478:
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437:
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110:
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44:
83:
63:
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in eastern North
America), including moist soil and damp crevices in old walls; they are found most commonly in shaded areas. Plants in full sun are usually stunted and yellowish in colour, while those in full shade are dark green and healthy. The disjunct populations of the North American variation
974:
is found across the vast majority of
European countries, in a wide variety of natural settings, across a multitude of habitats. It is also able to colonise manmade environments including growing in walls and brickwork crevices, particularly in limestone mortar. Furthermore, it is considered to be a
1571:
Kartesz, J.T. 1999. A synonymized checklist and atlas with biological attributes for the vascular flora of the United States, Canada, and
Greenland. First edition. In: Kartesz, J.T., and C.A. Meacham. Synthesis of the North American Flora, Version 1.0. North Carolina Botanical Garden, Chapel Hill,
2130:
Xu, Ke-Wang; Zhang, Liang; Rothfels, Carl J.; Smith, Alan R.; Viane, Ronald; Lorence, David; Wood, Kenneth R.; Cheng, Cheng-Wei; Knapp, Ralf; Zhou, Lin; Lu, Ngan Thi; Zhou, Xin-Mao; Wei, Hong-Jin; Fan, Qiang; Chen, Su-Fang; Cicuzza, Daniele; Gao, Xin-Fen; Li, Wen-Bo; Zhang, Li-Bing (2020).
983:
frequently grows on vertical faces and in crevices of lime mortared brick walls, the species may be threatened by wall renovation projects. Although the fern is often cultivated as a garden plant or used in landscape planting, commercially sold plants are sourced from horticultural stock.
978:
Across Europe as a whole, the IUCN assessed no immediate threats to this species in natural settings, and predicted that the species may benefit from milder wetter winters predicted under various climate change scenarios. In the urban environment, where
1128:
tonic (hence an archaic name for the genus, "spleenworts") and for other uses. "Hartstongue Ale" is mentioned in the recipe book of
Katharine Palmer (compiled from 1700). It was made by putting three leaves of hart's-tongue fern into a bottle of
1063:
of any region in the variation's distribution, with around 80% of all subpopulations and around 94% of all individuals. The fern was reported at more than 100 sites across the province, with around 75 still believed to be existing. Despite this,
2202:
A popular article on hart's tongue fern that includes several references and a discussion of cultivation possibilities for the
European and American varieties. The article strongly discourages collection and or cultivation of the North American
914:
is distributed in the
Russian Far East, north-eastern China and the Korean Peninsula, however it is considered as being generally rare on the mainland. In contrast, it is relatively abundant on many islands including the Japanese Archipelago.
2077:
The family herbal: or An account of all those
English plants, which are remarkable for their virtues, and of the drugs which are produced by vegetables of other countries; with their descriptions and their uses, as proved by
902:; this represents the first confirmed population of the species in North America west of the Mississippi. Genetic analyses and surveys are currently being performed to determine the population's variation and overall health.
1111:
The
American variety is reputed to be difficult to cultivate (making conservation efforts for it even more troublesome); due to this, most, if not all, cultivated individuals are derived from the Old World subspecies.
1335:
922:
has caught the attention of international botanists. In fact, the very existence of such varieties beg that "...these populations arose following colonization events involving a single spore".
1748:
Weston L. Testo, James E. Watkins Jr, John Wiley, Laura
Baumann, Eric Weaver "SHORTER NOTE," American Fern Journal, 111(3), 217-222, (2 September 2021). Retrieved March 17, 2023, from
1669:
994:
inhabits a large number of legally protected areas across the continent. Despite this, it is considered to be declining in numbers or a species at risk in certain parts of its range.
560:
subclade have undivided leaves with freely branching veins and single or paired sori. They are widely distributed through the
Northern Hemisphere. If defined to include the former
975:
gregarious species - it is able to spread both sexually and asexually, frequently forming large clumps and present in high densities - in some areas it is considered very common.
