55:
1065:
1077:
33:
239:, which is a diagnostic feature. Classification in the field is based on morphological characteristics, with considerable variations in colouration between insects of the same species (although for example,
259:
sucking mouthparts, they pierce plant tissues and cause damage ranging from leaf tattering and fruit blemishes, to complete death of shoots, branches or whole plants.
1228:
1254:
1202:
1241:
1002:
Stonedahl GM (1991) Oriental species of
Helopeltis (Heteroptera: Miridae): a review of economic literature and guide to identification.
1092:
1246:
977:
1069:
1163:
1150:
54:
1259:
236:
592:
622:
772:
402:
396:
1282:
852:
792:
732:
692:
682:
612:
482:
153:
862:
822:
802:
782:
752:
702:
572:
562:
1103:
890:
842:
722:
632:
532:
502:
492:
452:
1168:
1081:
832:
812:
762:
742:
712:
652:
642:
552:
542:
522:
442:
394:
by
Schmitz (1968). These species usually have a lesser pests status than their Asian counterparts, with
371:
582:
672:
662:
602:
512:
472:
294:
1287:
1031:
Signoret (Heteroptera, Miridae) avec un exposedes problemes relatifes aux structures genitales',
462:
49:
41:
1233:
1176:
433:
267:
1047:
341:
1141:
417:
307:
271:
207:
203:
178:
1087:
1276:
421:
283:
1181:
1155:
32:
1126:
1215:
1135:
962:
951:
914:
901:
407:
311:
191:
116:
1064:
412:
298:
263:
1189:
328:
320:
256:
199:
136:
106:
86:
66:
1076:
1120:
942:
356:
324:
223:
1207:
918:
365:
195:
126:
1220:
215:
211:
96:
76:
1097:
1194:
927:
337:
275:
286:
are important 'new encounter' pests of SE Asian cocoa including:
380:
1101:
1027:
Schmitz G (1968) 'Monographie des especes
Africaines du genre
219:
1018:
Entwistle PF (1972) Pests of Cocoa, Longmans, London, 779 pp
406:
causing greatest cocoa tree and crop damage in
Central and
1110:
390:, are the African species which were placed into
235:Mosquito bugs have a characteristic spine on the
386:Other cocoa Mirid pest species, very similar to
880:Entwistle (1972) lists and maps ten species of
8:
1098:
226:, or placed in its own genus. A number of
31:
20:
961:: widespread, but only found on cocoa in
970:: the most widespread in tropical Africa
995:
985:: widespread in west and central Africa
7:
379:as a sub-species): India through to
1004:Bulletin of Entomological Research
14:
900:: widely distributed east of the
1075:
1063:
935:: humid forest in central Africa
243:is characteristically green and
53:
420:, are similarly minor pests of
416:species, belonging to the same
1:
232:species are pests in Africa.
206:of various crops, including
168:
1050:(searched 20 December 2023)
926:: Ivory Coast and probably
1304:
186:, also sometimes known as
1088:DropData cocoa pest guide
1033:Ann. Mus. Roy. Afr. Cent.
