241:
209:
248:
800:
Fortuny, Josep; Arbez, Thomas; Mujal, Eudald; Steyer, J. Sébastien (2019-01-02). "Reappraisal of 'Metoposaurus hoffmani' Dutuit, 1978, and description of new temnospondyl specimens from the Middle–Late
Triassic of Madagascar (Morondava Basin)".
1396:
Flynn, John J.; Parrish, J. Michael; Rakotosamimanana, Berthe; Ranivoharimanana, Lovasoa; Simpson, William F.; Wyss, André R. (2000-09-25). "New
Traversodontids (Synapsida: Eucynodontia) from the Triassic of Madagascar".
1267:"The earliest-diverging avemetatarsalian: a new osteoderm-bearing taxon from the Triassic (?Earliest Late Triassic) of Madagascar and the composition of avemetatarsalian assemblages prior to the radiation of dinosaurs"
1334:
Kammerer, Christian F.; Flynn, John J.; Ranivoharimanana, Lovasoa; Wyss, André R. (2010-12-02). "The first record of a probainognathian (Cynodontia: Chiniquodontidae) from the
Triassic of Madagascar".
283:
It is described as "thick beds of mottled red or green clays associated with soft cross-bedded sandstones, light in colour and much finer-grained than the Isalo I sandstones." It is prominent in the
887:
Flynn, John J.; Parrish, J. Michael; Rakotosamimanana, Berthe; Simpson, William F.; Whatley, Robin L.; Wyss, André R. (1999-10-22). "A Triassic Fauna from
Madagascar, Including Early Dinosaurs".
1302:
1236:
1606:
1506:
Kammerer, Christian F.; Flynn, John J.; Ranivoharimanana, Lovasoa; Wyss, André R. (October 2012). "Ontogeny in the
Malagasy Traversodontid
1265:
Nesbitt, Sterling J.; Patellos, Emily; Kammerer, Christian F.; Ranivoharimanana, Lovasoa; Wyss, Andre´ R.; Flynn, John J. (25 July 2023).
1301:
Patellos, Emily A.; Nesbitt, Sterling J.; Kammerer, Christian F.; Wyss, Andre; Flynn, John J.; Ranivoharimanana, Lovasoa (October 2019).
1641:
1611:
859:"Paleogeographic and biostratigraphic implications of new early Mesozoic vertebrates from Poamay, central Morondava Basin, Madagascar"
1310:
1244:
939:
1162:"A tiny ornithodiran archosaur from the Triassic of Madagascar and the role of miniaturization in dinosaur and pterosaur ancestry"
240:
1237:"A new lagerpetid archosaur from the Triassic of Madagascar and the importance of miniaturization in ornithodiran evolution"
1631:
1303:"A new reptile from the ?Middle Triassic of Madagascar may represent the earliest-diverging Avemetatarsalia (Archosauria)"
413:
1636:
1616:
111:
1112:"Studies on continental Late Triassic tetrapod biochronology. II. The Ischigualastian and a Carnian global correlation"
1160:
Kammerer, Christian F.; Nesbitt, Sterling J.; Flynn, John J.; Ranivoharimanana, Lovasoa; Wyss, André R. (2020-07-02).
1235:
Kammerer, Christian F.; Nesbitt, Sterling J.; Flynn, John J.; Ranivoharimanana, Lovasoa; Wyss, Andre (October 2019).
1111:
1553:
Kammerer, Christian F.; Flynn, John J.; Ranivoharimanana, Lovasoa; Wyss, André R. (2008-06-12). "New material of
1440:
Ranivoharimanana, Lovasoa; Kammerer, Christian F.; Flynn, John J.; Wyss, André R. (2011-11-01). "New material of
1410:
726:(?Middle to Upper Triassic, Isalo Group, Madagascar) and its systematic position among stem archosaur reptiles"
1570:
1026:
961:
Flynn, John J.; Nesbitt, Sterling J.; Parrish, J. Michael; Ranivoharimanana, Lovasoa; Wyss, André R. (2010).
1621:
967:(Diapsida: Archosauromorpha) from the Triassic Isalo Group of southwestern Madagascar: cranium and mandible"
435:
1626:
157:
118:
495:
80:
1453:
1343:
1173:
1126:
1045:
978:
810:
720:
Nesbitt, S.J.; Flynn, J.J.; Pritchard, A.C.; Parrish, M.J.; Ranivoharimanana, L.; Wyss, A.R. (2015).