1749:
1726:
Lin YX, Viane R. 2013. Aspleniaceae. In: Wu ZY, Raven PH, Hong DY, eds. Flora of China, Vol. 2-3. Beijing: Science Press; St. Louis: Missouri Botanical Garden Press, 274.
1076:
in Ontario in May 2017, due to its extremely specific habitat requirements, relatively small distribution, and some subpopulations consisting of too little individuals.
1892:"Experimental Reintroduction of American Hart's-Tongue Fern (Asplenium scolopendrium var. americanum) : Factors Affecting Successful Establishment of Transplants"
2408:
2645:
948:
2470:
1916:
1534:
Pence, V. C. (2015, July). Propagation and cryopreservation of Asplenium scolopendrium var ... - JSTOR. American Fern Journal. Retrieved March 17, 2023, from
1006:
in 2010 (under Criterion D1, that is, having a very small population of under 1000 individuals). It is considered "critically threatened and rare" in the
2665:
1700:
959:
being listed as being a species of Least Concern (LC). Despite the positive status, this assessment is chiefly limited to the conservation situation of
1073:
1184:
2356:
1270:
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1170:
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2099:
A Collection of ye best Receipts most approved and fittest in Cookery, preserving, and all manner of Housewifery, physick & Chirurgery
740:). It is absent however in northeast Europe (Finland, Belarus, Latvia, Lithuania and European Russia). Beyond Europe, it is found in the
1659:
Snyder, D.B. 1990. Botanist, New Jersey Natural Heritage Program. Personal communication with Wayne Ostlie, MRO, The Nature Conservancy.
1236:
1156:
1750:
https://bioone.org/journals/american-fern-journal/volume-111/issue-3/0002-8444-111.3.217/SHORTER-NOTE/10.1640/0002-8444-111.3.217.short
2320:
1319:
860:
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selected with varying frond form, including with frilled frond margins, forked fronds and cristate forms. The species has gained the
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2189:
2058:
2036:
1644:
552:
clade have undivided or pinnatifid leaf blades with a thick, leathery texture, persistent scales on their stalk, and often possess
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2395:
477:
90:
70:
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in the southeastern US are found exclusively in sinkhole pits or limestone caves. These populations may be relics of cooler
990:
is not a specifically protected species in the majority of European states, however in part due to its wide distribution,
1211:
536:
published in 2020 divided the genus into eleven clades, which were given informal names pending further taxonomic study.
895:
1607:"Species Status Assessment : Report for the American Hart's-tongue Fern (Asplenium scolopendrium var. americanum)"
2558:
1101:
2262:
1917:"Ontario Species at Risk Evaluation Report for American Hart's-tongue Fern (Asplenium scolopendrium var. americanum)"
1261:
1855:
GÀrdenfors, U. 2010. Rödlistade arter i Sverige - The 2010 Red List of Swedish Species. ArtDatabanken, SLU, Uppsala.
1492:
Heo, Namjoo; Lomolino, Mark V; Watkins, James E; Yun, Seona; Weber-Townsend, Josh; Fernando, Danilo D (2022-09-01).
2545:
1891:
1144:
109:
2475:
2444:
1998:
1837:
KĂ„lĂ„s, J.A., Viken, Ă
. and Bakken, T. (eds). 2006. Norsk RĂždliste 2006 â 2006 Norwegian Red List. Artsdatabanken.
930:
The plants grow on neutral, calcium-rich, and/or lime-rich substrates under deciduous hardwood canopies (usually
899:
2550:
2675:
772:
of Europe is used for many horticultural uses. Specimens of this variety found in North America (such as in
1401:"Molecular phylogenetic assessment of three major taxa in the Asplenium scolopendrium complex (Aspleniaceae)"
864:
436:
2338:
1430:
2680:
2333:
2224:
894:
In 2020, a new population of hart's-tongue ferns was discovered inside a cave with basaltic lava flows in
868:
562:
1105:
511:
498:
217:
1547:
1467:
Christenhusz, M., Bento Elias, R., Dyer, R., Ivanenko, Y., Rouhan, G., Rumsey, F. & VĂ€re, H. 2017.