593:Helopeltis fasciaticollis
266:along the islands of the
166:
161:
50:Scientific classification
48:
39:
30:
23:
950:: north and east of the
623:Helopeltis hyalospilosus
436:currently (2023) lists:
198:(capsid bugs) and tribe
773:Helopeltis pseudomaynei
737:Stonedahl et al.2, 1995
403:Distantiella theobromae
397:Sahlbergella singularis
262:There has been evident
251:Damage and distribution
853:Helopeltis waterhousei
793:Helopeltis schoutedeni
733:Helopeltis pernicialis
693:Helopeltis melanescens
683:Helopeltis mayumbensis
613:Helopeltis ghesquierei
483:Helopeltis basilewskyi
978:São Tomé and Príncipe
863:Helopeltis westwoodii
823:Helopeltis sumatranus
803:Helopeltis seredensis
783:Helopeltis rauwolfiae
753:Helopeltis podagricus
703:Helopeltis obscuratus
573:Helopeltis couturieri
563:Helopeltis corbisieri
222:. Now in a different
1084:at Wikimedia Commons
843:Helopeltis villiersi
723:Helopeltis pellucida
633:Helopeltis insularis
533:Helopeltis cinchonae
503:Helopeltis bergrothi
493:Helopeltis bergevini
453:Helopeltis anacardii
833:Helopeltis theivora
813:Helopeltis sulawesi
763:Helopeltis poppiusi
743:Helopeltis plebejus
713:Helopeltis orophila
653:Helopeltis lalandei
643:Helopeltis labaumei
553:Helopeltis collaris
543:Helopeltis clavifer
523:Helopeltis carayoni
443:Helopeltis alluaudi
583:Helopeltis cuneata
463:Helopeltis antonii
42:Helopeltis antonii
16:Genus of true bugs
1270:
1269:
1104:Taxon identifiers
1080:Media related to
1048:Catalogue of life
884:attacking cocoa:
673:Helopeltis maynei
663:Helopeltis lemosi
603:Helopeltis gerini
513:Helopeltis bradyi
473:Helopeltis bakeri
467:V. Signoret, 1858
434:Catalogue of life
270:and there may be
268:Malay Archipelago
175:
174:
157:
1295:
1263:
1262:
1250:
1249:
1237:
1236:
1224:
1223:
1211:
1210:
1198:
1197:
1185:
1184:
1172:
1171:
1159:
1158:
1146:
1145:
1144:
1131:
1130:
1129:
1099:
1079:
1068:Data related to
1067:
1051:
1045:
1039:
1025:
1019:
1016:
1010:
1000:
913:): very common,
868:
858:
848:
838:
837:Waterhouse, 1886
828:
818:
808:
798:
788:
787:Ghesquiere, 1948
778:
768:
758:
757:(A. Costa, 1864)
748:
738:
728:
718:
717:Ghesquiere, 1939
708:
698:
688:
687:Ghesquiere, 1922
678:
677:Ghesquiere, 1922
668:
667:Ghesquiere, 1922
658:
648:
638:
628:
618:
608:
598:
588:
578:
568:
558:
548:
538:
528:
518:
517:Waterhouse, 1886
508:
498:
488:
478:
468:
458:
448:
342:Papua New Guinea
190:, is a group of
152:
58:
57:
35:
21:
1303:
1302:
1298:
1297:
1296:
1294:
1293:
1292:
1273:
1272:
1271:
1266:
1258:
1253:
1245:
1240:
1232:
1227:
1219:
1214:
1206:
1201:
1193:
1188:
1180:
1175:
1167:
1162:
1154:
1149:
1140:
1139:
1134:
1125:
1124:
1119:
1106:
1060:
1055:
1054:
1046:
1042:
1026:
1022:
1017:
1013:
1001:
997:
992:
878:
871:
866:
856:
846:
836:
826:
817:Stonedahl, 1991
816:
806:
796:
786:
776:
766:
756:
746:
736:
726:
716:
706:
696:
686:
676:
666:
656:
646:
636:
626:
616:
606:
596:
586:
576:
566:
556:
546:
536:
526:
516:
506:
496:
486:
476:
466:
456:
446:
430:
308:Malay Peninsula
272:cryptic species
253:
202:. They include
151:
52:
17:
12:
11:
5:
1301:
1299:
1291:
1290:
1285:
1283:Miridae genera
1275:
1274:
1268:
1267:
1265:
1264:
1251:
1238:
1225:
1212:
1199:
1186:
1173:
1160:
1147:
1132:
1116:
1114:
1108:
1107:
1102:
1096:
1095:
1090:
1085:
1073:
1072:at Wikispecies
1059:
1058:External links
1056:
1053:
1052:
1040:
1020:
1011:
994:
993:
991:
988:
987:
986:
980:
971:
968:A. schoutedeni
965:
956:
955:
954:
948:A. mayumbensis
945:
936:
930:
924:A. serendensis
921:
904:
877:
872:
870:
869:
859:
857:Kirkaldy, 1902
849:
847:Delattre, 1947
839:
829:
819:
809:
799:
789:
779:
769:
759:
749:
739:
729:
719:
709:
699:
689:
679:
669:
659:
649:
639:
637:Kirkaldy, 1902
629:
619:
609:
599:
589:
579:
569:
559:
549:
547:(Walker, 1871)
539:
529:
519:
509:
499:
489:
479:
469:
459:
449:
438:
429:
426:
422:Latin American
384:
383:
368:
359:
350:
344:
331:
314:
301:
252:
249:
194:in the family
173:
172:
164:
163:
159:
158:
144:
140:
139:
134:
130:
129:
124:
120:
119:
114:
110:
109:
104:
100:
99:
94:
90:
89:
84:
80:
79:
74:
70:
69:
64:
60:
59:
46:
45:
37:
36:
28:
27:
15:
13:
10:
9:
6:
4:
3:
2:
1300:
1289:
1286:
1284:
1281:
1280:
1278:
1261:
1256:
1252:
1248:
1243:
1239:
1235:
1230:
1226:
1222:
1217:
1213:
1209:
1204:
1200:
1196:
1191:
1187:
1183:
1178:
1174:
1170:
1165:
1161:
1157:
1152:
1148:
1143:
1137:
1133:
1128:
1122:
1118:
1117:
1115:
1113:
1109:
1105:
1100:
1094:
1093:PestInfo wiki
1091:
1089:
1086:
1083:
1078:
1074:
1071:
1066:
1062:
1061:
1057:
1049:
1044:
1041:
1037:
1034:
1030:
1024:
1021:
1015:
1012:
1008:
1005:
999:
996:
989:
984:
981:
979:
975:
972:
969:
966:
964:
960:
957:
953:
949:
946:
944:
940:
937:
934:
933:A. corbisieri
931:
929:
925:
922:
920:
916:
912:
908:
905:
903:
899:
896:
895:
893:
892:
887:
886:
885:
883:
876:
873:
867:(White, 1842)
865:
864:
860:
855:
854:
850:
845:
844:
840:
835:
834:
830:
825:
824:
820:
815:
814:
810:
807:Schmitz, 1968
805:
804:
800:
795:
794:
790:
785:
784:
780:
777:Schmitz, 1968
775:
774:
770:
767:Schmitz, 1968
765:
764:
760:
755:
754:
750:
747:Poppius, 1911
745:
744:
740:
735:
734:
730:
725:
724:
720:
715:
714:
710:
707:Poppius, 1915
705:
704:
700:
697:Schmitz, 1988
695:
694:
690:
685:
684:
680:
675:
674:
670:
665:
664:
660:
657:Carayon, 1949
655:
654:
650:
647:Poppius, 1911
645:
644:
640:
635:
634:
630:
627:Schmitz, 1988
625:
624:
620:
617:Schmitz, 1968
615:
614:
610:
607:Carayon, 1949
605:
604:
600:
597:Poppius, 1915
595:
594:
590:
587:Distant, 1903
585:
584:
580:
577:Schmitz, 1988
575:
574:
570:
567:Schmitz, 1968
565:
564:
560:
555:
554:
550:
545:
544:
540:
535:
534:
530:
527:Schmitz, 1968
525:
524:
520:
515:
514:
510:
505:
504:
500:
497:Poppius, 1914
495:
494:
490:
487:Schmitz, 1968
485:
484:
480:
477:Poppius, 1915
475:
474:
470:
465:
464:
460:
455:
454:
450:
445:
444:
440:
439:
437:
435:
427:
425:
423:
419:
415:
414:
409:
405:
404:
399:
398:
393:
389:
382:
378:
377:H. theobromae
374:
373:
369:
367:
363:
362:H. sumatranus
360:
358:
354:
351:
349:: Philippines
348:
345:
343:
339:
335:
332:
330:
326:
322:
318:
315:
313:
309:
305:
302:
300:
296:
292:
289:
288:
287:
285:
284:sensu stricto