618:
228:
45:
65:
1582:
1535:
1485:
1477:
1422:
1375:
1367:
1217:
1092:
1004:
938:
Flynn, John; Nesbitt, Sterling; Parrish, Michael; Ranivoharimanana, Lovasoa; Wyss, Andre (2008).
912:
834:
763:
660:
192:
55:
1574:
1527:
1469:
1414:
1359:
1209:
1191:
1142:
1061:
996:
920:
904:
858:
826:
755:
680:
1566:
1519:
1461:
1406:
1351:
1278:
1199:
1181:
1134:
1053:
986:
896:
818:
780:
Brenon, P. (1972). The
Geology of Madagascar. In: Battistini, R., Richard-Vindard, G. (eds)
745:
737:
669:
464:
398:
196:
649:
523:
477:
426:
224:
1457:
1347:
1177:
1130:
1049:
982:
814:
1204:
1161:
690:
652:
639:
628:
583:
563:
423:
353:
321:
150:
1057:
1600:
1586:
1489:
1426:
1221:
991:
962:
838:
721:
560:
533:
455:
450:
363:
328:
27:
1539:
1379:
1008:
767:
857:
Burmeister, Kurtis C.; Flynn, John J.; Parrish, J. Michael; Wyss, Andre R. (2006).
571:
553:
543:
344:
335:
284:
69:
822:
1465:
1355:
1283:
1266:
1138:
900:
741:
536:
503:
485:
445:
1523:
1025:
Langer, Max; Boniface, Michael; Cuny, Gilles; Barbieri, Laurent (2000-04-01).
785:
672:
580:
474:
277:
214:
1578:
1531:
1473:
1418:
1363:
1195:
1146:
1065:
1000:
940:"A new species of basal archosauromorph from the Late Triassic of Madagascar"
908:
830:
759:
172:
159:
133:
120:
1186:
1079:
Whatley, R.L.; Flynn, J.J.; Parrish, J.M.; Simpson, W.; Wyss, A.R. (1999). "
556:
95:
1307:
Society of
Vertebrate Paleontology Abstracts of Papers, 79th Annual Meeting
1241:
Society of
Vertebrate Paleontology Abstracts of Papers, 79th Annual Meeting
1213:
924:
631:
514:
273:
99:
31:
1481:
1371:
407:
Various indeterminate fragments, some of which may belong to dinosaurs.
1096:
916:
35:
750:
1083:
revisited: two rhynchosaur skeletons from southwestern
Madagascar".
342:
metoposaurids. Some fossils were the basis for the dubious species "
339:
1557:(Cynodontia: Traversodontidae) from the Triassic of Madagascar".
1444:(Cynodontia, Traversodontidae) from the Triassic of Madagascar".
441:
Numerous fossils representing parts of the skull and postcrania
429:, originally mistakenly identified as a "prosauropod" dinosaur.
404:
Teeth, a coracoid, a centrum fragment, and cranial fragments
1411:
10.1671/0272-4634(2000)020[0422:NTSEFT]2.0.CO;2
863:
New Mexico Museum of
Natural History & Science Bulletin
1510:
and a Reconsideration of its Phylogenetic Relationships".
1571:
10.1671/0272-4634(2008)28[445:NMOMBC]2.0.CO;2
784:. Monographiae Biologicae, vol 21. Springer, Dordrecht.
595:
458:
fossils are most likely juvenile rhynchosaurs instead.
376:
299:
882:
880:
878:
876:
220:
202:
188:
149:
110:
105:
91:
86:
76:
61:
51:
41:
21:
1329:
1327:
1296:
1294:
730:Bulletin of the American Museum of Natural History
1391:
1389:
1166:Proceedings of the National Academy of Sciences
1020:
1018:
16:Informal Triassic geological unit in Madagascar
1501:
1499:
1031:, a Late Triassic rhynchosaur from Madagascar"
795:
793:
715:
713:
711:
709:
8:
786:https://doi.org/10.1007/978-94-015-7159-3_2
18:
1282:
1271:Zoological Journal of the Linnean Society
1203:
1185:
990:
749:
454:. Possibly represents two taxa. Reported
1119:Journal of South American Earth Sciences
598:
419:Numerous skulls and postcranial fossils
379:
302:
705:
686:Skulls, jaws, and postcranial material
782:Biogeography and Ecology in Madagascar
366:remains. May be from Isalo II or III.