463:
2374:
1969:
1846:
Grulich, V. 2012. Red List of vascular plants of the Czech Republic: 3rd edition. Preslia 84: 631-645.
2622:
2511:
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888:
576:
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852:
52:
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880:
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104:
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2054:
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1517:
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506:
449:
1498:
complex (Aspleniaceae), a relictual fern with a northern pan-temperate disjunct distribution"
2586:
2498:
2144:
1582:
1509:
1412:
1093:
840:
836:
765:
407:
1765:
867:, where it has declined heavily due to illegal plant collecting, and an undisclosed pit in
1121:
844:
689:
Morphological differences between the varieties are minor, but the North American variety
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1944:
1400:
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The most striking and unusual feature of the fern is its simple, strap-shaped undivided
43:
2519:
1044:
1007:
761:
134:
17:
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2166:
2116:
952:
805:
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773:
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233:
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1606:
607:
599:
410:
184:
174:
164:
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2120:
784:
in the United States) are considered naturalized descendants of cultivated plants.
2563:
2047:
1548:"Asplenium scolopendrium L. | Plants of the World Online | Kew Science"
1513:
2596:
2457:
2436:
2382:
2256:
2097:
1015:
936:
931:
876:
792:, it occurs in rare, widely scattered populations located in different locales:
741:
603:
553:
426:
2247:
1713:
Lee, Y. M., and W. Y. Lee. "Illustrated rare and endangered species in Korea."
2213:
1735:
Dong, Shiyong, et al. "Red list assessment of lycophytes and ferns in China."
884:
817:
694:
645:
2325:
2197:
1521:
1493:
2307:
872:
813:
698:
526:
400:
194:
2576:
2158:
583:
Three subspecies were accepted in the most recent revision of the species:
875:(in just a single county); these southern populations are at dire risk of
2571:
2241:
2214:
Commentary and video on Hart's Tongue ferns at Lynn Glen, Dalry, Scotland
1097:
1043:
in 1989. The reasons for its rarity are currently being researched, with
848:
801:
781:
733:
430:
2348:
1535:
2361:
2133:"A global plastid phylogeny of the fern genus Asplenium (Aspleniaceae)"
999:
856:
797:
777:
753:
749:
745:
706:
629:
2387:
2149:
2132:
1416:
1125:
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809:
729:
679:
671:
641:
633:
595:
2400:
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1370:
1368:
1074:
Special Concern under the Committee on the Status of Species at Risk
610:. Populations in the eastern Mediterranean have been referred to as
1813:
1431:"Asplenium scolopendrium var. americanum in Flora of North America"
2312:
1274:
821:
757:
683:
675:
522:
476:
462:
448:
435:
422:
121:
403:
154:
144:
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2222:
1014:'s 2010 Red List. The species has been protected by law in the
998:
was first listed as "Vulnerable" in the National Red Lists for
1130:
425:. The supposed resemblance of the leaves to the tongue of a
2125:. Vol. II (1st ed.). Stockholm: Laurentii Salvii.
963:
in Europe, as opposed to other parts of its natural range.
1771:(Report). Atlanta, Georgia: U.S. Fish and Wildlife Service
863:, a wildlife refuge centered around an off-limits cave in
2180:
Hyde, H. A., Wade, A. E., & Harrison, S. G. (1978).
2102:. Unpublished manuscript (wellcome collection). p.
1047:
programs in New York and elsewhere also in development.
955:
was conducted in 2016, with the conservation status of
521:, meaning a centipede or millipede; this is due to the
1055:
Ontario, Canada has the highest population numbers of
883:
population descended from New York plants is found in
1475:
2017: e.T83504456A83506952. Accessed on 19 June 2024.
1399:
Heo, Namjoo; Yun, Seona; Fernando, Danilo D. (2023).
433:) gave rise to the common name "hart's-tongue fern".
