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1762:; Linton, M.; Birren, B.; Nusbaum, C.; Zody, C.; Baldwin, J.; Devon, K.; Dewar, K.; Doyle, M.; Fitzhugh, W.; Funke, R.; Gage, D.; Harris, K.; Heaford, A.; Howland, J.; Kann, L.; Lehoczky, J.; Levine, R.; McEwan, P.; McKernan, K.; Meldrim, J.; Mesirov, J. P.; Miranda, C.; Morris, W.; Naylor, J.; Raymond, C.; Rosetti, M.; Santos, R.; Sheridan, A.; et al. (February 2001).
437:, to care for him and his sister Madeleine. and home-tutored them until he was five. At age five he entered the local preparatory school, York House School, where he soon developed an aversion to games. He developed an early interest in science, having fun with dissecting animals and sectioning plants to observe their structure and function. Sulston won a scholarship to
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585:. His initial research was in the field of chemical synthesis of oligonucleotides. Sulston began his work on C. elegans in 1974 characterising its DNA. Since then he has carried out a wide range of genetical and developmental studies on the nematode but his major research has been on the developmental lineage and
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Sulston was in favour of free public access of scientific information. He wanted genome information freely available, and he described as "totally immoral and disgusting" the idea of profiteering from such research. He also wanted to change patent law, and argued that restrictions on drugs such as
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that affect it. In a series of studies, culminating in a paper published in 1983, Sulston has analysed and described the total cell lineage of the nematode making it the first organism for which the origin of every cell is exactly known. This work is the basis for the study of mutations affecting
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lineages and is the foundation on which detailed studies of development in this organism will be based. Sulston has now turned his attention to an analysis of the genome of C. elegans and was constructing a total physical map using a novel method of analysing cloned DNA fragments.
1024:; Favello, A.; Fraser, A.; Fulton, L.; Gardner, A.; Green, P.; Hawkins, T.; Hillier, L.; Jier, M.; Johnston, L.; Jones, M.; Kershaw, J.; Kirsten, J.; Laisster, N.; Latreille, P.; Lightning, J. (1994). "2.2 Mb of contiguous nucleotide sequence from chromosome III of C. Elegans".
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518:(LMB). Sulston soon produced the complete map of the worm's neurons. He continued work on its DNA and subsequently the whole genome sequencing. In 1998, the whole genome sequence was published in collaboration with the Genome Institute at
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In 2000, after the 'working draft' of the human genome sequence was completed, Sulston retired from directing the Sanger Centre. With
Georgina Ferry, he narrated his research career leading to the human genome sequence in
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John
Sulston met Daphne Bate, a research assistant in Cambridge. They got married in 1966 just before they left for US for postdoctoral research. Together they had two children. Their first child, Ingrid, was born in
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417:. Sulston was in favour of science in the public interest, such as free public access of scientific information and against the patenting of genes and the privatisation of genetic technologies.
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Although brought up in a
Christian family, Sulston lost his faith during his student life at Cambridge, and remained an atheist. He was a distinguished supporter of
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1020:; Ainscough, R.; Anderson, K.; Baynes, C.; Berks, M.; Bonfield, J.; Burton, J.; Connell, M.; Copsey, T.; Cooper, J.; Coulson, A.; Craxton, M.; Dear, S.; Du, Z.;
740:. Sulston forfeited £15,000 of the £20,000 pledged in June 2012, as Assange had entered the embassy of Ecuador to escape the jurisdiction of the English courts.
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429:, England to Arthur Edward Aubrey Sulston and Josephine Muriel Frearson, née Blocksidge. His father was an Anglican priest and administrator of the
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Sulston, J.; Mallett, F.; Staden, R.; Durbin, R.; Horsnell, T.; Coulson, A. (1988). "Software for genome mapping by fingerprinting techniques".
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Sulston, J. E.; Schierenberg, E.; White, J. G.; Thomson, J. N. (1983). "The embryonic cell lineage of the nematode
Caenorhabditis elegans".
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Although Orgel wanted
Sulston to remain with him, Sydney Brenner persuaded Sulston to return to Cambridge to work on the neurobiology of
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One of
Sulston's most important contributions during his research years at the LMB was to elucidate the precise order in which cells in
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where they were active members of the local community: John regularly volunteered in the local library and in working parties at
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1562:(2000). "Alexander Robertus Todd, O.M., Baron Todd of Trumpington. 2 October 1907 -- 10 January 1997: Elected F.R.S. 1942".
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to show that large-scale genome sequencing projects were feasible. As sequencing of the worm genome proceeded, the
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Sulston, J. E.; Horvitz, H. R. (1977). "Post-embryonic cell lineages of the nematode, Caenorhabditis elegans".
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divide. In fact, he and his team succeeded in tracing the nematode's entire embryonic cell lineage.
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Freeview Video of
Fredrick Sanger in conversation with John Sulston by the Vega Science Trust
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John
Sulston is distinguished for his work on the molecular and developmental genetics of
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Sulston was a leading campaigner against the patenting of human genetic information.
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383:(27 March 1942 – 6 March 2018) was a British biologist and academic who won the
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Sulston, J.E.; Horvitz, H.R. (1977). "Post-embryonic cell lineages of the nematode,
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489:, California. His academic advisor Colin Reese had arranged for him to work with
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John
Sulston profile from the Medical Research Council lab for Molecular Biology
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The Common Thread: A Story of
Science, Politics, Ethics, and the Human Genome
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1953:"Nobel Laureate, Dr John Sulston, Receives the TCD Dawson Prize in Genetics"
1877:
The Common Thread a Story of Science, Politics, Ethics, and the Human Genome
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1639:(1999). "How the worm was won: the C. elegans genome sequencing project".
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in 1967, and their second, Adrian, later in England. The couple lived in
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by Roche are a hindrance to patients whose lives are dependent on them.
674:
On 23 October 2017 he was awarded the Cambridge Chemistry Alumni Medal.
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sequencing projects. He had argued successfully for the sequencing of
501:, who worked in Cambridge. He became inclined to biological research.
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John Sulston interviewed by Alan Macfarlane 16 September 2008 (video)
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841:"The 2000 George W. Beadle Medal. John Sulston and Robert Waterston"
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by acting as a bail surety for him, according to Assange's attorney
545:
began. At this point he was made director of the newly established
2805:
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2152:"Julian Assange supporters ordered to forfeit £93,500 bail money"
648:. In 2006, he was awarded the George Dawson Prize in Genetics by
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Princess of Asturias Award for Technical and Scientific Research
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1825:"Frederick Sanger CBE CH OM. 13 August 1918 – 19 November 2013"
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Sequencing Consortium (1998). "Genome Sequence of the Nematode
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Sulston died on 6 March 2018 of stomach cancer, aged 75 years.
526:
became the first animal to have its complete genome sequenced.
5048:
Prince of Asturias Award for Technical and Scientific Research
1931:"Golden Plate Awardees of the American Academy of Achievement"
25:
1357:"Knight in Common Armor: An Interview with Sir John Sulston"
409:. He was a leader in human genome research and Chair of the
2668:
Sainsbury Wellcome Centre for Neural Circuits and Behaviour
1237:"John E. Sulston, 75, Dies; Found Clues to Genes in a Worm"
1256:"Sir John Edward Sulston CH. 27 March 1942 – 6 March 2018"
717:. In 2003 he was one of 22 Nobel Laureates who signed the
710:; he was a Trustee of Cambridge Past, Present and Future.
1902:"Certificate of Election EC/1986/35: John Edward Sulston"
1919:
John Sulston: Nobel Prize in Physiology or Medicine 2002
644:
In 2004, Sulston received the Golden Plate Award of the
660:, which he gave on the subject of population pressure.
