75:
33:
806:
is known only from Lagoa Santa, where the genus no longer occurs; the nearest records are about 70 km (43.5 mi) to the southeast. It is one of several Lagoa Santa fossil rodents that no longer occur in the area. Although the precise environmental background of the Lagoa Santa fossil
507:. Over many decades, the identity of most of Winge's species remained uncertain, and in many cases it was not until the 1990s that the original material was restudied to provide a definitive identification of the material. In a 2002 review of the fossil
647:
The zygomatic plates are almost completely vertical. As Winge already noted, the front margin of the zygomatic plate hardly extends forward before the antorbital bridge, which connects the plate to the body of the skull. This feature distinguishes
898:
Winge, 1888, p. 45; for current nomenclature: Voss and Myers, 1991, table 4; Musser and
Carleton, 2005, pp. 1111, 1145, 1149, 1154; Weksler and Bonvicino, 2005, p. 112; Weksler et al., 2006, pp. 8, 11, 14,
1068:
Pardiñas, U.F.J., D'Elía, G. and Ortiz, P.E. 2002. Sigmodontinos fósiles (Rodentia, Muroidea, Sigmodontinae) de América del sur: Estado actual de su conocimiento y prospectiva. Mastozoología
Neotropical 9(2):209–252 (in
290:(a bony plate on the side of the skull). It hardly extends forward in front of the connection between the plate and the main body of the skull, and that connection is relatively low on the skull. Furthermore, the
787:
behind the back end of the palate) is 4.29 mm long and 2.75 mm broad at the first molars. The upper molar row is 4.13 mm long and M1 is 1.19 mm broad. These measurements make
350:
of the collection in 1888. Winge described numerous new species, many of which received little attention from systematists afterward, and among these is the species he named
580:, both of which also occur in eastern Brazil, both have a broader interorbital region with better-developed ridges at the margins. One of three living species of
1179:
725:
have longer palates, extending beyond the third molars. Fine openings (foramina) are present on the palate. The back margin of the palate is squared;
532:, which currently includes three living species from southern Brazil and nearby Paraguay and Argentina. They recognized some features by which
1063:
1083:
Trouessart, E.L. 1898. Catalogus mammalium tam viventium quam fossilium. Tomus 2. Berlin: R. Friedländer and Sohn, 1469 pp. (in Latin).
555:
is known only from the front half of a skull, which was found in a cave known as "Lapa da Serra das
Abelhas". Compared to other species of
1159:
1174:
1169:
1154:
1101:
779:(gap) between the incisors and molars is 6.39 mm long. The incisive foramina are 5.25 mm long and 1.77 mm broad. The
526:" after re-examining the only known specimen in Copenhagen, and concluded that the animal was instead related to the living genus
676:
species, the front part of the zygomatic arches is bended forward and the zygomatic plates are bended outward. Furthermore, the
590:, also has a broader interorbital region. Behind the position of the nasals (which are missing in the only known skull) is an
807:
assemblage remains unclear, they may have been deposited in a period of climatic cooling that led to higher local diversity.
664:. In addition, the connection between the zygomatic plate and the antorbital bridge is inserted higher on the rostrum in
272:. The species remained unstudied and its affinities unclear until 2011, when it was recognized as a member of the genus
1052:
Musser, G.G. and
Carleton, M.D. 2005. Superfamily Muroidea. Pp. 894–1531 in Wilson, D.E. and Reeder, D.M. (eds.).
1164:
382:) "to look" and thus means "looking upward". Although Winge did not explain the name, it most likely refers to the
339:
261:
74:
1184:
591:
559:, it is large and has a more robust skull and teeth. Its rostrum (the front part) is broad and short and the
355:
784:
503:, conforming to the more conventional taxonomic arrangement; after that, the species was referred to as
408:
398:
180:
1106:
Bangs, 1900 (Rodentia, Sigmodontinae) of the
Brazilian Cerrado, with the description of two new species
776:
428:
1091:(Rodentia: Muridae) and the significance of Lund's collections from the caves of Lagoa Santa, Brazil
780:
1043:
Evolution of
Neotropical cricetine rodents (Muridae) with special reference to the phyllotine group
772:(a crest near the middle of the tooth). Although M3 is relatively large, its back part is reduced.
