260:
56:
1066:, have not been confirmed. It is rarely encountered, and has been collected in only one location in Rio Grande do Sul, but this may be due to insufficient efforts to locate it, rather than genuine rarity. Its distribution is generally limited to areas with mean winter temperatures over 12 °C (54 °F), mean annual temperatures over 18 °C (64 °F), annual rainfall over 1,100 mm (43 in), and a long rainy season averaging over 200 days. It is usually found in swamps or near streams.
120:
95:
840:, which occurs in the same area, the tail is longer than the head and body. It is sparsely haired and dark, and there is no difference in color between the upper and lower side. The coat, which is long, dense, and soft, is yellow–brown at the sides, but becomes darker on the upperparts and lighter on the underparts. The large hindfeet are characterized by conspicuous
960:. The main cusps are located opposite each other and have rounded edges. The enamel folds do not extend past the midlines of the molars. The mesoloph, an accessory crest on the upper molars that is usually well-developed in oryzomyines, is present but short on the first and second upper molar; it is much more reduced in
1012:, the first upper molar lacks an accessory fourth root. The first lower molar has four roots, including two small accessory roots located between larger anterior and posterior roots. The second molar has either two or three roots, with the anterior root split into two smaller roots in some specimens.
1167:
in some features of the first lower molar and may represent a distinct species. One lower first molar of this form has length 3.28 mm. Because the Bajo San José material does not contain lower first molars, it is impossible to determine whether this material also pertains to the later
Argentine
1205:
among reeds in up to 1.5 m (4.9 ft) deep water, usually about 20 cm (8 in) above the water. The material for the nest, which is 25 to 30 cm (10 to 12 in) in diameter and 9 to 11 cm (about 4 in) in height, comes from the surrounding reeds. Its wall consists of
868:
is absent, both important characters of oryzomyines. The head and body length is 160 to 230 mm (6.3 to 9.1 in), averaging 193 mm (7.6 in), the tail length is 195 to 255 mm (7.68 to 10.04 mm), averaging 235 mm (9.3 in), and the length of the hindfoot is 58 to
629:. When American zoologists Voss and Carleton restudied Winge's material in a 1993 paper, they were unable to find any consistent differences between the two and accordingly considered them to pertain to the same species. In addition, they reviewed the differences between this species and other
401:, with the head and body length averaging 193 mm (7.6 in), characterized by a long tail, large hindfeet, and long and dense fur. It builds nests above the water, supported by reeds, and it is not currently threatened.
904:
and the upper molars, are long, extending between the molars. The palate itself is also long, extending beyond the posterior margin of the maxillary bones, and it is perforated near the third molars by conspicuous
1182:. The length of the upper toothrow of one specimen from this locality is 8.50 mm (0.335 in) and the length of the upper first molar is 3.48 mm (0.137 in), slightly smaller than in living
1116:; there, it is known only from three skull fragments from a cave known as Laga da Escrivania Nr. 5. This cave also contains numerous remains of members of the extinct South American
860:
margins and between some of the digits. The forefeet also lack tufts on the digits and show very long claws, a character unique among oryzomyines. The female has four pairs of
2085:
Sigmodontinos (Mammalia, Rodentia) Pleistocenicos del sudoeste de la provincia de Buenos Aires (Argentina): Aspectos sistematicos, paleozoogeograficos y paleoambientales
1197:
is semiaquatic in habits, spending much of its time in the water, and is active during the night. An excellent swimmer, it is even more specialized for swimming than is
1905:
Amerigo
Vespucci and the rat of Fernando de Noronha: a new genus and species of Rodentia (Muridae, Sigmodontinae) from a volcanic island off Brazil's continental shelf
741:
has been suggested as an alternative. A comprehensive phylogenetic analysis of oryzomyines by
Marcelo Weksler, published in 2006, supported a close relationship among
3921:
1287:, reflecting a relatively wide distribution and the absence of evidence for a decline in populations. Several of the areas where it occurs are protected, but the
1284:
4085:
3986:
4080:
1206:
three layers, surrounding a central chamber, which is connected to the water by a ramp, also composed of reeds. Nests built by members of the related genus
945:, to which some of the chewing muscles are attached, are entirely separate, joining only at their anterior edges, which are located below the first molar.
4115:
2096:
Cricetidos (Mammalia: Rodentia) del
Pleistoceno tardio del nordeste de la provincia de Buenos Aires (Argentina). Aspectos sistematicos y paleoambientales
1307:
Measurements for head and body length and tail length are from 10 specimens, and those for hindfoot length are from 12 specimens, all from
Uruguay.
1159:, about 420,000 to 360,000 years ago, while younger specimens from other localities are as little as 30,000 years old. The younger Argentine
4050:
3895:
2222:
3934:
2191:
Phylogenetic relationships of oryzomyine rodents (Muroidea: Sigmodontinae): separate and combined analyses of morphological and molecular data
1893:
2029:
941:
of the lower incisor, a slight raising of the mandibular bone at the back end of the incisor, near the coronoid process, is small. The two
523:. Subsequently, it was rarely mentioned in the literature on South American rodents; those authors who did mention it placed it in either
4110:
1923:
1994:
Primer registro de ácaros (Gamasida: Laelapidae) parásitos de roedores silvestres en
Uruguay, con nuevos registros de hospedadores.
4100:
3960:
930:
549:
3978:
2021:
564:
found across much of South
America, and another unique to Uruguay and southern Brazil; he named the latter as a new species,
3939:
2116:
Ray, C.E. 1962. The
Oryzomyine Rodents of the Antillean Subregion. Doctor of Philosophy thesis, Harvard University, 211 pp.
2147:
Late
Pleistocene vertebrates from northern Uruguay: tools for biostratigraphic, climatic and environmental reconstruction
1992:
Lareschi, M., Gettinger, D., Venzal, J.M., Arzua, M., Nieri-Bastos, F.A., Barros-Battesti, D.M. and
Gonzalez, E.M. 2006.