5641:
People educated at Merchant Taylors' School, Northwood
5636:
Members of the European Molecular Biology Organization
2129:"Wikileaks' Julian Assange tells of 'smear campaign'"
1764:"Initial sequencing and analysis of the human genome"
1880:(1st ed.). Washington, DC: Joseph Henry Press.
1830:
Biographical Memoirs of Fellows of the Royal Society
1564:
Biographical Memoirs of Fellows of the Royal Society
1260:
Biographical Memoirs of Fellows of the Royal Society
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2170:"Julian Assange's backers lose £200,000 bail money"
630:, both of whom he had collaborated with at the MRC
433:. His mother quit her job as an English teacher at
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23:
2240:on the Stapleford Cambridge website where he lives
2030:
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2724:Wellcome Trust Centre for the History of Medicine
1535:. American Academy of Achievement. Archived from
803:. Archived from the original on 25 September 2015
224:Physiological Society Annual Review Prize Lecture
5691:Members of the Order of the Companions of Honour
1533:"John Sulston Biography Nobel Prize in Medicine"
1472:
1470:
1468:
821:: CS1 maint: bot: original URL status unknown (
455:Department of Chemistry, University of Cambridge
5381:European Organization for Nuclear Research CERN
2785:Coalition for Epidemic Preparedness Innovations
2105:. American Humanist Association. Archived from
1330:"Professor Sir John Sulston - personal details"
665:Member of the Order of the Companions of Honour
579:
1254:Waterston, Robert H.; Ferry, Georgina (2019).
801:"Royal Society Terms, conditions and policies"
5018:
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8:
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411:Institute for Science, Ethics and Innovation
1904:. London: The Royal Society. Archived from
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770:. European Molecular Biology Organization.
529:Sulston played a central role in both the
51:
20:
16:British biologist and academic (1942–2018)
5661:Academics of the University of Manchester
5656:Nobel laureates in Physiology or Medicine
2219:British Scientists share 2002 Nobel Prize
2009:"Nobel Laureate awarded our Alumni Medal"
1842:
1690:: A Platform for Investigating Biology".
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575:Fellow of the Royal Society (FRS) in 1986
431:Society for the Propagation of the Gospel
5651:Academics of the University of Cambridge
3157:Wellcome Trust Principal Research Fellow
1997:(Supplement). 17 June 2017. p. B26.
1421:The Yale Journal of Biology and Medicine
1202:Computer Applications in the Biosciences
896:Aspects of oligoribonucleotide synthesis
834:
832:
317:Aspects of oligoribonucleotide synthesis
1874:Sulston, John; Ferry, Georgina (2002).
1498:
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991: ed.). A & C Black.
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903:(PhD thesis). University of Cambridge.
758:
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2682:WTC for Gene Regulation and Expression
814:
5646:Alumni of Pembroke College, Cambridge
5037:for Technical and Scientific Research
3222:Nobel Prize in Physiology or Medicine
3161:Wellcome Trust Senior Research Fellow
1837:. Royal Society publishing: 437–466.
779:
777:
671:for services to science and society.
632:Laboratory of Molecular Biology (LMB)
620:Nobel Prize in Physiology or Medicine
577:. His certificate of election reads:
483:Salk Institute for Biological Studies
477:Between 1966 and 1969 he worked as a
387:for his work on the cell lineage and
385:Nobel Prize in Physiology or Medicine
7:
5035:Prince or Princess of Asturias Award
3126:Institutional Strategic Support Fund
1336:from the original on 11 October 2009
604:Royal Institution Christmas Lectures
5163:Instituto Nacional de Biodiversidad
1080:"The DNA of Caenorhabditis elegans"
652:. In 2013, Sulston was awarded the
439:Merchant Taylors' School, Northwood
407:MRC Laboratory of Molecular Biology
105:Merchant Taylors' School, Northwood
2247:Non-profit organization positions
2224:John Sulston: One man and his worm
1417:"A conversation with John Sulston"
997:10.1093/ww/9780199540884.013.36669
602:in 2000. In 2001 Sulston gave the
520:Washington University in St. Louis
14:
5187:Pedro Miguel Echenique Landiríbar
2595:Roger Makins, 1st Baron Sherfield
1740:. Wellcome Trust Sanger Institute
1477:Ferry, Georgina (11 March 2018).
1295:Ferry, Georgina (11 March 2018).
1078:Sulston, J.; Brenner, S. (1974).
732:In December 2010, Sulston backed
5696:People from South Bucks District
3171:
3170:
2741:
2214:The public servant: John Sulston
2150:Booth, Robert (8 October 2012).
1400:
1332:. The University of Manchester.
922:
689:The Sulston Laboratories of the
5681:Human Genome Project scientists
3120:Collaborative Awards in Science
2832:Malawi Liverpool Wellcome Trust
1939:American Academy of Achievement
691:Wellcome Trust Sanger Institute
646:American Academy of Achievement
616:Robert Burns Humanitarian Award
516:Laboratory of Molecular Biology
296:Laboratory of Molecular Biology
281:Wellcome Trust Sanger Institute
162:
3123:Investigator Awards in Science
2729:Wellcome Research Laboratories
2694:WTC for Molecular Parasitology
2691:WTC for Mitochondrial Research
2470:Michael Augustine Raftery
2079:. British Humanist Association
1969:. Royal Society of New Zealand
1967:"Rutherford Memorial Lecturer"
1235:Kolata, Gina (15 March 2018).
1214:10.1093/bioinformatics/4.1.125
786:"Sir John Sulston FMedSci FRS"
693:are named in Sulston's honour.
1:
1712:10.1126/science.282.5396.2012
1653:10.1016/S0168-9525(98)01666-7
893:Sulston, John Edward (1966).
457:, after being interviewed by
5616:Fellows of the Royal Society
2677:WTC for Cell-Matrix Research
2103:Humanism and Its Aspirations
1615:10.1016/0012-1606(77)90158-0
1376:10.1371/journal.pgen.0020225
1179:10.1016/0012-1606(77)90158-0
1144:10.1016/0012-1606(83)90201-4
1004:UK public library membership
978:"Sulston, Sir John (Edward)"
654:Royal Society of New Zealand
451:Natural Sciences (Chemistry)
397:in 2002 with his colleagues
4314:Christiane Nüsslein-Volhard
2822:Genome Reference Consortium
2770:Cambridge Biomedical Campus
1479:"Sir John Sulston obituary"
1297:"Sir John Sulston obituary"
764:"John Sulston EMBO profile"
658:Rutherford Memorial Lecture
443:Pembroke College, Cambridge
5712:
2703:WTC for Stem Cell Research
2453:Thomas Nelson Marsham
2077:"Distinguished Supporters"
1458:Ivan Oransky, Adam Marcus
954:, accessed 11 October 2020
857:10.1093/genetics/157.2.467
704:Stapleford, Cambridgeshire
445:graduating in 1963 with a
5631:English former Christians
5418:
5407:
5093:Antonio González González
5071:Luis Antonio Santaló Sors
5053:
5042:
4930:
3511:Charles Scott Sherrington
3487:Frederick Gowland Hopkins
3168:
2930:Executive leadership team
2739:
2708:Wellcome Sanger Institute
2270:
2257:
2251:
2246:
337:
245:
50:
5454:Scientific Collaboration
5159:Manuel Losada Villasante
5135:Francisco Bolívar Zapata
4501:Françoise Barré-Sinoussi
2672:Science Learning Centres
2532:Gordon Richard Wray
1510:. Soylent Communications
1462:7 April 2018, The Lancet
1355:Gitschier, Jane (2006).