1463:
1461:
1459:
1457:
1455:
1453:
1451:
1386:
1374:
1359:
648:. The tropical populations have been referred to as
2231:
1583:"Tropicos | Name - Asplenium scolopendrium L."
2046:
1639:. Tuscaloosa, AL: University of Alabama Press.
887:; it is a remnant of a 1936 effort to practice
949:International Union for Conservation of Nature
1790:"Asplenium scolopendrium, hart's tongue fern"
1635:Short, John W.; Spaulding, Daniel D. (2012).
8:
1010:'s 2012 plant Red List; and "Endangered" in
871:that is also off-limits and protected), and
1487:
1485:
1483:
1481:
1469:Asplenium scolopendrium (Europe assessment)
1177:A specimen growing in lime mortar on a wall
2219:
81:
61:
42:
31:
27:Species of fern in the family Aspleniaceae
2148:
2081:. C. Brightly and T. Kinnersley. p.
1759:
1757:
1715:Korea National Arboretum. Po-cheon, Korea
1502:Biological Journal of the Linnean Society
843:(present in 7 counties), and the eastern
1297:
1271:Botanical Society of Britain and Ireland
851:(present in 2 counties). Exceptions are
1630:
1628:
1473:The IUCN Red List of Threatened Species
1203:
1140:
953:The IUCN Red List of Threatened Species
768:. Unlike its American counterpart, the
254:
1699:: CS1 maint: archived copy as title (
1692:
764:). It has also been introduced to the
720:is a common species in the Old World:
497:first gave the hart's-tongue fern the
2646:NatureServe apparently secure species
1536:https://www.jstor.org/stable/44076071
1191:Example of the North American variety
1163:A cultivar with frilled frond margins
1120:This fern was used in the 1800s as a
1108:, as has the cultivar 'Angustatum'.
7:
2486:a93683ac-fe3b-4f25-8c45-c93b28c6d97e
2200:. Finger Lakes Native Plant Society.
1766:American hart's-tongue recovery plan
1764:Currie, Robert R. (September 1993).
1314:. United Kingdom: Mitchell Beazley.
570:makes up the former segregate genus
2196:Parker, Rosemarie (December 2009).
413:native to the Northern Hemisphere.
2551:urn:lsid:ipni.org:names:30216020-2
861:Fern Cave National Wildlife Refuge
25:
2666:Medicinal plants of North America
1183:
1169:
1155:
1143:
108:
1814:"The National Red List Project"
1336:"Flora Europaea Search Results"
1039:was declared endangered by the
525:pattern being reminiscent of a
308:(Fernald) Ă. Löve & D. Löve
264:(KĂŒmmerle) Brownsey & Jermy
2661:Garden plants of North America
2651:NatureServe vulnerable species
1041:U.S. Fish and Wildlife Service
703:A. scolopendrium scolopendrium
1:
2043:This book is a reprinting of
1945:"American Hart's Tongue Fern"
1494:"Evolutionary history of the
2184:. National Museum of Wales.
1072:was listed as a species of
896:El Malpais National Monument
429:(an archaic term for a male
2691:Taxa named by Carl Linnaeus
1310:Harrison, Lorraine (2012).
1102:Royal Horticultural Society
891:of populations in New York.
369:Scolopendrium scolopendrium
2707:
2096:Palmer, Katharine (1700).
2049:Ferns for American Gardens
2029:Ferns for American Gardens
1552:Plants of the World Online
1514:10.1093/biolinnean/blac080
1237:"NatureServe Explorer 2.0"
1212:"NatureServe Explorer 2.0"
517:is derived from the Greek
900:Cibola County, New Mexico
839:(present in 2 counties),
835:geological formations in
796:in the Mexican states of
361:Scolopendrium officinarum
247:
240:
223:
216:
105:Scientific classification
103:
79:
59:
50:
41:
34:
2671:Plants described in 1753
2045:Mickel, John T. (1994).
2027:Mickel, John T. (2003).