765:
737:
has the back margin U-shaped instead and has a longer palate, with the back margin behind the M3s.
618:
560:
50:
564:
707:
has even longer incisive foramina, extending between the molars, but the foramina are shorter in
608:
438:
418:
335:
257:
205:
69:
302:, and the palate is short, with its back margin between the third molars. The living species of
310:
in various characters, including shorter incisive foramina in two species and the shape of the
1149:
1059:
715:. The palate itself is wide and short, with its back margin between the M3s. Oryzomyines like
633:
586:
291:
280:
is probably a separate extinct species of the genus, which is no longer found at Lagoa Santa.
511:
rodents of South
America, Argentinean zoologist Ulyses Pardiñas and his coworkers wrote that
386:(a bone plate at the side of the skull), which is bended outward. Winge understood the genus
276:, which includes three other species from southern Brazil and nearby Argentina and Paraguay.
757:
692:
595:
346:
described Lund's collections in detail, among many others publishing a monograph about the
775:
The interorbital region is 4.14 mm long and the zygomatic plate is 2.38 mm. The
677:
536:
differs from the living species, and consequently kept it as a distinct, extinct species.
383:
287:
1042:
32:
669:
311:
1143:
700:
541:
508:
472:
299:
253:
146:
1127:
752:(with their cutting edge behind the vertical plane of the incisors). The molars are
703:) are broad and long, extending to the front margins of the first upper molar (M1).
544:, which is widespread and very diverse in South America and southern North America.
1128:
Jordfundne og nulevende
Gnavere (Rodentia) fra Lagoa Santa, Minas Geraes, Brasilien
1072:
Pardiñas, U.F.J. and Teta, P. 2011. On the taxonomic status of the
Brazilian mouse
745:
721:
599:
343:
265:
753:
749:
624:
764:, is divided into two smaller cusps on each side of the tooth by a valley, the
761:
603:
574:
569:
489:
480:
136:
59:
1058:. Baltimore: The Johns Hopkins University Press, 2 vols., 2142 pp.
680:, the notch between the zygomatic plate and arch, is deep, not shallow as in
392:
in a sense very different from that used today, including in it the species
86:
1055:
Mammal
Species of the World: a taxonomic and geographic reference. 3rd ed
769:
528:
499:
476:
286:
is similar to the other members of its genus in the configuration of its
248:
156:
106:
741:
448:
388:
696:
347:
295:
243:
126:
116:
96:
636:
forms a narrow projection towards the frontals, which is absent in
563:(between the skull) is hourglass-shaped, with squared margins. The
1116:
475:, which are characterized by the presence of a crest known as the
1087:
917:
Hershkovitz, 1962, p. 130; Voss and Myers, 1991, p. 417
783:(the portion of the palate between the incisive foramina and the
760:
arranged in pairs opposite each other. The front cusp of M1, the
729:
lacks a spine in the middle of the back margin, as is present in
264:, Brazil, in the first half of the 19th century and described by
1117:
Ten new genera of oryzomyine rodents (Cricetidae: Sigmodontinae)
926:
Trouessart, 1898, p. 527; Voss and Myers, 1991, p. 417
256:
known from a single broken skull. The specimen was collected by
1053:
471:
essentially included unspecialized species with pentalophodont
522:
In 2011, Pardiñas and Pablo Teta published another paper on "
519:", but this claim was based only on a cursory investigation.
373:
363:
338:
collected abundant remains of mammals around the village of
1011:
Pardiñas and Teta, 2011, p. 40; Winge, 1888, pp. 44–45
768:. Both M1 and the second molar (M2) have a well-developed
656:
and many oryzomyines, but the zygomatic plate of living
483:
on the lower molars, and excluded species now placed in
318:. The upper molar row is 4.13 mm long, which makes
540:
is a genus of unclear relations within the subfamily
1115:
Weksler, M., Percequillo, A.R. and Voss, R.S. 2006.