1096:10.1093/genetics/77.1.95
598:in 1989 and awarded the
512:Medical Research Council
465:in 1966 for research in
421:Early life and education
415:University of Manchester
301:University of Manchester
5676:English Nobel laureates
5611:British Nobel laureates
5545:Everett Peter Greenberg
5197:Enrique Moreno González
5141:Federico García Moliner
3129:Science Strategic Award
2888:Eliza Manningham-Buller
2751:Projects and facilities
2659:Francis Crick Institute
2476:Vulimiri Ramalingaswami
2131:. BBC. 17 December 2010
989:Oxford University Press
618:. Later, he shared the
479:postdoctoral researcher
427:Fulmer, Buckinghamshire
368:Sir John Edward Sulston
286:University of Cambridge
115:University of Cambridge
81:Fulmer, Buckinghamshire
5686:Caenorhabditis elegans
5424:Emmanuelle Charpentier
5083:David Vázquez Martínez
5077:Antonio Garcia-Bellido
4936:Nobel Prize recipients
4876:Physiology or Medicine
4867:(Italy, United States)
3282:Santiago Ramón y Cajal
3111:Awards and fellowships
2862:Wellcome Genome Campus
2790:COSMIC cancer database
2687:WTC for Human Genetics
2645:Centres and institutes
2559:Antonio Garcia-Bellido
2501:Charles J. M. Stirling
2029:Sulston, John (2002).
1844:10.1098/rsbm.2015.0013
1599:Caenorhabditis elegans
1576:10.1098/rsbm.1999.0099
1460:John Sulston. obituary
1273:10.1098/rsbm.2019.0014
694:
650:Trinity College Dublin
610:. In 2002, he won the
600:George W. Beadle Award
592:
583:Caenorhabditis elegans
573:Sulston was elected a
507:Caenorhabditis elegans
435:Watford Grammar School
394:Caenorhabditis elegans
188:W. Alden Spencer Award
131:Caenorhabditis elegans
5353:Arturo Álvarez-Buylla
3783:Dickinson W. Richards
3467:Julius Wagner-Jauregg
3153:Wellcome Trust Centre
3150:Wellcome Image Awards
2775:Cancer Genome Project
2481:Peter Richardson
1736:Caenorhabditis genome
1603:Developmental Biology
1167:Developmental Biology
1132:Developmental Biology
688:
669:2017 Birthday Honours
5315:George M. Whitesides
4206:Rita Levi-Montalcini
3567:Albert Szent-Györgyi
2827:Human Genome Project
2800:Diamond Light Source
2758:1000 Genomes Project
2698:WTC for Neuroimaging
2521:Elizabeth Warrington
2238:John Sulston profile
1415:Sulston, J. (2002).
901:repository.cam.ac.uk
725:the anti-viral drug
543:Human Genome Project
461:and was awarded his
425:Sulston was born in
5559:Jeffrey M. Friedman
5387:Avelino Corma Canós
5183:Emilio Méndez Pérez
4598:William C. Campbell
4509:Elizabeth Blackburn
4346:Robert F. Furchgott
4338:Stanley B. Prusiner
4330:Rolf M. Zinkernagel
4230:George H. Hitchings
4194:Joseph L. Goldstein
4154:Bengt I. Samuelsson
3983:Earl Sutherland Jr.
3146:Wellcome Book Prize
3138:Sir Henry Wellcome
2857:Wellcome Collection
2361:Dennis Chapman
2172:. 4 September 2012.
2037:. Bantam. pp.
1935:www.achievement.org
1821:Brownlee, George G.
1783:2001Natur.409..860L
1704:1998Sci...282.2012.
1698:(5396): 2012–2018.
1038:1994Natur.368...32W
839:Kimble, J. (2001).
663:He was appointed a
608:The Secrets of Life
67:John Edward Sulston
5357:Giacomo Rizzolatti
5287:Juan Ignacio Cirac
5087:Emilio Rosenblueth
4658:William Kaelin Jr.
4654:Peter J. Ratcliffe
4397:Leland H. Hartwell
4286:Richard J. Roberts
4166:Barbara McClintock
4110:Godfrey Hounsfield
4011:Nikolaas Tinbergen
3943:Marshall Nirenberg
3911:Charles B. Huggins
3700:Tadeusz Reichstein
3688:António Egas Moniz
3483:Christiaan Eijkman
2953:Peter Pereira Gray
2881:Board of governors
2554:Albert Eschenmoser
2516:David Wallace
2496:Michael Smith
2402:Martin Fleischmann
2355:Geoffrey Burnstock
2205:on Nobelprize.org
2063:"Our achievements"
1994:The London Gazette
1955:. 18 October 2006.
1641:Trends in Genetics
1637:Wilson, Richard K.
1241:The New York Times
948:on Nobelprize.org
917:uk.bl.ethos.648083
909:10.17863/CAM.16307
719:Humanist Manifesto
695:
594:He was elected an
569:Awards and honours
5621:British humanists
5583:
5582:
5579:
5578:
5555:Daniel J. Drucker
5541:Jeffrey I. Gordon
5479:Ingrid Daubechies
5469:Sandra Myrna Díaz
5403:
5402:
5367:Richard A. Lerner
5329:Raymond Tomlinson
5125:Santiago Grisolía
5033:Laureates of the
5000:
4999:
4907:Economic Sciences
4890:H. Robert Horvitz
4865:Riccardo Giacconi
4859:Masatoshi Koshiba
4853:Raymond Davis Jr.
4779:John Bennett Fenn
4727:
4726:
4692:Ardem Patapoutian
4541:Ralph M. Steinman
4537:Jules A. Hoffmann
4525:Robert G. Edwards
4493:Harald zur Hausen
4417:H. Robert Horvitz
4318:Eric F. Wieschaus
4254:E. Donnall Thomas
4238:J. Michael Bishop
4226:Gertrude B. Elion
4098:Hamilton O. Smith
4023:Christian de Duve
3759:Frederick Robbins
3636:Alexander Fleming
3575:Corneille Heymans
3421:Frederick Banting
3326:Allvar Gullstrand
3184:
3183:
2898:Tobias Bonhoeffer
2737:
2736:
2604:
2603:
2417:John Rodney Guest
2407:C. Robin Ganellin
2280:
2279:
2271:Succeeded by
2109:on 5 October 2012
2099:"Notable Signers"
2033:The Common Thread
1887:978-0-309-08409-3
1777:(6822): 860–921.
1504:"John E. Sulston"
1002:(Subscription or
796:on 10 April 2016.