2215:
952:) than in most oryzomyines, and many of the accessory crests are reduced, but they are sharply distinct from the highly
925:
is unusual in that the squamosal and the tegmen tympani usually overlap when viewed from the side. In the mandible, the
2084:
4095:
3973:
1993:
495:(lower jaw) from Lapa da Serra das Abelhas, but the latter later turned out to be from a different species, probably
2105:
Pardiñas, U.F.J., D'Elía, G. and Teta, P. 2008. Una introducción a los mayores sigmodontinos vivientes: Revisión de
2062:
2007:
729:
were discovered, providing additional evidence for this grouping. The allocation of the former, which is similar to
4090:
3991:
3421:
906:
2046:
Nava, S., Venzal, J.M., Labruna, M.B., Mastropaolo, M., González, E.M., Mangold, A.J. and Guglielmone, A.A. 2010.
3752:
3428:
1222:. A female caught in April was pregnant with three embryos, which were about 12 mm (0.47 in) long. The
484:
119:
968:. The corresponding structure on the lower molars, the mesolophid, is present on the first and second molars in
271:
in South America. The current distribution is in red, and fossil records outside the current range are in blue.
3745:
2534:
2419:
2298:
1227:
1156:
819:
2063:
Paleontologia e aspectos geológicos das sucessões do final do Neógeno no sudoeste do Rio Grande do Sul, Brasil
3774:
3701:
3463:
3407:
3393:
3386:
3379:
3178:
3171:
3098:
2877:
2641:
2208:
1929:
Freitas, T.R.O., Mattevi, M.S., Oliveira, L.F.B., Souza, M.J., Yonenaga-Yassuda, Y. and Salzano, F.M. 1983.
1109:
836:
410:
656:
valid way. Previously, Oryzomyini had been a somewhat loosely defined group defined among others by a long
633:
and concluded that these were significant enough to place the former in a distinct genus, which they named
3822:
3456:
3414:
3164:
3049:
2691:
2619:
2305:
1072:
fossils have been found throughout its current range and beyond it. In Uruguay and Rio Grande do Sul, the
537:
2095:
1233:
209:
3999:
3781:
3449:
3435:
3365:
3126:
3105:
3056:
2819:
2781:
2612:
2598:
2548:
2513:
1249:
1174:
1148:
830:
719:
607:
232:
3860:
487:, Brazil. Winge used four specimens for his description, including two skull fragments and an isolated
418:
history it has been confused with that species, but other features support its placement in a distinct
942:
3869:
3694:
3672:
3470:
3442:
3400:
3372:
3358:
3329:
3133:
3063:
2884:
2826:
2773:
2705:
2648:
2605:
2591:
2584:
2577:
2541:
2527:
2440:
2412:
841:
824:
770:
560:
in 1955, he recognized that the series from Uruguay contained two species, one close to the forms of
497:
405:
2180:(Rodentia: Muridae) and the significance of Lund's collections from the caves of Lagoa Santa, Brazil
4105:
3573:
3544:
3530:
3522:
3478:
3185:
3156:
3119:
2789:
2655:
2570:
2426:
1969:
1288:
1267:
1243:
1152:
893:
84:
2125:(Rodentia: Sigmodontinae) from the Middle Pleistocene of Bolivia and its phylogenetic significance
3336:
3245:
3231:
3091:
3027:
2765:
2719:
2684:
2520:
2405:
1255:
1086:
1024:
781:
closer together, as did the combined analysis of both morphological and IRPB data. Subsequently,
553:
480:
415:
306:
276:
114:
4004:
2146:
3926:
637:
after Lund, who had collected the original material. Since then, the species has been known as
613:
In a 1980 article, Argentine zoologist Elio Massoia recognized the resemblance between Winge's
3882:
3665:
3315:
3266:
3112:
3084:
3077:
3070:
3013:
2999:
2977:
2970:
2833:
2698:
2677:
2433:
2074:
A new genus of oryzomyine rodent (Cricetidae: Sigmodontinae) from the Pleistocene of Argentina
2035:
2025:
1919:
1059:
378:
4014:
1515:
Weksler, 2006, table 8; Voss and Myers, 1991, table 1; Ray, 1962, tables 7, 11
3559:
3551:
3322:
3287:
3280:
3259:
3006:
2956:
2949:
2942:
2935:
2928:
2855:
2845:
2797:
2447:
2015:
1964:
938:
897:
603:
3952:
3537:
3308:
3301:
3273:
3252:
3238:
3020:
2963:
2712:
2200:
1900:
11:1–120. Rio de Janeiro: Centro Pan-Americano de Febre Aftosa – OPAS/OMS (in Portuguese).
1077:
926:
918:
877:
853:
393:. The Argentine form may have been distinct from the living form from Brazil and Uruguay.
17:
3874:
1151:, Argentina, which suggests a warm and humid paleoclimate there. The oldest deposits, at
2073:
1978:
55:
3792:
3735:
3565:
1133:
1047:
is variable, ranging from nearly metacentric to acrocentric in five specimens studied.
1001:
914:
881:
1946:
1140:
rodents, but it is not certain that all remains from this cave are from the same age.
259:
4074:
3947:
3684:
3619:
3609:
2359:
2349:
2277:
2011:
1955:
1894:
Guia dos Roedores do Brasil, com chaves para gêneros baseadas em characteres externos
1261:
861:
653:
591:
513:
460:
433:
367:
191:
104:
99:
1186:, which ranges from 3.56 to 3.64 mm (0.140 to 0.143 in) in four specimens
610:, Argentina, and in 1982 it was recorded from Rio Grande do Sul in southern Brazil.