852:
850:
848:
7:
470:A maxilla and postcranial material
331:, skull fragments, and intercentra
1559:Journal of Vertebrate Paleontology
1446:Journal of Vertebrate Paleontology
1399:Journal of Vertebrate Paleontology
1336:Journal of Vertebrate Paleontology
1085:Journal of Vertebrate Paleontology
944:Journal of Vertebrate Paleontology
803:Journal of Vertebrate Paleontology
448:sometimes considered a species of
14:
1607:Geologic formations of Madagascar
1512:Fieldiana Life and Earth Sciences
645:Several skulls and jaw fragments
529:A jaw fragment and other remains
992:10.1111/j.1475-4983.2010.00954.x
348:". May be from Isalo II or III.
247:
246:
239:
207:
1027:"The phylogenetic position of
1:
1058:10.1016/S0753-3969(00)80002-6
823:10.1080/02724634.2019.1576701
724:Azendohsaurus madagaskarensis
414:Azendohsaurus madagaskarensis
1466:10.1080/02724634.2011.618154
1356:10.1080/02724634.2010.520784
1139:10.1016/j.jsames.2005.04.002
1110:Langer, Max Cardoso (2005).
901:10.1126/science.286.5440.763
338:remains similar to those of
151:Approximate paleocoordinates
742:10.1206/amnb-899-00-1-126.1
1658:
1642:Paleontology in Madagascar
1284:10.1093/zoolinnean/zlad038
722:"Postcranial osteology of
1612:Triassic System of Africa
1524:10.3158/2158-5520-5.1.112
234:
26:
1038:Annales de Paléontologie
1187:10.1073/pnas.1916631117
1029:Isalorhynchus genovefae
436:Isalorhynchus genovefae
486:Mambachiton fiandohana
1309:: 167. Archived from
1243:: 127. Archived from
345:Metoposaurus hoffmani
255:Isalo II (Madagascar)
1632:Sandstone formations
1277:(zlad038): 327–353.
619:Chiniquodon kalanoro
559:, though similar to
552:Likely referable to
268:, also known as the
46:Geological formation
1637:Mudstone formations
1617:Triassic Madagascar
1458:2011JVPal..31.1292R
1348:2010JVPal..30.1889K
1178:2020PNAS..11717932K
1172:(30): 17932–17936.
1131:2005JSAES..19..219L
1050:2000AnPal..86..101L
983:2010Palgy..53..669F
815:2019JVPal..39E6701F
491:A partial skeleton
276:geological unit in
169: /
130: /
66:Isalo III Formation
28:Stratigraphic range
963:"A new species of
661:Kannemeyeriiformes
22:Isalo II Formation
1555:Menadon besairiei
895:(5440): 763–765.
697:
696:
681:Menadon besairiei
589:
588:
498:avemetatarsalian
370:
369:
272:, is an informal
263:
262:
197:Toliara Provinces
81:Isalo I Formation
1649:
1591:
1590:
1550:
1544:
1543:
1503:
1494:
1493:
1452:(6): 1292–1302.
1437:
1431:
1430:
1393:
1384:
1383:
1342:(6): 1889–1894.
1331:
1322:
1321:
1319:
1318:
1298:
1289:
1288:
1286:
1262:
1256:
1255:
1253:
1252:
1232:
1226:
1225:
1207:
1189:
1157:
1151:
1150:
1116:
1107:
1101:
1100:
1076:
1070:
1069:
1035:
1022:
1013:
1012:
994:
958:
952:
951:
935:
929:
928:
884:
871:
870:
854:
843:
842:
797:
788:
778:
772:
771:
753:
717:
670:kannemeyeriiform
596:
478:avemetatarsalian
465:Kongonaphon kely
399:Archosauriformes
377:
300:
250:
249:
243:
213:
211:
210:
184:
183:
181:
180:
179:
174:
170:
167:
166:
165:
162:
145:
144:
142:
141:
140:
135:
131:
128:
127:
126:
123:
19:
1657:
1656:
1652:
1651:
1650:
1648:
1647:
1646:
1597:
1596:
1595:
1594:
1552:
1551:
1547:
1505:
1504:
1497:
1439:
1438:
1434:
1395:
1394:
1387:
1333:
1332:
1325:
1316:
1314:
1300:
1299:
1292:
1264:
1263:
1259:
1250:
1248:
1234:
1233:
1229:
1159:
1158:
1154:
1114:
1109:
1108:
1104:
1078:
1077:
1073:
1033:
1024:
1023:
1016:
960:
959:
955:
937:
936:
932:
886:
885:
874:
856:
855:
846:
809:(1): e1576701.