1241:Explorer.natureserve.org
1216:Explorer.natureserve.org
918:The unique dispersal of
395:, commonly known as the
2512:Asplenium scolopendrium
2263:Asplenium scolopendrium
2233:Asplenium scolopendrium
2001:Asplenium scolopendrium
1972:Asplenium scolopendrium
1496:Asplenium scolopendrium
1312:RHS Latin for Gardeners
1090:Asplenium scolopendrium
996:Asplenium scolopendrium
988:Asplenium scolopendrium
972:Asplenium scolopendrium
957:Asplenium scolopendrium
920:Asplenium scolopendrium
865:Jackson County, Alabama
744:, and North Africa (in
718:Asplenium scolopendrium
670:, is native to eastern
548:clade". Members of the
502:Asplenium scolopendrium
483:Asplenium scolopendrium
468:Asplenium scolopendrium
454:Asplenium scolopendrium
441:Asplenium scolopendrium
392:Asplenium scolopendrium
341:Phyllitis scolopendrium
329:Phyllitis scolopendrium
321:Phyllitis scolopendrium
257:Asplenium scolopendrium
227:Asplenium scolopendrium
36:Asplenium scolopendrium
18:Hart's-tongue fern
1970:"RHS Plant Selector -
1896:Digitalcommons.esf.edu
1124:in folk medicine as a
1031:In the United States,
951:(IUCN) assessment for
556:veins. Members of the
532:A global phylogeny of
486:
471:
457:
443:
353:Scolopendrium lindenii
269:Biropteris antri-jovis
2686:Ferns of the Americas
1106:Award of Garden Merit
1092:is often grown as an
908:A. scolopendrium
660:A. scolopendrium
650:A. scolopendrium
622:A. scolopendrium
612:A. scolopendrium
588:A. scolopendrium
568:A. scolopendrium
480:
466:
452:
439:
377:Scolopendrium vulgare
285:Phyllitis fernaldiana
277:Phyllitis antri-jovis
209:A. scolopendrium
1866:"Flora- en faunawet"
1737:Biodiversity Science
1340:Websites.rbge.org.uk
889:ex-situ conservation
853:disjunct populations
2074:Hill, John (1812).
1818:Nationalredlist.org
1586:Legacy.tropicos.org
510:of 1753. The Latin
53:Conservation status
1870:Wetten.overheid.nl
833:Niagara Escarpment
829:Onondaga Limestone
728:occurs throughout
577:A. sagittatum
544:subclade" of the "
487:
472:
458:
444:
397:hart's-tongue fern
313:Phyllitis lindenii
301:Phyllitis japonica
293:Phyllitis japonica
67:Apparently Secure
2633:
2632:
2494:Open Tree of Life
2225:Taxon identifiers
2150:10.1111/cla.12384
2122:Species Plantarum
1890:Michael Serviss.
1739:25.7 (2017): 765.
1417:10.1002/tax.12890
812:, as well as the
760:, as well as the
668:A. komarovii
574:and is sister to
563:A. komarovii
507:Species Plantarum
388:
387:
381:
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16:(Redirected from
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2031:. Timber Press.
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1675:. Archived from
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1637:Ferns of Alabama
1632:
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1273:. Archived from
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1233:
1227:
1226:
1224:
1222:
1208:
1187:
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1094:ornamental plant
1066:A. scolopendrium
1057:A. scolopendrium
1033:A. scolopendrium
992:A. scolopendrium
981:A. scolopendrium
961:A. scolopendrium
947:The most recent
841:southern Ontario
837:Central New York
766:Falkland Islands
540:belongs to the "
538:A. scolopendrium
512:specific epithet
379:
371:
363:
355:
347:
335:
323:
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307:
295:
287:
280:(KĂŒmmerle) Seitz
279:
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113:
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88:
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68:
65:
64:
46:
32:
21:
2706:
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2676:Ferns of Europe
2636:
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2255:
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2210:
2198:"A Real Rarity"
2195:
2177:
2175:Further reading
2129:
2115:
2112:
2111:
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2090:
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1670:"Archived copy"
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1614:
1613:. November 2019
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1309:
1308:
1304:
1300:, p. 1079.
