
602:
600:
596:
595:
593:
592:
589:
588:Nucleophagales
586:
582:
576:
570:
569:
567:
566:
564:Rozellomycetes
560:
558:
549:
545:
544:
541:
540:
539:
538:
537:
536:
533:Mesomycetozoea
530:
525:
515:
505:
504:
503:
498:
493:
488:
483:
482:
481:
469:Diaphoretickes
466:
461:
456:
451:
446:
441:
433:
432:
429:
428:
415:
413:
412:
405:
398:
390:
382:
381:
346:
310:
287:
280:
254:
233:
232:
230:
227:
136:
135:
134:
133:
128:
120:
119:
115:
114:
98:
94:
93:
84:
80:
79:
74:
70:
69:
64:
60:
59:
54:
50:
49:
44:
40:
39:
26:
25:
16:Order of fungi
15:
13:
10:
9:
6:
4:
3:
2:
1857:
1846:
1843:
1841:
1840:Fungus orders
1838:
1836:
1833:
1832:
1830:
1819:
1814:
1812:
1807:
1805:
1800:
1799:
1793:
1791:
1787:
1783:
1778:
1775:
1771:
1766:
1753:
1748:
1744:
1740:
1735:
1731:
1727:
1722:
1718:
1713:
1707:
1703:
1698:
1692:
1688:
1687:
1685:
1683:
1679:
1671:
1666:
1662:
1658:
1653:
1649:
1645:
1639:
1635:
1630:
1626:
1622:
1617:
1613:
1609:
1604:
1600:
1596:
1591:
1587:
1583:
1578:
1574:
1570:
1565:
1561:
1557:
1552:
1548:
1544:
1539:
1535:
1531:
1526:
1522:
1518:
1513:
1509:
1505:
1500:
1496:
1492:
1487:
1483:
1478:
1472:
1468:
1463:
1457:
1453:
1452:
1450:
1448:
1444:
1440:
1435:
1424:
1418:
1400:
1397:
1395:
1392:
1390:
1387:
1385:
1382:
1380:
1377:
1376:
1374:
1372:
1368:
1362:
1359:
1357:
1354:
1352:
1349:
1347:
1344:
1342:
1339:
1338:
1336:
1334:
1330:
1324:
1321:
1319:
1316:
1314:
1311:
1309:
1306:
1304:
1301:
1299:
1296:
1294:
1291:
1289:
1286:
1284:
1281:
1279:
1276:
1275:
1273:
1271:
1267:
1264:
1261:
1256:
1255:Basidiomycota
1252:
1238:
1235:
1233:
1230:
1228:
1225:
1223:
1222:Leotiomycetes
1220:
1218:
1215:
1213:
1212:Xylonomycetes
1210:
1209:
1207:
1205:
1201:
1195:
1192:
1190:
1187:
1185:
1182:
1180:
1177:
1175:
1172:
1170:
1167:
1165:
1162:
1160:
1157:
1155:
1152:
1151:
1149:
1147:
1143:
1137:
1136:Pezizomycetes
1134:
1132:
1129:
1128:
1126:
1122:
1119:
1117:
1113:
1107:
1104:
1102:
1099:
1097:
1094:
1092:
1091:Pichiomycetes
1089:
1087:
1084:
1082:
1079:
1077:
1074:
1073:
1071:
1069:
1065:
1059:
1056:
1054:
1051:
1049:
1046:
1044:
1043:Novakomycetes
1041:
1039:
1036:
1034:
1031:
1030:
1028:
1026:
1022:
1019:
1015:
1011:
1005:
1002:
1001:
999:
997:
993:
990:
988:
984:
974:
971:
969:
966:
964:
961:
959:
955:
954:
952:
950:
949:Glomeromycota
946:
940:
939:Mucoromycetes
937:
935:
932:
931:
929:
927:
923:
917:
914:
913:
911:
909:
905:
899:
896:
895:
893:
891:
887:
881:
878:
876:
873:
871:
868:
866:
863:
862:
860:
858:
854:
848:
845:
843:
840:
839:
837:
835:
831:
825:
822:
821:
819:
817:
813:
807:
804:
803:
801:
799:
798:Olpidiomycota
795:
792:
788:
784:
774:
771:
769:
766:
765:
763:
761:
757:
751:
748:
746:
743:
741:
737:
735:
731:
730:
728:
726:
722:
716:
713:
711:
708:
707:
705:
703:
699:
693:
690:
689:
687:
685:
681:
678:
673:
669:
666:
664:
660:
650:
647:
646:
644:
642:
638:
635:
631:
617:
614:
612:
609:
607:
604:
603:
601:
599:Microsporidia
597:
590:
587:
584:
583:
580:
577:
575:
571:
565:
562:
561:
559:
557:
556:Rozellomycota
553:
550:
548:Rozellomyceta
546:
534:
531:
529:
526:
524:
521:
520:
519:
516:
514:
511:
510:
509:
506:
502:
499:
497:
496:Stramenopiles
494:
492:
489:
487:
484:
480:
477:
476:
475:
472:
471:
470:
467:
465:
462:
459:(major groups
457:
455:
452:
450:
447:
445:
442:
439:
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425:fungal orders
422:
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362:(1): 149–62.