281:
277:
273:
269:
265:
260:
258:
255:With typical
250:
248:
246:
242:
238:
233:
231:
230:
225:
221:
217:
213:
209:
205:
201:
197:
193:
192:heteropterans
189:
188:mosquito bugs
185:
184:
180:
171:
170:
165:
160:
155:
150:
149:
145:
142:
141:
138:
135:
132:
131:
128:
125:
122:
121:
118:
115:
112:
111:
108:
105:
102:
101:
98:
95:
92:
91:
88:
85:
82:
81:
78:
75:
72:
71:
68:
65:
62:
61:
56:
51:
47:
44:
43:
38:
34:
29:
26:
22:
19:
1111:
1043:
1035:
1032:
1028:
1023:
1014:
1006:
1003:
998:
983:A. westwoodi
982:
973:
967:
958:
947:
938:
932:
923:
911:H. bergevini
910:
906:
898:A. bergrothi
897:
891:A. bergrothi
889:
881:
879:
874:
861:
851:
841:
831:
827:Roepke, 1916
821:
811:
801:
797:Reuter, 1906
791:
781:
771:
761:
751:
741:
731:
721:
711:
701:
691:
681:
671:
661:
651:
641:
631:
621:
611:
601:
591:
581:
571:
561:
551:
541:
531:
521:
511:
507:Reuter, 1892
501:
491:
481:
471:
461:
457:Miller, 1954
451:
447:Reuter, 1905
441:
431:
411:
401:
395:
391:
387:
385:
376:
370:
361:
352:
346:
333:
316:
303:
290:
279:
261:
254:
247:red-brown).
244:
240:
234:
228:
227:
187:
182:
181:
176:
167:
147:
146:
40:
24:
18:
1216:iNaturalist
1136:Wikispecies
963:Ivory Coast
959:A. poppiusi
952:Congo River
941:: southern
915:Ivory Coast
907:A. lalendei
902:Niger River
408:West Africa
372:H. theivora
353:H. sulawesi
347:H. collaris
334:H. clavifer
312:Philippines
297:through to
241:H. theivora
117:Heteroptera
1277:Categories
1156:Helopeltis
1142:Helopeltis
1112:Helopeltis
1082:Helopeltis
1070:Helopeltis
1029:Helopeltis
990:References
882:Afropeltis
875:Afropeltis
727:Stal, 1871
557:Stal, 1871
537:Mann, 1907
413:Monalonion
392:Afropeltis
388:Helopeltis
299:West Irian
291:H. antonii
280:Helopeltis
264:speciation
257:Hemipteran
245:H. antonii
229:Afropeltis
183:Helopeltis
148:Helopeltis
113:Suborder:
87:Arthropoda
25:Helopeltis
1288:Dicyphini
1127:Q16982956
1038:: 247 pp.
1009:: 465–490
974:A. lemosi
939:A. gerini
329:Indonesia
321:Sri Lanka
317:H. bradyi
304:H. bakeri
237:scutellum
200:Dicyphini
137:Dicyphini
107:Hemiptera
73:Kingdom:
67:Eukaryota
1121:Wikidata
943:Cameroun
357:Sulawesi
325:Malaysia
274:in this
224:subgenus
169:see text
162:Species
154:Signoret
123:Family:
83:Phylum:
77:Animalia
63:Domain:
1234:1111292
1208:3265690
919:Nigeria
894:group:
428:Species
424:cocoa.
366:Sumatra
196:Miridae
143:Genus:
133:Tribe:
127:Miridae
103:Order:
97:Insecta
93:Class:
1260:237374
1247:113163
1221:474380
1195:1HELOG
1169:190058
375:(with
216:cotton
212:cashew
156:, 1858
1229:IRMNG
1182:62VFS
928:Ghana
418:tribe
338:Sabah
295:India
282:spp.
276:genus
208:cacao
204:pests
179:genus
1255:NCBI
1242:ITIS
1203:GBIF
1190:EPPO
1164:BOLD
888:the
432:The
400:and
381:Java
340:and
310:and
218:and
177:The
1177:CoL
1151:AFD
1036:168
917:to
909:(=
220:tea
1279::
1257::
1244::
1231::
1218::
1205::
1192::
1179::
1166::
1153::
1138::
1123::
1007:81
976::
410:.
364::
355::
336::
327:,
323:,
319::
306::
293::
278:.
214:,
210:,
Text is available under the Creative Commons Attribution-ShareAlike License. Additional terms may apply.