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1165:
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1160:
1151:
1148:
1139:
1122:medicinal plant
1118:
1116:Herbal medicine
1096:, with several
1087:
1082:
1053:
1029:
1024:
969:
945:
928:
845:Upper Peninsula
780:in Canada, and
732:(including the
724:The subspecies
715:
492:
474:
460:
446:
419:
384:
253:
252:
236:
231:
225:
212:
107:
99:
86:
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75:
66:
62:
55:
28:
23:
22:
15:
12:
11:
5:
2704:
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2663:
2658:
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2610:wfo-0001110391
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2208:External links
2206:
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1508:(2): 183â199.
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1411:(2): 245â260.
1391:
1387:Xu et al. 2020
1379:
1375:Xu et al. 2020
1364:
1360:Xu et al. 2020
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1321:978-1845337315
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1008:Czech Republic
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906:In East Asia,
904:
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855:that exist in
825:
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2053:. MacMillan.
2051:
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2038:9780881925982
2034:
2030:
2023:
2020:
2008:
2004:
2003:'Angustatum'"
2002:
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1682:on 2021-11-11
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1402:
1395:
1392:
1389:, p. 41.
1388:
1383:
1380:
1377:, p. 30.
1376:
1371:
1369:
1365:
1362:, p. 27.
1361:
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1317:
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1298:Linnaeus 1753
1294:
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1280:on 2015-06-26
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869:Morgan County
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790:North America
783:
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774:New Brunswick
771:
770:scolopendrium
767:
763:
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755:
751:
747:
743:
739:
738:British Isles
735:
731:
727:
726:scolopendrium
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638:United States
635:
631:
628:is native to
627:
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617:
613:
609:
605:
601:
597:
594:is native to
593:
592:scolopendrium
589:
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515:scolopendrium
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345:scolopendrium
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316:(Hook.) Maxon
314:
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278:
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218:Binomial name
215:
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198:
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183:
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135:Tracheophytes
133:
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117:
116:
111:
106:
102:
96:
92:
78:
72:
58:
54:
49:
45:
40:
37:
33:
30:
19:
2525:tro-26603800
2232:
2181:
2143:(1): 22â71.
2140:
2136:
2121:
2117:Linnaeus, C.
2098:
2091:
2076:
2069:
2048:
2028:
2022:
2010:. Retrieved
2006:
2000:
1993:
1981:. Retrieved
1977:
1971:
1964:
1952:. Retrieved
1948:
1939:
1927:. Retrieved
1923:
1911:
1899:. Retrieved
1895:
1885:
1873:. Retrieved
1869:
1860:
1851:
1842:
1833:
1821:. Retrieved
1817:
1808:
1797:. Retrieved
1793:
1784:
1773:. Retrieved
1744:
1736:
1731:
1722:
1714:
1709:
1684:. Retrieved
1677:the original
1664:
1655:
1636:
1615:. Retrieved
1611:Ecos.fws.gov
1610:
1601:
1589:. Retrieved
1585:
1577:
1567:
1555:. Retrieved
1551:
1542:
1530:
1505:
1501:
1495:
1472:
1468:
1438:. Retrieved
1434:
1425:
1408:
1404:
1394:
1382:
1355:
1343:. Retrieved
1339:
1330:
1311:
1305:
1293:
1282:. Retrieved
1275:the original
1262:
1256:
1244:. Retrieved
1240:
1231:
1219:. Retrieved
1215:
1206:
1119:
1110:
1089:
1088:
1069:
1065:
1060:
1056:
1054:
1036:
1032:
1030:
1018:since 1998.
1002:in 2014 and
995:
991:
987:
986:
980:
977:
971:
970:
960:
956:
946:
943:Conservation
932:sugar maples
929:
919:
917:
911:
907:
905:
787:
769:
725:
717:
716:
713:Distribution
702:
697:, while the
690:
688:
667:
663:
659:
653:
649:
625:
621:
615:
611:
608:Central Asia
600:North Africa
591:
590:subsp.