799:
798:
791:
779:
775:
719:
718:
707:
702:
650:massetognathine
594:
564:crocodylomorphs
524:Procolophonidae
427:archosauromorph
424:allokotosaurian
375:
359:Interclavicles
298:
293:
270:Makay Formation
259:
258:
257:
256:
253:
252:
251:
229:Mahajanga Basin
225:Morondava Basin
208:
206:
177:
175:
171:
168:
163:
160:
158:
156:
155:
138:
136:
132:
129:
124:
121:
119:
117:
116:
17:
12:
11:
5:
1655:
1653:
1645:
1644:
1639:
1634:
1629:
1624:
1622:Ladinian Stage
1619:
1614:
1609:
1599:
1598:
1593:
1592:
1565:(2): 445–462.
1545:
1518:(5): 112–125.
1508:Dadadon isaloi
1495:
1442:Dadadon isaloi
1432:
1405:(3): 422–427.
1385:
1323:
1290:
1257:
1227:
1152:
1125:(2): 219–239.
1102:
1071:
1044:(2): 101–127.
1014:
977:(3): 669–688.
953:
930:
872:
844:
789:
773:
704:
703:
701:
698:
695:
694:
691:traversodontid
687:
684:
676:
675:
666:
664:
657:
656:
653:traversodontid
646:
643:
640:Dadadon isaloi
635:
634:
629:chiniquodontid
625:
624:Part of a jaw
622:
614:
613:
610:
607:
603:
602:
593:
590:
587:
586:
584:dinosauromorph
577:
575:
568:
567:
550:
547:
540:
539:
530:
527:
520:
519:
510:
507:
500:
499:
492:
489:
481:
480:
471:
468:
460:
459:
442:
439:
431:
430:
420:
417:
409:
408:
405:
402:
395:
394:
391:
388:
384:
383:
374:
371:
368:
367:
362:Indeterminate
360:
357:
354:Stereospondyli
350:
349:
334:Indeterminate
332:
325:
322:Metoposauridae
318:
317:
314:
311:
307:
306:
297:
294:
292:
291:Fossil content
289:
261:
260:
254:
245:
244:
238:
237:
236:
235:
232:
231:
222:
218:
217:
204:
200:
199:
190:
186:
185:
153:
147:
146:
114:
108:
107:
103:
102:
93:
89:
88:
84:
83:
78:
74:
73:
63:
59:
58:
53:
49:
48:
43:
39:
38:
24:
23:
15:
13:
10:
9:
6:
4:
3:
2:
1654:
1643:
1640:
1638:
1635:
1633:
1630:
1628:
1627:Carnian Stage
1625:
1623:
1620:
1618:
1615:
1613:
1610:
1608:
1605:
1604:
1602:
1588:
1584:
1580:
1576:
1572:
1568:
1564:
1560:
1556:
1549:
1546:
1541:
1537:
1533:
1529:
1525:
1521:
1517:
1513:
1509:
1502:
1500:
1496:
1491:
1487:
1483:
1479:
1475:
1471:
1467:
1463:
1459:
1455:
1451:
1447:
1443:
1436:
1433:
1428:
1424:
1420:
1416:
1412:
1408:
1404:
1400:
1392:
1390:
1386:
1381:
1377:
1373:
1369:
1365:
1361:
1357:
1353:
1349:
1345:
1341:
1337:
1330:
1328:
1324:
1313:on 2020-12-15
1312:
1308:
1304:
1297:
1295:
1291:
1285:
1280:
1276:
1272:
1268:
1261:
1258:
1247:on 2020-12-15
1246:
1242:
1238:
1231:
1228:
1223:
1219:
1215:
1211:
1206:
1201:
1197:
1193:
1188:
1183:
1179:
1175:
1171:
1167:
1163:
1156:
1153:
1148:
1144:
1140:
1136:
1132:
1128:
1124:
1120:
1113:
1106:
1103:
1098:
1094:
1091:(sup3): 84A.