1095:Bulletin of the American Museum of Natural History
493:instead. Since 1898, authors have placed Winge's
672:(cheekbones) spread broadly. More than in other
602:is incompletely closed there. This interfrontal
935:Voss and Myers, 1991; Musser and Carleton, 2005
628:, a Brazilian rodent with some similarities to
1076:Winge, 1887 (Mammalia, Rodentia, Cricetidae).
962:Musser and Carleton, 2005, pp. 1086–1088, 1121
8:
342:in Brazil. After his death, his fellow Dane
31:
20:
748:at their front surfaces and are slightly
456:was the most divergent of the species of
44:, seen from the right, above, and below.
334:Between 1835 and 1849, Danish zoologist
998:
996:
994:
992:
982:
980:
978:
976:
974:
972:
970:
968:
815:
1100:Weksler, M. and Bonvicino, C.R. 2005.
867:
865:
863:
861:
859:
857:
855:
853:
851:
594:, a lowered portion of the skull; the
298:, extend to a point between the first
832:
830:
828:
7:
452:, species uncertain). He wrote that
294:, openings in the front part of the
871:Pardiñas and Teta, 2011, p. 41
1102:Taxonomy of pygmy rice rats genus
944:Pardiñas et al., 2002, p. 242
756:(low-crowned) and bear crests and
660:species closely resembles that of
14:
889:Voss and Myers, 1991, pp. 426–428
880:Voss and Myers, 1991, p. 415
1180:Extinct mammals of South America
1020:Pardiñas and Teta, 2011, table 1
1002:Pardiñas and Teta, 2011, table 2
73:
1086:Voss, R.S. and Myers, P. 1991.
1029:Pardiñas and Teta, 2011, p. 43
986:Pardiñas and Teta, 2011, p. 40
953:Pardiñas and Teta, 2011, p. 38
1:
845:Trouessart, 1898, p. 527
791:the largest known species of
699:between the incisors and the
616:, nor with most specimens of
598:(connection) between the two
467:Winge's concept of the genus
460:, but that it was similar to
322:the largest known species of
822:Winge, 1888, pl. III, fig. 4
40:Front half of the skull of
1201:
1160:Taxa named by Herluf Winge
1110:Arquivos do Museu Nacional
374:
364:
1175:Extinct animals of Brazil
1170:Endemic mammals of Brazil
1155:Mammals described in 1888
1121:American Museum Novitates
740:The well-developed upper
487:, which he classified in
362:, derives from the Greek
262:Lagoa Santa, Minas Gerais
229:: Pardiñas and Teta, 2011
211:
204:
186:
179:
70:Scientific classification
68:
57:
48:
39:
30:
23:
799:Distribution and habitat
592:interlacrymal depression
1134:1(3):1–200 (in Danish).
908:Winge, 1888, p. 45
836:Winge, 1888, p. 44
1041:Hershkovitz, P. 1962.