365:
364:
357:/honorary-faculty
265:Molecular biology
247:Scientific career
176:1 son, 1 daughter
155:Daphne Edith Bate
126:Genome sequencing
5703:
5666:Knights Bachelor
5626:English atheists
5409:
5377:François Englert
5235:Lawrence Roberts
5229:Jean Weissenbach
5153:Manuel Patarroyo
5129:Salvador Moncada
5113:Marcos Moshinsky
5065:Manuel Ballester
5044:
5027:
5020:
5013:
5004:
4938:
4898:(United Kingdom)
4886:(United Kingdom)
4754:
4747:
4740:
4731:
4720:
4705:
4694:
4679:
4673:Michael Houghton
4660:
4650:Gregg L. Semenza
4644:
4638:James P. Allison
4632:
4630:Michael W. Young
4616:
4614:Yoshinori Ohsumi
4608:
4592:
4576:
4574:Thomas C. Südhof
4560:
4548:
4527:
4519:
4513:Carol W. Greider
4503:
4487:
4471:
4459:
4447:
4435:
4423:
4407:
4380:
4364:
4356:
4340:
4332:
4326:Peter C. Doherty
4320:
4304:
4298:Alfred G. Gilman
4292:
4280:
4268:
4256:
4244:
4242:Harold E. Varmus
4232:
4216:
4208:
4196:
4184:
4168:
4160:
4144:
4128:
4118:Baruj Benacerraf
4112:
4100:
4084:
4068:
4045:
4029:
4013:
3997:
3985:
3977:
3961:
3945:
3935:Robert W. Holley
3929:
3913:
3901:
3885:
3873:
3857:
3841:
3839:Georg von Békésy
3833:
3821:
3809:
3807:Joshua Lederberg
3793:
3785:
3779:Werner Forssmann
3769:
3761:
3745:
3733:
3725:
3706:
3690:
3678:
3670:
3668:Bernardo Houssay
3654:
3646:
3630:
3618:
3606:
3605:
3599:
3598:
3592:
3591:
3585:
3577:
3569:
3561:
3549:
3541:
3525:
3517:
3505:
3497:
3495:Karl Landsteiner
3489:
3477:
3469:
3461:
3459:Johannes Fibiger
3442:
3441:
3435:
3433:Willem Einthoven
3427:
3415:
3403:
3402:
3396:
3388:
3380:
3379:
3373:
3372:
3366:
3365:
3359:
3358:
3352:
3344:
3336:
3328:
3320:
3312:
3304:
3298:Élie Metchnikoff
3292:
3290:Alphonse Laveran
3284:
3272:
3264:
3256:
3248:
3240:
3211:
3204:
3197:
3188:
3174:
3173:
3020:Other key people
2985:Former directors
2959:Chonnettia Jones
2893:Michael Ferguson
2867:Wellcome Library
2745:
2664:Gurdon Institute
2649:
2631:
2624:
2617:
2608:
2491:Raymond Smallman
2459:William Mitchell
2367:John Clarke
2306:
2299:
2292:
2283:
2268:1993–2000
2264:Sanger Institute
2252:Preceded by
2244:
2210:
2174:
2173:
2166:
2160:
2159:
2147:
2141:
2140:
2138:
2136:
2125:
2119:
2118:
2116:
2114:
2095:
2089:
2088:
2086:
2084:
2073:
2067:
2066:
2059:
2053:
2052:
2036:
2026:
2020:
2019:
2017:
2015:
2005:
1999:
1998:
1985:
1979:
1978:
1976:
1974:
1963:
1957:
1956:
1949:
1943:
1942:
1927:
1921:
1916:
1910:
1909:
1898:
1892:
1891:
1871:
1865:
1864:
1846:
1817:
1811:
1810:
1791:10.1038/35057062
1768:
1756:
1750:
1749:
1747:
1745:
1730:
1724:
1723:
1679:
1673:
1672:
1633:
1627:
1626:
1594:
1588:
1587:
1555:
1549:
1548:
1546:
1544:
1539:on 23 April 2014
1529:
1520:
1519:
1517:
1515:
1500:
1487:
1486:
1474:
1463:
1456:
1447:
1446:
1436:
1427:(5–6): 299–306.
1412:
1406:
1405:
1404:
1398:
1388:
1378:
1352:
1346:
1345:
1343:
1341:
1326:
1320:
1319:
1317:
1315:
1292:
1286:
1285:
1275:
1251:
1245:
1244:
1232:
1226:
1225:
1197:
1191:
1190:
1162:
1156:
1155:
1127:
1118:
1117:
1107:
1075:
1066:
1065:
1046:10.1038/368032a0
1014:
1008:
1007:
1000:
980:
972:
955:
953:
943:
928:
927:
926:
920:
890:
879:
878:
868:
836:
827:
826:
820:
812:
810:
808:
797:
792:. Archived from
781:
772:
771:
760:
453:. He joined the
447:Bachelor of Arts
382:
361:
358:
356:
354:
352:
350:
348:
329:Doctoral advisor
323:
166:
164:
95:
76:
74:
55:
45:
21:
5711:
5710:
5706:
5705:
5704:
5702:
5701:
5700:
5586:
5585:
5584:
5575:
5571:Svetlana Mojsov
5567:Jens Juul Holst
5563:Joel F. Habener
5523:Geoffrey Hinton
5487:Emmanuel Candès
5448:Barry C. Barish
5428:Jennifer Doudna
5414:
5399:
5395:Galen D. Stucky
5281:Antonio Damasio
5275:Robert Weinberg
5271:Bert Vogelstein
5247:Tim Berners-Lee
5221:Francis Collins
5170:Valentín Fuster
5049:
5038:
5031:
5001:
4996:
4934:
4926:
4923:(United States)
4921:Vernon L. Smith
4915:Daniel Kahneman
4901:
4896:John E. Sulston
4892:(United States)
4870:
4855:(United States)
4839:
4836:(United States)
4820:
4796:
4781:(United States)
4765:
4758:
4728:
4723:
4708:
4697:
4682:
4677:Charles M. Rice
4669:Harvey J. Alter
4663:
4647:
4635:
4626:Michael Rosbash
4622:Jeffrey C. Hall
4619:
4611:
4595:
4586:May-Britt Moser
4579:
4563:
4558:Shinya Yamanaka
4551:
4530:
4522:
4517:Jack W. Szostak
4506:
4490:
4485:Oliver Smithies
4474:
4462:
4450:
4438:
4433:Peter Mansfield
4426:
4421:John E. Sulston
4410:
4394:
4383:
4367:
4359:
4343:
4335:
4323:
4310:Edward B. Lewis
4307:
4295:
4283:
4271:
4259:
4247:
4235:
4219:
4214:Susumu Tonegawa
4211:
4199:
4187:
4171:
4163:
4147:
4131:
4115:
4103:
4087:
4074:Roger Guillemin
4071:
4066:Daniel Gajdusek
4062:Baruch Blumberg
4059:
4048:
4039:Renato Dulbecco
4035:David Baltimore
4032:
4016:
4003:Karl von Frisch
4000:
3988:
3980:
3964:
3948:
3932:
3923:Haldan Hartline
3916:
3904:
3888:
3876:
3860:
3855:Maurice Wilkins
3844:
3836:
3824:
3819:Arthur Kornberg
3812:
3796:
3788:
3772:
3764:
3748:
3736:
3728:
3720:
3709:
3693:
3681:
3673:
3657:
3649:
3633:
3624:Joseph Erlanger
3621:
3609:
3603:
3602:
3596:
3595:
3589:
3588:
3580:
3572:
3564:
3552:
3544:
3528:
3520:
3508:
3500:
3492:
3480:
3475:Charles Nicolle
3472:
3464:
3456:
3445:
3439:
3438:
3430:
3418:
3406:
3400:
3399:
3391:
3383:
3377:
3376:
3370:
3369:
3363:
3362:
3356:
3355:
3347:
3339:
3331:
3323:
3318:Albrecht Kossel
3315:
3307:
3295:
3287:
3275:
3267:
3259:
3251:
3243:
3235:
3224:
3215:
3185:
3180:
3164:
3132:Sir Henry Dale
3106:
3037:Dominic Cadbury
3032:William Castell
3015:
2998:Bridget Ogilvie
2980:
2944:Stephen Caddick
2925:
2876:
2746:
2733:
2712:
2640:
2635:
2605:
2600:
2583:
2537:
2442:Chris J. Leaver
2382:Simon Donaldson
2335:Roy M. Anderson
2323:
2310:
2276:
2267:
2255:
2234:
2203:John E. Sulston
2183:
2178:
2177:
2168:
2167:
2163:
2149:
2148:
2144:
2134:
2132:
2127:
2126:
2122:
2112:
2110:
2097:
2096:
2092:
2082:
2080:
2075:
2074:
2070:
2061:
2060:
2056:
2049:
2028:
2027:
2023:
2013:
2011:
2007:
2006:
2002:
1987:
1986:
1982:
1972:
1970:
1965:
1964:
1960:
1951:
1950:
1946:
1929:
1928:
1924:
1917:
1913:
1908:on 8 July 2019.