4041:
3887:
3764:
3348:
2867:
2503:
2491:
2481:
2383:
2372:
2337:
2329:
2318:
2134:
Teta, P. and Pardiñas, U.F.J. 2006. "Pleistocene record of marsh rats of the genus
1121:
1098:
1063:
1044:
1040:
953:
783:
698:
677:
476:
454:
386:
249:
2166:
Hershkovitz (Mammalia, Muridae) with an analysis of its phylogenetic relationships
753:; the other species of the group were not included. Data from the sequence of the
1283:
The species' conservation status is currently assessed as "least concern" by the
572:
as one of the members of a "sigmodont" group of American rodents, also including
3965:
3908:
3854:
3500:
3490:
3294:
3145:
2560:
2469:
2459:
1129:
1069:
1032:
1028:
997:
865:
845:
359:
41:
3845:
2047:
1172:
form. The morphology of the upper and lower jaw precludes an identification as
884:, is expansive and produced into a spinous process at the anterior margin. The
3803:
3723:
3713:
3597:
3586:
3209:
3198:
3039:
2631:
2395:
2288:
2271:
2232:
2003:
1137:
1125:
1036:
1020:
981:
808:
725:
702:—which also lack complete mesoloph(id)s has been universally supported since.
665:
649:
580:
574:
507:
448:
442:
437:
429:
181:
806:
is among the largest living oryzomyines, rivaled only by some large forms of
2809:
2754:
2667:
2039:
1215:
1117:
1092:
1016:
949:
910:
595:
390:
131:
63:
2109:
Hershkovitz, 1966 y descripción de un nuevo género (Rodentia: Cricetidae).
705:
Voss and Carleton had found some support for a close relationship between
4035:
3839:
3655:
3643:
3632:
3512:
2989:
2918:
2906:
2896:
2253:
1073:
814:
661:
586:
525:
492:
151:
75:
45:
4055:
3900:
3221:
1930:
1211:
1104:
901:
857:
519:
488:
374:
363:
67:
491:(upper jaw) from the cave chamber Lapa da Escrivania Nr. 5 and a
3913:
2265:
2259:
2247:
1084:, in addition to such other mammals as the extinct saber-toothed cat
869:
68 mm (2.3 to 2.7 in), averaging 62 mm (2.4 in).
657:
398:
382:
171:
161:
141:
3816:
2120:
1062:; records of live specimens from eastern Argentina and Lagoa Santa,
1039:
pairs, which have two major arms, and a small metacentric pair. The
818:, but it is substantially smaller than some of the recently extinct
2190:
2157:
1904:
1604:
Voss and Carleton, 1993, p. 16; Weksler, 2006, pp. 34–35
1178:, a fossil species with molar traits almost identical to those of
885:
648:, Voss and Carleton also, for the first time, diagnosed the tribe
621:, and recommended that the former be reclassified as a species of
548:) in his 1929 report on the collection. When his successor at the
419:
2176:
2017:
Mammal Species of the World: A Taxonomic and Geographic Reference
1556:
Voss and Carleton, 1993, p. 7; Weksler, 2006, pp. 24–25
1027: = 58). The non-sex chromosomes (autosomes) are mostly
1661:
Voss and Carleton, 1993, p. 20; Weksler, 2006, fig. 25
1223:
1202:
1112:, Lagoa Santa, lies far northeast of the nearest record of live
1000:, contains a deep pit. Each of the three upper molars has three
754:
3820:
2204:
1931:
Chromosome relationships in three representatives of the genus
1241:
in Uruguay. Other rodents found in association with it include
1670:
Voss and Carleton, 1993, p. 20; Weksler, 2006, p. 49
1622:
Weksler, 2006, p. 40; Voss and Carleton, 1993, p. 17
849:
1547:
Voss and Carleton, 1993, p. 6; Weksler, 2006, p. 23
1058:
has been found as a living animal only in Uruguay and nearby
717:
within Oryzomyini. In subsequent years, the related species
2145:
Ubilla, M., Perea, D., Aguilar, C.G. and Lorenzo, N. 2004.
540:
collected some rodents in Uruguay, which he identified as
27:
A semiaquatic rat species from southeastern South America.
2048:
Hosts, distribution and genetic divergence (16S rDNA) of
1892:
Bonvicino, C.R., Oliveira, J.A. and D'Andrea, P.S. 2008.
1715:
Freitas et al., 1983; Voss and Carleton, 1993, p. 10
2138:
in southern South America: Paleoclimatic significance".
1844:
Sierra de Soriano in Voss and Carleton, 1993, p. 34
1362:
González et al., 2016; Duff and Lawson, 2004, p. 56
984:, is present, though small, on the first upper molar in
668:
on the lower molars; this crest is absent or reduced in
1023:
with a total of 58 major arms (2n = 52,
852:
are reduced. As in some other semiaquatic oryzomyines,
737:
is controversial, and placement as a second species of
1871:
Lareschi et al., 2006; Nava et al., 2010, table 1
1214:
contained green plant material, suggesting that it is
1035:, having only one arm, but there are also three large
896:
of the skull is narrow and flanked by high beads. The
1574:
Voss and Carleton, 1993; Weksler, 2006, table 5
872:
The front part of the skull is notably broad. As in
598:(the crown consists of transverse ridges). In 1981,
4025:
3829:
3791:
3762:
3733:
3711:
3682:
3653:
3629:
3607:
3583:
3510:
3488:
3346:
3219:
3195:
3143:
3037:
2987:
2916:
2894:
2865:
2843:
2807:
2751:
2729:
2665:
2629:
2558:
2501:
2479:
2457:
2393:
2369:
2347:
2315:
2286:
2195:Bulletin of the American Museum of Natural History
2184:Bulletin of the American Museum of Natural History
996:, the anterior cusp on the first lower molar, the
988:, but entirely absent in both other genera. As in
913:bone lacks a suspensory process that contacts the
1147:have been found at six Pleistocene localities in
1945:Gonzalez, E.; D'elia, G.; Pardinas, U. (2017) .
475:was first described in 1888 by Danish zoologist
1751:Voss and Myers, 1991, table 5, p. 429
1349:
1347:
1210:are similar in many details. Several dissected
1970:10.2305/IUCN.UK.2016-3.RLTS.T10219A22328078.en
1742:Oliveira and Kerber, 2009; Ubilla et al., 2004
1291:may pose a threat to its continued existence.