1089:Pseudoryzomys simplex
462:C. longicaudatus
409:Euryoryzomys russatus
399:Oligoryzomys nigripes
394:Calomys longicaudatus
429:Cerradomys subflavus
785:mesopterygoid fossa
766:anteromedian flexus
619:Juliomys ossitenuis
561:interorbital region
51:Conservation status
804:Juliomys anoblepas
709:J. ossitenuis
682:J. ossitenuis
609:Juliomys rimofrons
553:Juliomys anoblepas
505:Oryzomys anoblepas
439:Hylaeamys laticeps
419:Sooretamys angouya
336:Peter Wilhelm Lund
284:Juliomys anoblepas
258:Peter Wilhelm Lund
239:Juliomys anoblepas
227:Juliomys anoblepas
223:: Trouessart, 1898
221:Oryzomys anoblepas
193:Juliomys anoblepas
42:Juliomys anoblepas
25:Juliomys anoblepas
1165:Rodents of Brazil
1074:Calomys anoblepas
1064:978-0-8018-8221-0
1047:Fieldiana Zoology
789:J. anoblepas
727:J. anoblepas
695:(openings in the
693:incisive foramina
686:J. rimofrons
662:J. anoblepas
650:J. anoblepas
638:J. anoblepas
634:premaxillary bone
630:J. anoblepas
614:Juliomys pictipes
587:Juliomys pictipes
524:Calomys anoblepas
515:was "possibly an
513:O. anoblepas
479:on the upper and
454:C. anoblepas
404:Calomys coronatus
352:Calomys anoblepas
320:J. anoblepas
316:J. anoblepas
308:J. anoblepas
292:incisive foramina
278:J. anoblepas
270:Calomys anoblepas
252:of the subfamily
235:
234:
215:Calomys anoblepas
172:J. anoblepas
63:
16:Species of rodent
1192:
1126:Winge, H. 1888.
1036:Literature cited
1030:
1027:
1021:
1018:
1012:
1009:
1003:
1000:
987:
984:
963:
960:
954:
951:
945:
942:
936:
933:
927:
924:
918:
915:
909:
906:
900:
896:
890:
887:
881:
878:
872:
869:
846:
843:
837:
834:
823:
820:
713:J. pictipes
670:zygomatic arches
444:Calomys plebejus
434:Calomys saltator
424:Calomys laticeps
377:
376:
372:) "upwards" and
367:
366:
260:in the caves of
195:
191:
78:
77:
62:
35:
21:
1200:
1199:
1195:
1194:
1193:
1191:
1190:
1189:
1185:Extinct rodents
1140:
1139:
1038:
1033:
1028:
1024:
1019:
1015:
1010:
1006:
1001:
990:
985:
966:
961:
957:
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948:
943:
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912:
907:
903:
897:
893:
888:
884:
879:
875:
870:
849:
844:
840:
835:
826:
821:
817:
813:
801:
678:zygomatic notch
612:, but not with
606:is shared with
550:
384:zygomatic plate
332:
314:(cheekbone) in
288:zygomatic plate
200:
197:
189:
188:
175:
72:
64:
53:
17:
12:
11:
5:
1198:
1196:
1188:
1187:
1182:
1177:
1172:
1167:
1162:
1157:
1152:
1142:
1141:
1136:
1135:
1132:E Museo Lundii
1124:
1113:
1112:63(1):113–130.
1098:
1084:
1081:
1070:
1066:
1050:
1037:
1034:
1032:
1031:
1022:
1013:
1004:
988:
964:
955:
946:
937:
928:
919:
910:
901:
891:
882:
873:
847:
838:
824:
814:
812:
809:
800:
797:
781:palatal bridge
549:
546:
331:
328:
312:zygomatic arch
233:
232:
231:
230:
224:
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209:
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202:
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55:
54:
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37:
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28:
27:
15:
13:
10:
9:
6:
4:
3:
2:
1197:
1186:
1183:
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1178:
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1161:
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983:
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979:
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947:
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938:
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831:
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786:
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751:
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655:
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645:
643:
639:
635:
631:
627:
626:
621:
620:
615:
611:
610:
605:
601:
600:frontal bones
597:
593:
589:
588:
583:
579:
576:
572:
571:
566:
562:
558:
554:
547:
545:
543:
542:Sigmodontinae
539:
535:
531:
530:
525:
520:
518:
514:
510:
506:
502:
501:
496:
492:
491:
486:
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385:
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371:
361:
357:
356:specific name
353:
349:
345:
341:
337:
329:
327:
325:
321:
317:
313:
309:
305:
301:
297:
293:
289:
285:
281:
279:
275:
271:
267:
263:
259:
255:
254:Sigmodontinae
251:
250:
246:in the genus
245:
241:
240:
228:
225:
222:
219:
216:
213:
212:
210:
207:
203:
199:(Winge, 1888)
196:
194:
185:
182:
181:Binomial name
178:
174:
173:
167:
164:
163:
160:
159:
155:
152:
151:
148:
147:Sigmodontinae
145:
142:
141:
138:
135:
132:
131:
128:
125:
122:
121:
118:
115:
112:
111:
108:
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101:
98:
95:
92:
91:
88:
85:
82:
81:
76:
71:
67:
61:
56:
52:
47:
43:
38:
34:
29:
26:
22:
19:
1137:
1131:
1120:
1109:
1104:Oligoryzomys
1103:
1097:206:414–432.