587:
582:
575:
571:
567:
561:
557:
554:anastomosing
549:
545:
541:
537:
533:
531:
518:
514:
505:
501:
493:
482:
473:
467:
459:
453:
445:
440:
420:
411:Aspleniaceae
396:
391:
390:
389:
376:
368:
360:
352:
344:
340:
332:
328:
320:
312:
304:
300:
292:
284:
276:
268:
260:
256:
226:
224:
208:
207:
195:
185:Aspleniaceae
175:Aspleniineae
165:Polypodiales
128:
94:
35:
29:
2458:NatureServe
2383:iNaturalist
2257:Wikispecies
2182:Welsh Ferns
2012:20 February
1983:12 February
1717:(1997): 41.
1435:Efloras.org
1085:Cultivation
1016:Netherlands
937:Pleistocene
877:extirpation
742:Middle East
701:subspecies
666:, formerly
640:, southern
616:antri-jovis
604:Middle East
519:skolopendra
417:Description
348:(L.) Newman
324:(L.) Newman
261:antri-jovis
91:NatureServe
87:Vulnerable
71:NatureServe
2640:Categories
2520:Plant List
2401:17051170-1
2137:Cladistics
2078:experience
2007:Rhs.org.uk
1978:Rhs.org.uk
1949:Ontario.ca
1799:2024-06-27
1775:2010-02-28
1686:2021-11-11
1284:2014-10-17
1198:References
1070:americanum
1061:americanum
1037:americanum
939:climates.
885:New Jersey
881:introduced
827:along the
818:Hispaniola
816:island of
806:Nuevo LeĂłn
695:tetraploid
691:americanum
652:var.
646:Hispaniola
626:americanum
614:var.
485:sporophyte
470:prothallus
171:Suborder:
141:Division:
95:americanum
2656:Asplenium
2339:233500198
2167:201197385
1522:0024-4066
1098:cultivars
912:japonicum
873:Tennessee
814:Caribbean
699:Old World
664:japonicum
572:Phyllitis
558:Phyllitis
550:Phyllitis
546:Phyllitis
542:Phyllitis
534:Asplenium
529:'s legs.
401:evergreen
333:americana
305:americana
203:Species:
196:Asplenium
118:Kingdom:
2594:VASCAN:
2577:26603800
2572:Tropicos
2463:2.129927
2414:10886824
2248:Q1130272
2242:Wikidata
2203:variety.
2159:34618950
2119:(1753).
1954:16 March
1929:16 March
1901:16 March
1875:16 March
1823:16 March
1695:cite web
1617:16 March
1591:16 March
1557:16 March
1440:16 March
1345:16 March
1246:16 March
1221:16 March
849:Michigan
802:Coahuila
782:Maryland
736:and the
734:Caucasus
654:lindenii
527:myriapod
499:binomial
495:Linnaeus
490:Taxonomy
431:red deer
399:, is an
272:KĂŒmmerle
242:Synonyms
181:Family:
93:) (var.
2445:MoBotPF
2362:2650669
1150:Habitat
1137:Gallery
1000:Albania
926:Habitat
910:subsp.
857:Alabama
798:Chiapas
778:Ontario
754:Tunisia
750:Algeria
746:Morocco
707:diploid
662:subsp.
630:Ontario
624:subsp.
504:in his
406:in the
336:Fernald
303:subsp.
288:Ă. Löve
259:subsp.
191:Genus:
161:Order:
151:Class:
122:Plantae
89: (
69: (
2623:426033
2587:601691
2584:uBio:
2538:ASSC11
2533:PLANTS
2499:905814
2483:NZOR:
2450:285846
2427:192155
2388:129105
2349:ASPSCO
2346:FoIO:
2326:150133
2300:594778
2274:130489
2271:APDB:
2188:
2165:
2157:
2057:
2035:
1643:
1520:
1318:
1126:spleen
1051:Canada
1012:Sweden
1004:Norway
967:Europe
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