1900:
1899:
1895:
1888:
1873:
1872:
1868:
1819:
1818:
1814:
1766:
1758:
1757:
1753:
1743:
1741:
1732:
1731:
1727:
1688:C. elegans
1681:
1680:
1676:
1635:
1634:
1630:
1596:
1595:
1591:
1557:
1556:
1552:
1542:
1540:
1531:
1530:
1523:
1513:
1511:
1502:
1501:
1490:
1476:
1475:
1466:
1457:
1450:
1414:
1413:
1409:
1399:
1354:
1353:
1349:
1339:
1337:
1328:
1327:
1323:
1313:
1311:
1294:
1293:
1289:
1253:
1252:
1248:
1234:
1233:
1229:
1199:
1198:
1194:
1164:
1163:
1159:
1129:
1128:
1121:
1077:
1076:
1069:
1032:(6466): 32–38.
1016:
1015:
1011:
1001:
974:
973:
958:
946:John E. Sulston
944:
931:
921:
892:
891:
882:
838:
837:
830:
813:
806:
804:
799:
783:
782:
775:
768:people.embo.org
762:
761:
754:
749:
683:
612:Dan David Prize
571:
475:
423:
370:
345:
321:
305:
269:
241:
218:Edinburgh Medal
212:Knight Bachelor
194:EMBO Membership
168:
165: 1966)
160:
156:
142:
138:
111:Alma mater
93:
84:
78:
72:
70:
69:
68:
58:
57:Sulston in 2008
46:
33:
31:
28:
17:
12:
11:
5:
5709:
5707:
5699:
5698:
5693:
5688:
5683:
5678:
5673:
5671:Wellcome Trust
5668:
5663:
5658:
5653:
5648:
5643:
5638:
5633:
5628:
5623:
5618:
5613:
5608:
5603:
5598:
5588:
5587:
5581:
5580:
5577:
5576:
5574:
5573:
5551:
5549:Bonnie Bassler
5537:
5535:Demis Hassabis
5519:
5501:Philip Felgner
5493:Katalin Karikó
5489:
5471:
5461:
5455:
5436:
5430:
5419:
5416:
5415:
5412:
5405:
5404:
5401:
5400:
5398:
5397:
5383:
5369:
5363:Gregory Winter
5359:
5345:
5331:
5321:
5307:Shuji Nakamura
5299:
5293:Peter Lawrence
5289:
5283:
5277:
5255:
5249:
5239:Robert E. Kahn
5231:
5225:Hamilton Smith
5209:
5207:Luc Montagnier
5199:
5193:Ricardo Miledi
5189:
5179:
5172:
5166:
5155:
5149:
5143:
5137:
5131:
5121:
5115:
5109:Manuel Cardona
5105:
5099:Jacinto Convit
5095:
5089:
5079:
5073:
5067:
5061:
5054:
5051:
5050:
5047:
5040:
5039:
5032:
5030:
5029:
5022:
5015:
5007:
4998:
4997:
4995:
4994:
4989:
4984:
4979:
4974:
4969:
4964:
4959:
4954:
4949:
4944:
4939:
4931:
4928:
4927:
4925:
4924:
4918:
4911:
4909:
4903:
4902:
4900:
4899:
4893:
4887:
4884:Sydney Brenner
4880:
4878:
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4856:
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4773:
4767:
4766:
4759:
4757:
4756:
4749:
4742:
4734:
4725:
4724:
4722:
4721:
4714:Katalin Karikó
4706:
4695:
4680:
4661:
4645:
4633:
4617:
4609:
4593:
4577:
4570:Randy Schekman
4561:
4549:
4528:
4520:
4504:
4497:Luc Montagnier
4488:
4477:Mario Capecchi
4472:
4460:
4453:Barry Marshall
4448:
4436:
4429:Paul Lauterbur
4424:
4413:Sydney Brenner
4408:
4391:
4389:
4385:
4384:
4382:
4381:
4374:Paul Greengard
4370:Arvid Carlsson
4365:
4357:
4341:
4333:
4321:
4305:
4302:Martin Rodbell
4293:
4281:
4278:Edwin G. Krebs
4274:Edmond Fischer
4269:
4257:
4245:
4233:
4222:James W. Black
4217:
4209:
4197:
4185:
4182:César Milstein
4178:Georges Köhler
4169:
4161:
4150:Sune Bergström
4145:
4142:Torsten Wiesel
4138:David H. Hubel
4129:
4113:
4101:
4094:Daniel Nathans
4085:
4078:Andrew Schally
4069:
4056:
4054:
4050:
4049:
4047:
4046:
4030:
4014:
3998:
3991:Gerald Edelman
3986:
3978:
3975:Julius Axelrod
3962:
3959:Salvador Luria
3955:Alfred Hershey
3946:
3930:
3914:
3902:
3891:François Jacob
3886:
3874:
3858:
3842:
3834:
3822:
3810:
3794:
3786:
3775:André Cournand
3770:
3762:
3746:
3734:
3731:Selman Waksman
3726:
3717:
3715:
3711:
3710:
3708:
3707:
3696:Edward Kendall
3691:
3679:
3671:
3655:
3652:Hermann Muller
3647:
3631:
3628:Herbert Gasser
3619:
3607:
3600:
3593:
3586:
3583:Gerhard Domagk
3578:
3570:
3562:
3550:
3542:
3539:William Murphy
3531:George Whipple
3526:
3518:
3506:
3498:
3490:
3478:
3470:
3462:
3453:
3451:
3447:
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3444:
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3436:
3428:
3416:
3409:Archibald Hill
3404:
3397:
3389:
3381:
3374:
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3360:
3353:
3345:
3342:Charles Richet
3337:
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3159:
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3136:
3130:
3127:
3124:
3121:
3118:
3117:Capital Awards
3114:
3112:
3108:
3107:
3105:
3104:
3102:Henry Wellcome
3099:
3094:
3089:
3084:
3079:
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3049:
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3023:
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3017:
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3007:
3004:Michael Dexter
3001:
2995:
2992:Peter Williams
2988:
2986:
2982:
2981:
2979:
2978:
2975:
2972:
2969:
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2956:Mark Henderson
2954:
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2948:
2945:
2942:
2939:
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2927:
2926:
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2911:Bryan Grenfell
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2669:
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2646:
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2638:Wellcome Trust
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2508:
2503:
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2478:
2473:
2466:
2461:
2456:
2449:
2447:George Lorimer
2444:
2439:
2434:
2429:
2424:
2419:
2414:
2409:
2404:
2399:
2397:Peter Fellgett
2394:
2392:John M. Edmond
2389:
2384:
2379:
2374:
2372:Peter Day
2369:
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2182:
2181:External links
2179:
2176:
2175:
2161:
2142:
2120:
2090:
2068:
2065:. 10 May 2018.
2054:
2048:978-0309084093
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1944:
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1911:
1893:
1886:
1866:
1812:
1751:
1725:
1674:
1628:
1609:(1): 110–156.
1589:
1558:Brown, D. M.;
1550:
1521:
1488:
1464:
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1407:
1347:
1321:
1287:
1246:
1227:
1208:(1): 125–132.
1192:
1173:(1): 110–156.
1157:
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1009:
956:
929:
880:
851:(2): 467–468.