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36:
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1790:expanding it
1779:
1764:
1681:
1446:
1283:Mixiomycetes
1232:Sareomycetes
926:Mucoromycota
740:Coenomycetes
518:Opisthokonta
359:
355:
349:
324:
320:
313:
307:(3): 441–42.
304:
300:
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257:
246:. Retrieved
243:"Zygomycota"
237:
220:
214:
208:
202:
200:
193:
181:
174:Harpellaceae
168:or (rarely)
141:
139:
126:Harpellaceae
100:
86:
24:Harpellales
18:
1706:Wikispecies
1564:iNaturalist
1477:Harpellales
1471:Wikispecies
1447:Harpellales
1086:Lipomycetes
1017:(sac fungi)
616:Microsporea
417:Opisthokont
142:Harpellales
101:Harpellales
1835:Zygomycota
1829:Categories
1782:Zygomycota
1421:See also:
1014:Ascomycota
787:Zygomycota
421:True fungi
327:: 106550.
248:2009-03-07
229:References
162:Zygospores
63:Division:
1697:Q54281948
1670:100364847
672:Zoosporic
649:Aphelidea
513:Amoebozoa
491:Alveolata
479:Cryptista
454:Eukaryota
356:Mycologia
301:Mycotaxon
195:Mycotaxon
158:basipetal
118:Families
53:Kingdom:
47:Eukaryota
1726:11914456
1691:Wikidata
1665:Tropicos
1603:MycoBank
1538:Fungorum
1491:60013271
1486:AusFungi
1462:Q1055276
1456:Wikidata
663:Eumycota
508:Amorphea
486:Rhizaria
474:Hacrobia
464:Excavata
449:Bacteria
376:18488361
341:31279967
188:and 200
176:and the
43:Domain:
1752:7510250
1739:2219693
1260:basidia
987:Dikarya
523:Animals
444:Archaea
190:species
170:Isopoda
166:Insecta
144:are an
106:Lichtw.
97:Order:
83:Class:
1657:138164
1641:NZOR:
1595:936428
1569:153136
1530:1HARPO
1258:(with
501:Plants
438:Domain
374:
339:
278:
186:genera
112:(1978)
110:Manier
108:&
1780:This
1721:IRMNG
1634:61421
1608:90427
1582:10082
1577:IRMNG
1543:90427
1504:23739
1124:Other
674:fungi
528:Fungi
150:fungi
146:order
57:Fungi
1786:stub
1734:NCBI
1629:NCBI
1590:ITIS
1551:GBIF
1525:EPPO
1499:BOLD
372:PMID
337:PMID
276:ISBN
219:and
140:The
1616:NBN
1556:747
1517:3BM
1512:CoL
364:doi
360:100
329:doi
325:139
272:305
148:of
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1817:e
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1262:)
956:†
738:?
732:?
535:)
409:e
402:t
395:v
378:.
366::
343:.
331::
305:7
284:.
251:.
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