1094:
1088:
1077:
1073:
1054:
1046:
1025:
1016:
1007:
958:
949:
940:
931:
922:
913:
904:
894:
885:
876:
841:
818:
803:
802:
792:
788:
774:
744:have orange
739:
734:
730:
726:
722:Oligoryzomys
720:
716:
712:
708:
704:
690:
685:
681:
673:
665:
661:
657:
653:
649:
646:
641:
637:
629:
623:
617:
613:
607:
585:
581:
577:
568:
565:thomasomyine
556:
552:
551:
537:
534:J. anoblepas
533:
527:
523:
521:
516:
512:
509:sigmodontine
504:
498:
494:
488:
484:
468:
466:
461:
457:
453:
447:
443:
437:
433:
427:
423:
417:
413:
407:
403:
397:
393:
387:
379:
369:
359:
351:
344:Herluf Winge
333:
323:
319:
315:
307:
306:differ from
303:
283:
282:
277:
273:
269:
266:Herluf Winge
247:
238:
237:
236:
226:
220:
214:
192:
187:
171:
170:
157:
41:
24:
18:
1080:2788:38–44.
754:brachyodont
750:opisthodont
735:Wilfredomys
705:Wilfredomys
666:Wilfredomys
654:Wilfredomys
625:Wilfredomys
548:Description
497:species in
446:(currently
436:(currently
426:(currently
416:(currently
414:Calomys rex
406:(currently
396:(currently
340:Lagoa Santa
268:in 1888 as
217:Winge, 1888
143:Subfamily:
1144:Categories
1123:3537:1–29.
811:References
762:anterocone
731:Rhipidomys
640:and other
604:fontanelle
575:oryzomyine
570:Rhipidomys
490:Hesperomys
481:mesolophid
137:Cricetidae
1069:Spanish).
1049:46:1–524.
644:species.
360:anoblepas
165:Species:
93:Kingdom:
87:Eukaryota
1150:Juliomys
793:Juliomys
777:diastema
770:mesoloph
742:incisors
674:Juliomys
658:Juliomys
642:Juliomys
582:Juliomys
573:and the
557:Juliomys
538:Juliomys
529:Juliomys
500:Oryzomys
477:mesoloph
330:Taxonomy
324:Juliomys
304:Juliomys
274:Juliomys
249:Juliomys
206:Synonyms
158:Juliomys
133:Family:
127:Rodentia
117:Mammalia
107:Chordata
103:Phylum:
97:Animalia
83:Domain:
1078:Zootaxa
717:Oecomys
578:Oecomys
567:rodent
517:Oecomys
495:Calomys
485:Calomys
469:Calomys
458:Calomys
449:Delomys
442:), and
389:Calomys
348:rodents
190:†
169:†
153:Genus:
123:Order:
113:Class:
60:Extinct
1062:
746:enamel
701:molars
697:palate
668:. The
632:, the
596:suture
473:molars
354:. The
300:molars
296:palate
244:rodent
758:cusps
652:from
622:. In
380:blepo
375:βλέπω
242:is a
1060:ISBN
719:and
711:and
691:The
684:and
432:),
422:),
412:),
402:),
370:ano
365:ἄνω
1146::
1130:.
1119:.
1108:.
1093:.
1045:.
991:^
967:^
899:23
850:^
827:^
795:.
733:.
688:.
584:,
464:.
358:,
326:.
378:(
368:(
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