828:
773:
751:
750:
748:
745:
734:Julian Assange
682:
679:
628:Robert Horvitz
624:Sydney Brenner
570:
567:
555:Cambridgeshire
553:), located in
499:Sydney Brenner
474:
471:
459:Alexander Todd
422:
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403:Robert Horvitz
399:Sydney Brenner
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233:
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122:Known for
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98:
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96:(aged 75)
90:
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5609:
5607:
5606:Nematologists
5604:
5602:
5599:
5597:
5594:
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5591:
5572:
5568:
5564:
5560:
5556:
5552:
5550:
5546:
5542:
5538:
5536:
5532:
5531:Yoshua Bengio
5528:
5524:
5520:
5518:
5517:Sarah Gilbert
5514:
5513:Derrick Rossi
5510:
5506:
5502:
5498:
5497:Drew Weissman
5494:
5490:
5488:
5484:
5480:
5476:
5472:
5470:
5466:
5462:
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5456:
5453:
5449:
5445:
5444:Kip S. Thorne
5441:
5437:
5435:
5431:
5429:
5425:
5421:
5420:
5417:
5410:
5406:
5396:
5392:
5391:Mark E. Davis
5388:
5384:
5382:
5378:
5374:
5370:
5368:
5364:
5360:
5358:
5354:
5350:
5349:Joseph Altman
5346:
5344:
5343:Linda Watkins
5340:
5336:
5332:
5330:
5326:
5325:Martin Cooper
5322:
5320:
5316:
5312:
5311:Robert Langer
5308:
5304:
5300:
5298:
5294:
5290:
5288:
5284:
5282:
5278:
5276:
5272:
5268:
5267:Joan Massagué
5264:
5260:
5259:Judah Folkman
5256:
5254:
5250:
5248:
5244:
5240:
5236:
5232:
5230:
5226:
5222:
5218:
5214:
5210:
5208:
5204:
5200:
5198:
5194:
5190:
5188:
5184:
5180:
5178:research team
5177:
5173:
5171:
5167:
5165:of Costa Rica
5164:
5160:
5156:
5154:
5150:
5148:
5144:
5142:
5138:
5136:
5132:
5130:
5126:
5122:
5120:
5116:
5114:
5110:
5106:
5104:
5103:Pablo Rudomín
5100:
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5066:
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4827:
4823:
4816:
4813:
4812:
4810:
4807:
4803:
4799:
4793:(Switzerland)
4792:
4791:Kurt Wüthrich
4789:
4786:
4785:Koichi Tanaka
4783:
4780:
4777:
4776:
4774:
4772:
4768:
4763:
4755:
4750:
4748:
4743:
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4732:
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4718:Drew Weissman
4715:
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4602:Satoshi Ōmura
4599:
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4587:
4583:
4578:
4575:
4571:
4567:
4566:James Rothman
4562:
4559:
4555:
4550:
4546:
4542:
4538:
4534:
4533:Bruce Beutler
4529:
4526:
4521:
4518:
4514:
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4505:
4502:
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4486:
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4449:
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4445:Linda B. Buck
4442:
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4434:
4430:
4425:
4422:
4418:
4414:
4409:
4406:
4402:
4398:
4393:
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4390:
4386:
4379:
4375:
4371:
4366:
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4362:Günter Blobel
4358:
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4351:
4350:Louis Ignarro
4347:
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4339:
4334:
4331:
4327:
4322:
4319:
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4299:
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4290:Phillip Sharp
4287:
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4279:
4275:
4270:
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4263:
4258:
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4250:Joseph Murray
4246:
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4218:
4215:
4210:
4207:
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4202:Stanley Cohen
4198:
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4190:Michael Brown
4186:
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4106:Allan Cormack
4102:
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4082:Rosalyn Yalow
4079:
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4040:
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4027:George Palade
4024:
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4019:Albert Claude
4015:
4012:
4008:
4007:Konrad Lorenz
4004:
3999:
3996:
3995:Rodney Porter
3992:
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3971:Ulf von Euler
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3868:
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3847:Francis Crick
3843:
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3835:
3832:
3831:Peter Medawar
3828:
3823:
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3816:
3811:
3808:
3804:
3800:
3799:George Beadle
3795:
3792:
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3767:Hugo Theorell
3763:
3760:
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3755:Thomas Weller
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3413:Otto Meyerhof
3410:
3405:
3398:
3395:
3390:
3387:
3382:
3375:
3368:
3361:
3354:
3351:
3350:Róbert Bárány
3346:
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3334:Alexis Carrel
3330:
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3319:
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3311:
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3299:
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3067:Richard Hynes
3065:
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3052:Oliver Franks
3050:
3048:
3045:
3043:
3040:
3038:
3035:
3033:
3030:
3028:
3027:Damon Buffini
3025:
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3018:
3011:
3008:
3005:
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2999:
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2990:
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2947:Simon Chaplin
2946:
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2940:
2938:
2937:Jeremy Farrar
2935:
2934:
2932:
2928:
2922:
2919:
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2916:Naguib Kheraj
2914:
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2906:William Burns
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2657:
2656:
2654:
2650:
2647:
2643:
2639:
2632:
2627:
2625:
2620:
2618:
2613:
2612:
2609:
2596:
2593:
2592:
2590:
2586:
2580:
2577:
2575:
2572:
2570:
2569:Edwin H. Land
2567:
2565:
2564:Joseph Keller
2562:
2560:
2557:
2555:
2552:
2550:
2547:
2546:
2544:
2540:
2534:
2533:
2529:
2527:
2524:
2522:
2519:
2517:
2514:
2512:
2509:
2507:
2504:
2502:
2499:
2497:
2494:
2492:
2489:
2487:
2486:Harold Ridley
2484:
2482:
2479:
2477:
2474:
2472:
2471:
2467:
2465:
2464:Keith Moffatt
2462:
2460:
2457:
2455:
2454:
2450:
2448:
2445:
2443:
2440:
2438:
2435:
2433:
2432:Alec Jeffreys
2430:
2428:
2427:Werner Israel
2425:
2423:
2420:
2418:
2415:
2413:
2410:
2408:
2405:
2403:
2400:
2398:
2395:
2393:
2390:
2388:
2385:
2383:
2380:
2378:
2377:Richard Dixon
2375:
2373:
2370:
2368:
2365:
2363:
2362:
2358:
2356:
2353:
2351:
2348:
2346:
2343:
2341:
2338:
2336:
2333:
2332:
2330:
2326:
2322:
2318:
2317:Royal Society
2314:
2307:
2302:
2300:
2295:
2293:
2288:
2287:
2284:
2275:
2274:Allan Bradley
2266:
2265:
2261:
2250:
2245:
2239:
2236:
2235:
2231:
2230:
2225:
2222:
2220:
2217:
2215:
2212:
2209:
2204:
2201:
2199:
2196:
2194:
2191:
2189:
2186:
2185:
2180:
2171:
2165:
2162:
2157:
2153:
2146:
2143:
2130:
2124:
2121:
2108:
2104:
2100:
2094:
2091:
2078:
2072:
2069:
2064:
2058:
2055:
2050:
2044:
2040:
2035:
2034:
2025:
2022:
2010:
2004:
2001:
1996:
1995:
1990:
1984:
1981:
1968:
1962:
1959:
1954:
1948:
1945:
1940:
1936:
1932:
1926:
1923:
1920:
1915:
1912:
1907:
1903:
1897:
1894:
1889:
1883:
1879:
1878:
1870:
1867:
1862:
1858:
1854:
1850:
1845:
1840:
1836:
1832:
1831:
1826:
1822:
1816:
1813:
1808:
1804:
1800:
1796:
1792:
1788:
1784:
1780:
1776:
1772:
1765:
1761:
1760:Lander, E. S.
1755:
1752:
1739:
1737:
1729:
1726:
1721:
1717:
1713:
1709:
1705:
1701:
1697:
1693:
1689:
1685:
1678:
1675:
1670:
1666:
1662:
1658:
1654:
1650:
1646:
1642:
1638:
1632:
1629:
1624:
1620:
1616:
1612:
1608:
1604:
1600:
1593:
1590:
1585:
1581:
1577:
1573:
1569:
1565:
1561:
1554:
1551:
1538:
1534:
1528:
1526:
1522:
1509:
1505:
1499:
1497:
1495:
1493:
1489:
1484:
1480:
1473:
1471:
1469:
1465:
1461:
1455:
1453:
1449:
1444:
1440:
1435:
1430:
1426:
1422:
1418:
1411:
1408:
1403:
1396:
1392:
1387:
1382:
1377:
1372:
1368:
1364:
1363:
1362:PLOS Genetics
1358:
1351:
1348:
1335:
1331:
1325:
1322:
1310:
1306:
1302:
1298:
1291:
1288:
1283:
1279:
1274:
1269:
1265:
1261:
1257:
1250:
1247:
1242:
1238:
1231:
1228:
1223:
1219:
1215:
1211:
1207:
1203:
1196:
1193:
1188:
1184:
1180:
1176:
1172:
1168:
1161:
1158:
1153:
1149:
1145:
1141:
1138:(1): 64–119.