1285:International Union for Conservation of Nature
2216:
1862:Tuttle in Voss and Carleton, 1993, p. 32
1853:Barley in Voss and Carleton, 1993, p. 34
1724:González et al., 2016; Bonvicino et al., 2008
757:gene supported a closer relationship between
8:
1778:Pardiñas and Lezcano, 1995, pp. 258–259
1918:. New Haven: A & C Black, 312 pp.
948:The molars are slightly more high-crowned (
787:was described as an additional relative of
436:, it is a member of a group of specialized
3817:
2223:
2209:
2201:
1796:Pardiñas and Deschamps, 1995, table 2
1163:specimens are subtly distinct from living
680:for the group, all of which are shared by
644:In the same paper in which they described
385:, but it previously ranged northward into
258:
93:
54:
31:
2094:Pardiñas, U.F.J. and Lezcano, M.J. 1995.
2083:Pardiñas, U.F.J. and Deschamps, C. 1996.
1968:
1787:Pardiñas and Deschamps, 1995, p. 850
900:, which perforate the palate between the
660:and the presence of a crest known as the
1461:Musser and Carleton, 2005; Weksler, 2006
888:bone is small, but less reduced than in
1880:Voss and Carleton, 1993, pp. 32–34
1813:
1811:
1648:
1646:
1591:
1589:
1479:Steppan, 1996; Carleton and Olson, 1999
1439:
1437:
1413:Oliveira et al. in Freitas et al., 1983
1380:Pardiñas et al., 2008, pp. 556–557
1353:Musser and Carleton, 2005, p. 1124
1319:
1300:
1583:Voss and Carleton, 1993, table 2.
1534:
1532:
1530:
1422:Voss and Carleton, 1993, pp. 3, 6
1400:
1398:
1325:
1323:
373:Its distribution is now restricted to
1983:, with a summary of sigmodont rodents
1903:Carleton, M.D. and Olson, S.L. 1999.
1388:
1386:
1337:
1335:
1076:(Late Pleistocene to Early Holocene)
880:, the flattened front portion of the
414:, and over the course of its complex
389:, Brazil, and southward into eastern
7:
4086:Extant Pleistocene first appearances
2156:Voss, R.S. and Carleton, M.D. 1993.
2061:Oliveira, É.V. and Kerber, L. 2009.
1826:Carleton and Olson, 1999, p. 52
1769:Teta and Pardiñas, 2006, p. 180
1733:Teta and Pardiñas, 2006, p. 179
1136:, in addition to 16 species of
1031:, having a long and a short arm, or
676:. Voss and Carleton recognized five
4081:IUCN Red List least concern species
2140:Current Research in the Pleistocene
1956:IUCN Red List of Threatened Species
1817:Voss and Carleton, 1993, p. 34
1688:Voss and Carleton, 1993, p. 20
1652:Voss and Carleton, 1993, p. 19
1631:Voss and Carleton, 1993, p. 17
1613:Voss and Carleton, 1993, p. 16
1595:Voss and Carleton, 1993, p. 15
1524:Voss and Carleton, 1993, p. 13
1443:Voss and Carleton, 1993, p. 31
1404:Voss and Carleton, 1993, p. 10
590:, on the basis of its flat-crowned
2129:Journal of Vertebrate Paleontology
2056:Experimental and Applied Acarology
1538:Voss and Carleton, 1993, p. 7
1470:Voss and Carleton, 1993, p. 1
1431:Voss and Carleton, 1993, p. 4
1392:Voss and Carleton, 1993, p. 3
1371:Voss and Carleton, 1993, p. 6
1341:Voss and Carleton, 1993, p. 5
25:
4116:Taxa named by Michael D. Carleton
1979:South American marsh rats, genus
1933:Holochilus (Rodentia, Cricetidae)
1916:Mammals of the World: A checklist
848:on the digits and several of the
733:in features of the dentition, to
684:; the placement in Oryzomyini of
505:and placed it in the same genus (
1237:have been found on specimens of
933:are less well-developed than in
118:
78:represents a different species.
2175:Voss, R.S. and Myers, P. 1991.
980:. Another accessory crest, the
550:Field Museum of Natural History
2022:Johns Hopkins University Press
1914:Duff, A. and Lawson, A. 2004.
1835:Hershkovitz, 1955, p. 658
1565:Weksler, 2006, pp. 19, 23
483:had collected in the caves of
1:
1697:Weksler, 2006, pp. 42–43
909:. As in all oryzomyines, the
479:, who reviewed the materials
1805:Pardiñas, 2008, table 1
1488:Pardiñas, 2008, p. 1275
769:more distantly related, but
536:In 1926, American zoologist
2102:32(3):249–265 (in Spanish).
1999:35(5):596–601 (in Spanish).
907:posterolateral palatal pits
602:was also recognized in the
440:rodents that also includes
4132:
2111:Arquivos do Museu Nacional
2006:; Carleton, M. D. (2005).
1289:destruction of its habitat
688:and of three other genera—
18:Lund's Amphibious Rat
4111:Mammals described in 1888
2242:
2170:American Museum Novitates
2149:(subscription required).
2127:(subscription required).
2076:(subscription required).
2054:(subscription required).
1937:(subscription required).
1909:American Museum Novitates
1898:Série de Manuais Técnicos
1706:Weksler, 2006, p. 43
1679:Weksler, 2006, p. 45
1640:Weksler, 2006, p. 47
1228:Gigantolaelaps wolffsohni
568:. Hershkovitz identified
485:Lagoa Santa, Minas Gerais
282:
275:
266:
257:
238:
231:
115:Scientific classification
113:
91:
82:
62:
53:
34:
2151:Quaternary International
2091:52:367–379 (in Spanish).
2069:5(1):21–34 (in Spanish).
2014:; Reeder, D. M. (eds.).
1977:Hershkovitz, P.M. 1955.
1201:. It builds a spherical
1157:Marine Isotopic Stage 11
1051:Distribution and ecology
664:on the upper molars and
501:. Lund named the animal
4101:Monotypic rodent genera
2072:Pardiñas, U.F.J. 2008.