1090:
1086:
1082:
1081:Isalorhynchus
1075:
1072:
1067:
1063:
1059:
1055:
1051:
1047:
1043:
1039:
1032:
1030:
1021:
1019:
1015:
1010:
1006:
1002:
998:
993:
988:
984:
980:
976:
972:
971:Palaeontology
968:
966:
965:Azendohsaurus
957:
954:
949:
945:
941:
934:
931:
926:
922:
918:
914:
910:
906:
902:
898:
894:
890:
883:
881:
879:
877:
873:
868:
864:
860:
853:
851:
849:
845:
840:
836:
832:
828:
824:
820:
816:
812:
808:
804:
796:
794:
790:
787:
783:
777:
774:
769:
765:
761:
757:
752:
747:
743:
739:
735:
731:
727:
725:
716:
714:
712:
710:
706:
699:
692:
688:
685:
683:
682:
678:
677:
674:
671:
667:
665:
662:
659:
658:
654:
651:
647:
644:
642:
641:
637:
636:
633:
630:
626:
623:
621:
620:
616:
615:
611:
608:
605:
604:
601:
597:
591:
585:
582:
578:
576:
573:
570:
569:
565:
562:
561:goniopholidid
558:
555:
551:
548:
545:
542:
541:
538:
535:
534:procolophonid
531:
528:
525:
522:
521:
517:
516:
511:
508:
505:
502:
501:
497:
493:
490:
488:
487:
483:
482:
479:
476:
472:
469:
467:
466:
462:
461:
457:
456:sphenodontian
453:
452:
451:Hyperodapedon
447:
443:
440:
438:
437:
433:
432:
428:
425:
421:
418:
416:
415:
411:
410:
406:
403:
400:
397:
396:
392:
389:
386:
385:
382:
378:
372:
365:
364:stereospondyl
361:
358:
355:
352:
351:
347:
346:
341:
337:
333:
330:
329:interclavicle
326:
323:
320:
319:
315:
312:
309:
308:
305:
301:
295:
290:
288:
286:
281:
279:
275:
271:
267:
242:
233:
230:
226:
223:
219:
216:
205:
201:
198:
194:
191:
187:
182:
173:23.1°S 24.7°E
154:
152:
148:
143:
134:20.3°S 45.4°E
115:
113:
109:
104:
101:
97:
94:
90:
85:
82:
79:
75:
71:
67:
64:
60:
57:
54:
50:
47:
44:
40:
37:
33:
29:
25:
20:
1562:
1558:
1554:
1548:
1515:
1511:
1507:
1449:
1445:
1441:
1435:
1402:
1398:
1339:
1335:
1315:. Retrieved
1311:the original
1306:
1274:
1270:
1260:
1249:. Retrieved
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70:unconformity
668:An unnamed
579:An unnamed
549:Osteoderms
537:parareptile
532:An unnamed
512:Similar to
504:Phytosauria
494:An armored
446:rhynchosaur
178:-23.1; 24.7
176: /
139:-20.3; 45.4
137: /
112:Coordinates
56:Isalo Group
1601:Categories
1317:2020-06-27
1251:2020-06-27
869:: 457–475.
700:References
673:dicynodont
581:silesaurid
475:lagerpetid
304:Amphibians
296:Amphibians
278:Madagascar
215:Madagascar
1587:140720709
1579:0272-4634
1532:2158-5520
1490:220437281
1474:0272-4634
1427:130041740
1419:0272-4634
1364:0272-4634
1222:220386589
1196:0027-8424
1147:0895-9811
1066:0753-3969
1001:1475-4983
909:0036-8075
839:146075668
831:0272-4634
760:0003-0090
751:2246/6624
736:: 1–126.
693:cynodont
655:cynodont
609:Material
600:Synapsids
592:Synapsids
566:as well.
557:aetosaurs
390:Material
381:Synapsids
313:Material
193:Mahajanga
96:Sandstone
87:Lithology
62:Underlies
1540:84651546
1482:41407685
1380:85868922
1372:25790814
1214:32631980
1009:82341339
925:10531059
768:86289421
632:cynodont
515:Rutiodon
373:Reptiles
274:Triassic
266:Isalo II
106:Location
100:mudstone
77:Overlies
32:Ladinian
1454:Bibcode
1344:Bibcode
1205:7395432
1174:Bibcode
1127:Bibcode
1097:4524027
1046:Bibcode
979:Bibcode
917:2899382
889:Science
811:Bibcode
546:indet.
518:teeth.
506:indet.
473:A tiny
356:indet.
324:indet.
203:Country
164:24°42′E
161:23°06′S
125:45°24′E
122:20°18′S
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509:Teeth
393:Notes
387:Taxon
340:Indian
316:Notes
310:Taxon
221:Extent
212:
195:&
189:Region
1583:S2CID
1536:S2CID
1486:S2CID
1478:JSTOR
1423:S2CID
1376:S2CID
1368:JSTOR
1218:S2CID
1115:(PDF)
1093:JSTOR
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1005:S2CID
913:JSTOR
835:S2CID
764:S2CID
496:basal
1575:ISSN
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1516:2012
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