1137:
1133:
1126:
1124:
1120:
1115:
1111:
1106:
1101:
1097:
1093:
1090:(1): 95–104.
1089:
1085:
1081:
1074:
1072:
1068:
1063:
1059:
1055:
1051:
1047:
1043:
1039:
1035:
1031:
1027:
1023:
1019:
1013:
1010:
1005:
998:
994:
990:
986:
985:
979:
975:Anon (2015).
971:
969:
967:
965:
963:
961:
957:
952:
947:
942:
940:
938:
936:
934:
930:
925:
918:
914:
910:
906:
902:
898:
897:
889:
887:
885:
881:
876:
872:
867:
862:
858:
854:
850:
846:
842:
835:
833:
829:
824:
818:
802:
795:
791:
790:Royal Society
787:
784:Anon (1986).
780:
778:
774:
769:
765:
759:
757:
753:
746:
744:
741:
739:
738:Mark Stephens
735:
730:
728:
722:
720:
716:
711:
709:
705:
701:
692:
687:
681:Personal life
680:
678:
675:
672:
670:
666:
661:
659:
655:
651:
647:
642:
640:
635:
633:
629:
625:
621:
617:
613:
609:
605:
601:
597:
591:
588:
584:
578:
576:
568:
566:
564:
558:
556:
552:
549:(named after
548:
547:Sanger Centre
544:
540:
536:
532:
527:
525:
521:
517:
513:
509:
508:
502:
500:
496:
495:Francis Crick
492:
488:
484:
480:
472:
470:
468:
464:
460:
456:
452:
448:
444:
440:
436:
432:
428:
420:
418:
416:
412:
408:
404:
400:
396:
395:
390:
386:
381:
377:
373:
369:
360:
359:/john-sulston
344:
340:
336:
332:
330:
326:
319:
318:
314:
312:
308:
302:
299:
297:
294:
292:
289:
287:
284:
282:
279:
278:
276:
272:
266:
263:
261:
258:
257:
255:
251:
248:
244:
237:
234:
231:
228:
225:
222:
219:
216:
213:
210:
207:
204:
201:
198:
195:
192:
189:
186:
185:
183:
179:
175:
171:
152:
148:
145:
141:
140:Sulston score
137:
133:
132:
127:
124:
120:
116:
113:
109:
106:
103:
99:
91:
87:
82:
77:27 March 1942
65:
61:
54:
49:
44:
40:
36:
27:
22:
19:
5509:Özlem Türeci
5465:Joanne Chory
5459:Svante Pääbo
5440:Rainer Weiss
5339:Baruch Minke
5335:David Julius
5303:Sumio Iijima
5297:Ginés Morata
5253:Jane Goodall
5217:John Sulston
5216:
5213:Craig Venter
5203:Robert Gallo
5147:Amable Liñán
5059:Alberto Sols
4966:
4895:
4834:Jimmy Carter
4815:Imre Kertész
4703:Svante Pääbo
4688:David Julius
4642:Tasuku Honjo
4590:Edvard Moser
4582:John O'Keefe
4545:posthumously
4544:
4481:Martin Evans
4457:Robin Warren
4441:Richard Axel
4420:
4388:2001–present
4266:Bert Sakmann
4134:Roger Sperry
4126:George Snell
4122:Jean Dausset
4090:Werner Arber
4043:Howard Temin
3967:Bernard Katz
3951:Max Delbrück
3907:Francis Rous
3883:Feodor Lynen
3879:Konrad Bloch
3867:Alan Hodgkin
3851:James Watson
3827:Frank Burnet
3815:Severo Ochoa
3803:Edward Tatum
3791:Daniel Bovet
3704:Philip Hench
3616:Edward Doisy
3547:Hans Spemann
3535:George Minot
3515:Edgar Adrian
3503:Otto Warburg
3425:John Macleod
3394:August Krogh
3386:Jules Bordet
3302:Paul Ehrlich
3254:Niels Finsen
3238:Emil Behring
3175:
3140:Postdoctoral
3097:John Sulston
3096:
3092:David Stuart
3072:Anne Johnson
3010:Mark Walport
2974:James Thomas
2921:Fiona Powrie
2804:
2762:
2579:Vivian Fuchs
2574:Shosaku Numa
2530:
2526:Allan Wilson
2506:John Sulston
2505:
2468:
2451:
2422:Gabriel Horn
2359:
2340:John Argyris
2258:
2229:The Guardian
2227:
2164:
2156:The Guardian
2155:
2145:
2133:. Retrieved
2123:
2111:. Retrieved
2107:the original
2102:
2093:
2081:. Retrieved
2071:
2057:
2032:
2024:
2012:. Retrieved
2003:
1992:
1983:
1973:11 September
1971:. Retrieved
1961:
1947:
1934:
1925:
1914:
1906:the original
1896:
1876:
1869:
1834:
1828:
1815:
1774:
1770:
1754:
1742:. Retrieved
1735:
1728:
1695:
1691:
1687:
1683:
1677:
1647:(2): 51–58.
1644:
1640:
1631:
1606:
1602:
1598:
1592:
1567:
1563:
1560:Kornberg, H.
1553:
1541:. Retrieved
1537:the original
1512:. Retrieved
1507:
1483:The Guardian
1482:
1424:
1420:
1410:
1369:(12): e225.
1366:
1360:
1350:
1338:. Retrieved
1324:
1312:. Retrieved
1301:The Guardian
1300:
1290:
1263:
1259:
1249:
1240:
1230:
1205:
1201:
1195:
1170:
1166:
1160:
1135:
1131:
1087:
1083:
1029:
1025:
1012:
982:
900:
895:
848:
844:
805:. Retrieved
794:the original
767:
742:
731:
723:
715:Humanists UK
712:
696:
676:
673:
667:(CH) in the
662:
643:
638:
636:
607:
593:
582:
580:
572:
562:
559:
538:
535:human genome
530:
528:
523:
505:
503:
491:Leslie Orgel
476:
441:and then to
424:
392:
391:of the worm
367:
366:
315:
274:Institutions
246:
206:Beadle Award
200:Darwin Medal
129:
94:(2018-03-06)
92:6 March 2018
30:John Sulston
18:
5601:2018 deaths
5596:1942 births
5483:Terence Tao
5373:Peter Higgs
5319:Tobin Marks
5263:Tony Hunter
5243:Vinton Cerf
5119:Guido Münch
4762:Nobel Prize
4554:John Gurdon
4469:Craig Mello
4465:Andrew Fire
4378:Eric Kandel
4354:Ferid Murad
4262:Erwin Neher
4174:Niels Jerne
3939:Har Khorana
3927:George Wald
3895:André Lwoff
3863:John Eccles
3751:John Enders
3723:Max Theiler
3684:Walter Hess
3676:Paul Müller
3640:Ernst Chain
3310:Emil Kocher
3270:Robert Koch
3262:Ivan Pavlov
3246:Ronald Ross
3087:David Steel
3082:Peter Rigby
3057:Roger Gibbs
3042:Harold Cook
3012:(2003–2013)
3006:(1998–2008)
3000:(1991–1998)
2994:(1965–1991)
2968:Kathy Poole
2965:Nick Moakes
2764:Big Picture
2511:Jean Thomas
2412:Adrian Gill
2387:John Dowell
2350:Alec Broers
1989:"No. 61962"
1738:sequencing"
1570:: 515–532.