1760:Teta and Pardiñas, 2006
1452:Voss and Carleton, 1993
1273:Holochilus brasiliensis
1080:has yielded remains of
1043:is metacentric and the
837:Holochilus brasiliensis
546:Holochilus brasiliensis
411:Holochilus brasiliensis
336:Voss and Carleton, 1993
2008:"Superfamily Muroidea"
1997:Neotropical Entomology
1963:: e.T10219A115096149.
956:, hypsodont molars of
856:are present along the
538:Colin Campbell Sanborn
408:is similar to that of
4000:Paleobiology Database
2178:Pseudoryzomys simplex
1329:González et al., 2016
1250:Oligoryzomys nigripes
1175:Holochilus primigenus
1149:Buenos Aires Province
972:, but absent in both
831:Megalomys desmarestii
720:Holochilus primigenus
608:Buenos Aires Province
514:Pseudoryzomys simplex
352:Lund's amphibious rat
40:Temporal range: Late
3804:Ekbletomys hypenemus
2338:A. praeuniversitatis
2119:Steppan, S.J. 1996.
2078:Journal of Mammalogy
842:interdigital webbing
825:Ekbletomys hypenemus
726:Noronhomys vespuccii
498:Gyldenstolpia fronto
428:. Within the family
2089:Estudios Geológicos
2050:Amblyomma dubitatum
1279:Conservation status
1268:Oxymycterus nasutus
1244:Scapteromys tumidus
1234:Amblyomma dubitatum
894:interorbital region
784:Carletonomys cailoi
542:Holochilus vulpinus
517:and two species of
85:Conservation status
4096:Mammals of Uruguay
4027:Hesperomys molitor
2189:Weksler, M. 2006.
2160:Hesperomys molitor
1256:Reithrodon auritus
1087:Smilodon populator
931:coronoid processes
917:, the roof of the
627:Holochilus molitor
617:and Hershkovitz's
615:Hesperomys molitor
554:Philip Hershkovitz
503:Hesperomys molitor
481:Peter Wilhelm Lund
366:from southeastern
323:Holochilus molitor
286:Hesperomys molitor
74:. The illustrated
4091:Rodents of Brazil
4066:
4065:
3823:Taxon identifiers
3814:
3813:
2236:
2231:Species of tribe
2164:Holochilus magnus
2121:A new species of
2052:(Acari: Ixodidae)
2031:978-0-8018-8221-0
1987:Fieldiana Zoology
1060:Rio Grande do Sul
1019:contains 52
1004:; unlike in both
943:masseteric ridges
898:incisive foramina
820:Antillean species
619:Holochilus magnus
566:Holochilus magnus
511:) as what is now
379:Rio Grande do Sul
356:greater marsh rat
343:
342:
337:
328:
319:
318:Hershkovitz, 1962
310:
303:Holochilus magnus
299:
290:
213:
108:
16:(Redirected from
4123:
4059:
4058:
4046:
4045:
4044:
4018:
4017:
4008:
4007:
3995:
3994:
3982:
3981:
3969:
3968:
3956:
3955:
3943:
3942:
3930:
3929:
3917:
3916:
3904:
3903:
3891:
3890:
3878:
3877:
3875:Lundomys_molitor
3865:
3864:
3863:
3861:Lundomys molitor
3850:
3849:
3848:
3831:Lundomys molitor
3818:
3422:O. longicaudatus
2846:Microakodontomys
2470:D. albimaculatus
2420:C. maracajuensis
2299:A. galapagoensis
2234:
2225:
2218:
2211:
2202:
2158:A new genus for
2113:66(3–4):509–594.
2080:89(5):1270–1278.
2043:
2024:. p. 1124.
2020:(3rd ed.).
1974:
1972:
1949:Lundomys molitor
1887:Literature cited
1881:
1878:
1872:
1869:
1863:
1860:
1854:
1851:
1845:
1842:
1836:
1833:
1827:
1824:
1818:
1815:
1806:
1803:
1797:
1794:
1788:
1785:
1779:
1776:
1770:
1767:
1761:
1758:
1752:
1749:
1743:
1740:
1734:
1731:
1725:
1722:
1716:
1713:
1707:
1704:
1698:
1695:
1689:
1686:
1680:
1677:
1671:
1668:
1662:
1659:
1653:
1650:
1641:
1638:
1632:
1629:
1623:
1620:
1614:
1611:
1605:
1602:
1596:
1593:
1584:
1581:
1575:
1572:
1566:
1563:
1557:
1554:
1548:
1545:
1539:
1536:
1525:
1522:
1516:
1513:
1507:
1504:
1498:
1495:
1489:
1486:
1480:
1477:
1471:
1468:
1462:
1459:
1453:
1450:
1444:
1441:
1432:
1429:
1423:
1420:
1414:
1411:
1405:
1402:
1393:
1390:
1381:
1378:
1372:
1369:
1363:
1360:
1354:
1351:
1342:
1339:
1330:
1327:
1308:
1305:
1195:Lundomys molitor
1056:Lundomys molitor
939:capsular process
844:, but they lack
804:Lundomys molitor
654:phylogenetically
639:Lundomys molitor
604:Late Pleistocene
473:Lundomys molitor
350:, also known as
347:Lundomys molitor
335:
332:Lundomys molitor
326:
317:
305:
298:Trouessart, 1898
297:
294:Oryzomys molitor
288:
269:Lundomys molitor
267:Distribution of
262:
244:
242:Lundomys molitor
207:
123:
122:
102:
97:
96:
58:
32:
21:
4131:
4130:
4126:
4125:
4124:
4122:
4121:
4120:
4071:
4070:
4067:
4062:
4054:
4049:
4040:
4039:
4034:
4021:
4013:
4011:
4003:
3998:
3990:
3985:
3977:
3972:
3964:
3959:
3951:
3946:
3938:
3933:
3925:
3920:
3912:
3907:
3899:
3894:
3886:
3881:
3873:
3868:
3859:
3858:
3853:
3844:
3843:
3838:
3825:
3815:
3810:
3787:
3758:
3729:
3707:
3678:
3649:
3625:
3603:
3579:
3506:
3484:
3429:O. magellanicus
3342:
3330:O. sydandersoni
3215:
3191:
3139:
3099:N. maculiventer
3033:
2983:
2912:
2890:
2861:
2856:M. transitorius
2839:
2803:
2747:
2725:
2692:H. megacephalus
2661:
2642:H. brasiliensis
2625:
2554:
2497:
2475:
2453:
2389:
2365:
2343:
2311:
2282:
2238:
2229:
2032:
2002:
1944:
1889:
1884:
1879:
1875:
1870:
1866:
1861:
1857:
1852:
1848:
1843:
1839:
1834:
1830:
1825:
1821:
1816:
1809:
1804:
1800:
1795:
1791:
1786:
1782:
1777:
1773:
1768:
1764:
1759:
1755:
1750:
1746:
1741:
1737:
1732:
1728:
1723:
1719:
1714:
1710:
1705:
1701:
1696:
1692:
1687:
1683:
1678:
1674:
1669:
1665:
1660:
1656:
1651:
1644:
1639:
1635:
1630:
1626:
1621:
1617:
1612:
1608:
1603:
1599:
1594:
1587:
1582:
1578:
1573:
1569:
1564:
1560:
1555:
1551:
1546:
1542:
1537:
1528:
1523:
1519:
1514:
1510:
1505:
1501:
1496:
1492:
1487:
1483:
1478:
1474:
1469:
1465:
1460:
1456:
1451:
1447:
1442:
1435:
1430:
1426:
1421:
1417:
1412:
1408:
1403:
1396:
1391:
1384:
1379:
1375:
1370:
1366:
1361:
1357:
1352:
1345:
1340:
1333:
1328:
1321:
1317:
1312:
1311:
1306:
1302:
1297:
1281:
1239:L. molitor
1192:
1190:Natural history
1090:and species of
1078:Sopas Formation
1053:
919:tympanic cavity
878:zygomatic plate
854:fringes of hair
801:
470:
395:L. molitor
314:Calomys molitor
253:
246:
240:
227:
224:L. molitor
206:
117:
109:
98:
94:
87:
49:
38:
28:
23:
22:
15:
12:
11:
5:
4129:
4127:
4119:
4118:
4113:
4108:
4103:
4098:
4093:
4088:
4083:
4073:
4072:
4064:
4063:
4061:
4060:
4047:
4031:
4029:
4023:
4022:
4020:
4019:
4009:
3996:
3983:
3970:
3957:
3944:
3931:
3918:
3905:
3892:
3879:
3866:
3851:
3835:
3833:
3827:
3826:
3821:
3812:
3811:
3809:
3808:
3797:
3795:
3793:Incertae sedis
3789:
3788:
3786:
3785:
3778:
3770:
3768:
3760:
3759:
3757:
3756:
3749:
3741:
3739:
3736:Transandinomys
3731:
3730:
3728:
3727:
3719:
3717:
3709:
3708:
3706:
3705:
3698:
3690:
3688:
3680:
3679:
3677:
3676:
3673:S. ucayalensis
3669:
3661:
3659:
3651:
3650:
3648:
3647:
3638:
3636:
3627:
3626:
3624:
3623:
3615:
3613:
3605:
3604:
3602:
3601:
3592:
3590:
3581:
3580:
3578:
3577:
3570:
3563:
3555:
3548:
3541:
3534:
3526:
3518:
3516:
3508:
3507:
3505:
3504:
3496:
3494:
3486:
3485:
3483:
3482:
3474:
3467:
3460:
3453:
3446:
3439:
3432:
3425:
3418:
3411:
3404:
3397:
3390:
3383:
3376:
3369:
3362:
3354:
3352:
3344:
3343:
3341:
3340:
3333:
3326:
3319:
3312:
3305:
3298:
3291:
3284:
3277:
3270:
3263:
3256:
3249:
3242:
3235:
3227:
3225:
3217:
3216:
3214:
3213:
3204:
3202:
3193:
3192:
3190:
3189:
3182:
3179:N. narboroughi
3175:
3172:N. fernandinae
3168:
3160:
3151:
3149:
3141:
3140:
3138:
3137:
3130:
3123:
3116:
3109:
3102:
3095:
3088:
3081:
3074:
3067:
3060:
3053:
3050:N. albigularis
3045:
3043:
3035:
3034:
3032:
3031:
3024:
3017:
3010:
3003:
2995:
2993:
2985:
2984:
2982:
2981:
2974:
2967:
2960:
2953:
2946:
2939:
2932:
2924:
2922:
2914:
2913:
2911:
2910:
2902:
2900:
2892:
2891:
2889:
2888:
2881:
2873:
2871:
2863:
2862:
2860:
2859:
2851:
2849:
2841:
2840:
2838:
2837:
2830:
2823:
2820:M. caliginosus
2815:
2813:
2805:
2804:
2802:
2801:
2793:
2785:
2782:M. desmarestii
2777:
2769:
2760:
2758:
2749:
2748:
2746:
2745:
2737:
2735:
2727:
2726:
2724:
2723:
2716:
2709:
2702:
2695:
2688:
2681:
2673:
2671:
2663:
2662:
2660:
2659:
2652:
2645:
2637:
2635:
2627:
2626:
2624:
2623:
2616:
2609:
2602:
2595:
2588:
2581:
2574:
2566:
2564:
2556:
2555:
2553:
2552:
2545:
2538:
2535:E. macconnelli
2531:
2524:
2517:
2509:
2507:
2499:
2498:
2496:
2495:
2487:
2485:
2477:
2476:
2474:
2473:
2465:
2463:
2455:
2454:
2452:
2451:
2444:
2437:
2430:
2423:
2416:
2409:
2401:
2399:
2391:
2390:
2388:
2387:
2378:
2376:
2367:
2366:
2364:
2363:
2355:
2353:
2345:
2344:
2342:
2341:
2333:
2324:
2322:
2313:
2312:
2310:
2309:
2306:A. xanthaeolus
2302:
2294:
2292:
2284:
2283:
2281:
2280:
2274:
2268:
2262:
2256:
2250:
2243:
2240:
2239:
2230:
2228:
2227:
2220:
2213:
2205:
2199:
2198:
2187:
2173:
2154:
2143:
2132:
2131:16(3):522–530.