1266:: 421–447.
596:EMBO Member
557:, England.
551:Fred Sanger
522:, so that
469:chemistry.
333:Colin Reese
230:Nobel prize
5590:Categories
5527:Yann LeCun
5505:Uğur Şahin
5475:Yves Meyer
5450:, and the
4802:Literature
4405:Paul Nurse
3739:Hans Krebs
3664:Gerty Cori
3612:Henrik Dam
3559:Otto Loewi
3555:Henry Dale
3142:Fellowship
3134:Fellowship
3077:Roy Porter
3062:Henry Dale
3047:Kay Davies
2977:Ed Whiting
2962:Tim Livett
2950:Alyson Fox
2941:Chris Bird
2902:Alan Brown
2852:UK Biobank
2588:Statute 12
2549:Piet Borst
2437:Allen Kerr
2345:Ian Axford
1684:C. elegans
1340:6 November
1022:Durbin, R.
1018:Wilson, R.
1006:required.)
788:. London:
747:References
708:Magog Down
639:C. elegans
539:C. elegans
531:C. elegans
524:C. elegans
467:nucleotide
449:degree in
73:1942-03-27
5434:Hugh Herr
5176:Atapuerca
4817:(Hungary)
4771:Chemistry
4764:laureates
4606:Tu Youyou
4158:John Vane
4053:1976–2000
3714:1951–1975
3660:Carl Cori
3450:1926–1950
3229:1901–1925
3218:Laureates
2971:Jim Smith
2113:4 October
2083:4 October
1853:0080-4606
1799:0028-0836
1661:0168-9525
1309:0261-3077
1282:186212646
984:Who's Who
587:mutations
260:Chemistry
144:Apoptosis
117:(BA, PhD)
101:Education
83:, England
4917:(Israel)
4401:Tim Hunt
3177:Category
2872:WormBase
2817:Farmcare
2795:DECIPHER
2319:elected
2260:Director
2135:21 April
2014:10 March
1861:86297413
1823:(2015).
1807:11237011
1744:22 April
1669:10098407
1584:73076704
1543:21 April
1514:21 April
1443:14580111
1395:17196043
1334:Archived
1084:Genetics
1062:21450455
987:(online
875:11370623
845:Genetics
817:cite web
700:La Jolla
614:and the
565:(2002).
487:La Jolla
355:/faculty
173:Children
4861:(Japan)
4845:Physics
4787:(Japan)
3220:of the
2812:Ensembl
2652:Current
2542:Foreign
2328:Fellows
2321:in 1986
2315:of the
2313:Fellows
2262:of the
1779:Bibcode
1720:9851916
1700:Bibcode
1692:Science
1434:2588810
1386:1756915
1314:14 July
1222:2838135
1152:6684600
1114:4858229
1105:1213121
1054:7906398
1034:Bibcode
866:1461515
807:9 March
727:Tamiflu
510:at the
481:at the
413:at the
405:at the
353:/people
342:Website
167:
159:
5569:, and
5553:2024:
5547:, and
5539:2023:
5533:, and
5521:2022:
5515:, and
5491:2021:
5485:, and
5473:2020:
5463:2019:
5457:2018:
5438:2017:
5432:2016:
5422:2015:
5393:, and
5385:2014:
5379:, and
5371:2013:
5361:2012:
5355:, and
5347:2011:
5341:, and
5333:2010:
5323:2009:
5317:, and
5301:2008:
5291:2007:
5285:2006:
5279:2005:
5273:, and
5257:2004:
5251:2003:
5245:, and
5233:2002:
5227:, and
5211:2001:
5201:2000:
5191:1999:
5181:1998:
5174:1997:
5168:1996:
5157:1995:
5151:1994:
5145:1993:
5139:1992:
5133:1991:
5123:1990:
5117:1989:
5107:1988:
5097:1987:
5091:1986:
5081:1985:
5075:1984:
5069:1983:
5063:1982:
5057:1981:
4648:2019:
4636:2018:
4620:2017:
4612:2016:
4596:2015:
4580:2014:
4564:2013:
4552:2012:
4531:2011:
4523:2010:
4507:2009:
4491:2008:
4475:2007:
4463:2006:
4451:2005:
4439:2004:
4427:2003:
4411:2002:
4395:2001:
4368:2000:
4360:1999:
4344:1998:
4336:1997:
4324:1996:
4308:1995:
4296:1994:
4284:1993:
4272:1992:
4260:1991:
4248:1990:
4236:1989:
4220:1988:
4212:1987:
4200:1986:
4188:1985:
4172:1984:
4164:1983:
4148:1982:
4132:1981:
4116:1980:
4104:1979:
4088:1978:
4072:1977:
4060:1976:
4033:1975:
4017:1974:
4001:1973:
3989:1972:
3981:1971:
3965:1970:
3949:1969:
3933:1968:
3917:1967:
3905:1966:
3889:1965:
3877:1964:
3861:1963:
3845:1962:
3837:1961:
3825:1960:
3813:1959:
3797:1958:
3789:1957:
3773:1956:
3765:1955:
3749:1954:
3737:1953:
3729:1952:
3721:1951:
3694:1950:
3682:1949:
3674:1948:
3658:1947:
3650:1946:
3634:1945:
3622:1944:
3610:1943:
3581:1939:
3573:1938:
3565:1937:
3553:1936:
3545:1935:
3529:1934:
3521:1933:
3509:1932:
3501:1931:
3493:1930:
3481:1929:
3473:1928:
3465:1927:
3457:1926:
3431:1924:
3419:1923:
3407:1922:
3392:1920:
3384:1919:
3348:1914:
3340:1913:
3332:1912:
3324:1911:
3316:1910:
3308:1909:
3296:1908:
3288:1907:
3276:1906:
3268:1905:
3260:1904:
3252:1903:
3244:1902:
3236:1901:
2837:MEROPS
2780:ChEMBL
2717:Former
2597:
2045:
1884:
1859:
1851:
1805:
1797:
1771:Nature
1718:
1667:
1659:
1623:838129
1621:
1582:
1441:
1431:
1393:
1383:
1307:
1280:
1220:
1187:838129
1185:
1150:
1112:
1102:
1060:
1052:
1026:Nature
915:
873:
863:
514:(MRC)
473:Career
389:genome
347:sanger
322:(1966)
320:
311:Thesis
253:Fields
238:(2002)
232:(2002)
226:(2002)
220:(2001)
214:(2001)
208:(2000)
202:(1996)
196:(1989)
190:(1986)
181:Awards
150:Spouse
136:humans
4967:2002
4826:Peace
4760:2002
2806:eLife
2226:from
1857:S2CID
1767:(PDF)
1580:S2CID
1278:S2CID
1058:S2CID
913:EThOS
622:with
378:
374:
161:(
157:
41:
37:
5467:and
5452:LIGO
5426:and
5365:and
5327:and
5295:and
5205:and
5195:and
5185:and
5161:and
5127:and
5111:and
5101:and
5085:and
4992:2007
4987:2006
4982:2005
4977:2004
4972:2003
4962:2001
4957:2000
4952:1999
4947:1998
4942:1997
4806:2002
4710:2023
4699:2022
4684:2021
4665:2020
3604:1942
3597:1941
3590:1940
3440:1925
3401:1921
3378:1918
3371:1917
3364:1916
3357:1915
2847:Rfam
2842:Pfam
2137:2014
2115:2012
2085:2012
2043:ISBN
2016:2018
1975:2013
1882:ISBN
1849:ISSN
1803:PMID
1795:ISSN
1746:2014
1716:PMID
1682:The
1665:PMID
1657:ISSN
1619:PMID
1545:2014
1516:2014
1508:NNDB
1439:PMID
1391:PMID
1342:2014
1316:2023
1305:ISSN
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