2117:
2114:
2103:
2092:
2081:
2070:
2059:
2058:51(4):335–351.
2044:
2030:
2000:
1990:
1975:
1942:
1927:
1912:
1901:
1888:
1885:
1883:
1882:
1873:
1864:
1855:
1846:
1837:
1828:
1819:
1807:
1798:
1789:
1780:
1771:
1762:
1753:
1744:
1735:
1726:
1717:
1708:
1699:
1690:
1681:
1672:
1663:
1654:
1642:
1633:
1624:
1615:
1606:
1597:
1585:
1576:
1567:
1558:
1549:
1540:
1526:
1517:
1508:
1506:Pardiñas, 2008
1499:
1490:
1481:
1472:
1463:
1454:
1445:
1433:
1424:
1415:
1406:
1394:
1382:
1373:
1364:
1355:
1343:
1331:
1318:
1316:
1313:
1310:
1309:
1299:
1298:
1296:
1293:
1280:
1277:
1191:
1188:
1052:
1049:
915:tegmen tympani
800:
797:
678:synapomorphies
469:
466:
432:and subfamily
341:
340:
339:
338:
329:
320:
311:
300:
291:
280:
279:
273:
272:
264:
263:
255:
254:
247:
236:
235:
229:
228:
221:
219:
215:
214:
199:
195:
194:
189:
185:
184:
179:
175:
174:
169:
165:
164:
159:
155:
154:
149:
145:
144:
139:
135:
134:
129:
125:
124:
111:
110:
92:
89:
88:
83:
80:
79:
60:
59:
51:
50:
39:
26:
24:
14:
13:
10:
9:
6:
4:
3:
2:
4128:
4117:
4114:
4112:
4109:
4107:
4104:
4102:
4099:
4097:
4094:
4092:
4089:
4087:
4084:
4082:
4079:
4078:
4076:
4069:
4057:
4052:
4048:
4043:
4037:
4033:
4032:
4030:
4028:
4024:
4016:
4010:
4006:
4001:
3997:
3993:
3988:
3984:
3980:
3975:
3971:
3967:
3962:
3958:
3954:
3949:
3945:
3941:
3936:
3932:
3928:
3923:
3919:
3915:
3910:
3906:
3902:
3897:
3893:
3889:
3884:
3880:
3876:
3871:
3867:
3862:
3856:
3852:
3847:
3841:
3837:
3836:
3834:
3832:
3828:
3824:
3819:
3807:
3805:
3799:
3798:
3796:
3794:
3790:
3784:
3783:
3779:
3777:
3776:
3775:Z. brevicauda
3772:
3771:
3769:
3767:
3766:
3761:
3755:
3754:
3753:T. talamancae
3750:
3748:
3747:
3743:
3742:
3740:
3738:
3737:
3732:
3726:
3725:
3721:
3720:
3718:
3716:
3715:
3710:
3704:
3703:
3699:
3697:
3696:
3692:
3691:
3689:
3687:
3686:
3685:Sigmodontomys
3681:
3675:
3674:
3670:
3668:
3667:
3663:
3662:
3660:
3658:
3657:
3652:
3646:
3645:
3644:R. primigenus
3640:
3639:
3637:
3635:
3634:
3628:
3622:
3621:
3617:
3616:
3614:
3612:
3611:
3610:Pseudoryzomys
3606:
3600:
3599:
3594:
3593:
3591:
3589:
3588:
3582:
3576:
3575:
3574:O. peninsulae
3571:
3569:
3568:
3567:O. palustris
3564:
3562:
3561:
3556:
3554:
3553:
3549:
3547:
3546:
3545:O. dimidiatus
3542:
3540:
3539:
3535:
3533:
3532:
3531:O. antillarum
3527:
3525:
3524:
3523:O. albiventer
3520:
3519:
3517:
3515:
3514:
3509:
3503:
3502:
3498:
3497:
3495:
3493:
3492:
3487:
3481:
3480:
3475:
3473:
3472:
3468:
3466:
3465:
3464:O. stramineus
3461:
3459:
3458:
3454:
3452:
3451:
3447:
3445:
3444:
3440:
3438:
3437:
3433:
3431:
3430:
3426:
3424:
3423:
3419:
3417:
3416:
3412:
3410:
3409:
3408:O. fulvescens
3405:
3403:
3402:
3398:
3396:
3395:
3394:O. flavescens
3391:
3389:
3388:
3387:O. destructor
3384:
3382:
3381:
3380:O. chacoensis
3377:
3375:
3374:
3370:
3368:
3367:
3363:
3361:
3360:
3356:
3355:
3353:
3351:
3350:
3345:
3339:
3338:
3337:O. trinitatis
3334:
3332:
3331:
3327:
3325:
3324:
3320:
3318:
3317:
3313:
3311:
3310:
3306:
3304:
3303:
3299:
3297:
3296:
3292:
3290:
3289:
3285:
3283:
3282:
3278:
3276:
3275:
3271:
3269:
3268:
3264:
3262:
3261:
3257:
3255:
3254:
3250:
3248:
3247:
3246:O. catherinae
3243:
3241:
3240:
3236:
3234:
3233:
3232:O. auyantepui
3229:
3228:
3226:
3224:
3223:
3218:
3212:
3211:
3206:
3205:
3203:
3201:
3200:
3194:
3188:
3187:
3183:
3181:
3180:
3176:
3174:
3173:
3169:
3167:
3166:
3165:N. indefessus
3161:
3159:
3158:
3153:
3152:
3150:
3148:
3147:
3142:
3136:
3135:
3131:
3129:
3128:
3127:N. pectoralis
3124:
3122:
3121:
3117:
3115:
3114:
3110:
3108:
3107:
3106:N. meridensis
3103:
3101:
3100:
3096:
3094:
3093:
3089:
3087:
3086:
3082:
3080:
3079:
3075:
3073:
3072:
3068:
3066:
3065:
3061:
3059:
3058:
3057:N. auriventer
3054:
3052:
3051:
3047:
3046:
3044:
3042:
3041:
3036:
3030:
3029:
3025:
3023:
3022:
3018:
3016:
3015:
3011:
3009:
3008:
3004:
3002:
3001:
2997:
2996:
2994:
2992:
2991:
2986:
2980:
2979:
2975:
2973:
2972:
2968:
2966:
2965:
2961:
2959:
2958:
2954:
2952:
2951:
2947:
2945:
2944:
2940:
2938:
2937:
2933:
2931:
2930:
2926:
2925:
2923:
2921:
2920:
2915:
2909:
2908:
2904:
2903:
2901:
2899:
2898:
2893:
2887:
2886:
2882:
2880:
2879:
2878:M. altissimus
2875:
2874:
2872:
2870:
2869:
2864:
2858:
2857:
2853:
2852:
2850:
2848:
2847:
2842:
2836:
2835:
2831:
2829:
2828:
2827:M. robustulus
2824:
2822:
2821:
2817:
2816:
2814:
2812:
2811:
2806:
2800:
2799:
2794:
2792:
2791:
2786:
2784:
2783:
2778:
2776:
2775:
2774:M. curazensis
2770:
2768:
2767:
2762:
2761:
2759:
2757:
2756:
2750:
2744:
2743:
2739:
2738:
2736:
2734:
2733:
2728:
2722:
2721:
2717:
2715:
2714:
2710:
2708:
2707:
2706:H. perenensis
2703:
2701:
2700:
2696:
2694:
2693:
2689:
2687:
2686:
2682:
2680:
2679:
2675:
2674:
2672:
2670:
2669:
2664:
2658:
2657:
2653:
2651:
2650:
2646:
2644:
2643:
2639:
2638:
2636:
2634:
2633:
2628:
2622:
2621:
2620:H. saturatior
2617:
2615:
2614:
2610:
2608:
2607:
2603:
2601:
2600:
2596:
2594:
2593:
2589:
2587:
2586:
2582:
2580:
2579:
2575:
2573:
2572:
2568:
2567:
2565:
2563:
2562:
2557:
2551:
2550:
2546:
2544:
2543:
2539:
2537:
2536:
2532:
2530:
2529:
2525:
2523:
2522:
2518:
2516:
2515:
2511:
2510:
2508:
2506:
2505:
2500:
2494:
2493:
2489:
2488:
2486:
2484:
2483:
2478:
2472:
2471:
2467:
2466:
2464:
2462:
2461:
2456:
2450:
2449:
2445:
2443:
2442:
2438:
2436:
2435:
2431:
2429:
2428:
2424:
2422:
2421:
2417:
2415:
2414:
2410:
2408:
2407:
2403:
2402:
2400:
2398:
2397:
2392:
2386:
2385:
2380:
2379:
2377:
2375:
2374:
2368:
2362:
2361:
2357:
2356:
2354:
2352:
2351:
2350:Amphinectomys
2346:
2340:
2339:
2334:
2332:
2331:
2326:
2325:
2323:
2321:
2320:
2314:
2308:
2307:
2303:
2301:
2300:
2296:
2295:
2293:
2291:
2290:
2285:
2279:
2278:Sigmodontinae
2275:
2273:
2269:
2267:
2263:
2261:
2257:
2255:
2251:
2249:
2245:
2244:
2241:
2237:
2226:
2221:
2219:
2214:
2212:
2207:
2206:
2203:
2196:
2192:
2188:
2185:
2181:
2179:
2174:
2171:
2167:
2165:
2161:
2155:
2152:
2148:
2144:
2141:
2137:
2133:
2130:
2126:
2124:
2118:
2115:
2112:
2108:
2104:
2101:
2097:
2093:
2090:
2086:
2082:
2079:
2075:
2071:
2068:
2064:
2060:
2057:
2053:
2051:
2045:
2041:
2037:
2033:
2027:
2023:
2019:
2018:
2013:
2012:Wilson, D. E.
2009:
2005:
2004:Musser, G. G.
2001:
1998:
1995:
1991:
1988:
1984:
1982:
1976:
1971:
1966:
1962:
1958:
1957:
1952:
1950:
1943:
1940:
1936:
1934:
1928:
1925:
1924:0-7136-6021-X
1921:
1917:
1913:
1910:
1906:
1902:
1899:
1895:
1891:
1890:
1886:
1877:
1874:
1868:
1865:
1859:
1856:
1850:
1847:
1841:
1838:
1832:
1829:
1823:
1820:
1814:
1812:
1808:
1802:
1799:
1793:
1790:
1784:
1781:
1775:
1772:
1766:
1763:
1757:
1754:
1748:
1745:
1739:
1736:
1730:
1727:
1721:
1718:
1712:
1709:
1703:
1700:
1694:
1691:
1685:
1682:
1676:
1673:
1667:
1664:
1658:
1655:
1649:
1647:
1643:
1637:
1634:
1628:
1625:
1619:
1616:
1610:
1607:
1601:
1598:
1592:
1590:
1586:
1580:
1577:
1571:
1568:
1562:
1559:
1553:
1550:
1544:
1541:
1535:
1533:
1531:
1527:
1521:
1518:
1512:
1509:
1503:
1500:
1497:Weksler, 2006
1494:
1491:
1485:
1482:
1476:
1473:
1467:
1464:
1458:
1455:
1449:
1446:
1440:
1438:
1434:
1428:
1425:
1419:
1416:
1410:
1407:
1401:
1399:
1395:
1389:
1387:
1383:
1377:
1374:
1368:
1365:
1359:
1356:
1350:
1348:
1344:
1338:
1336:
1332:
1326:
1324:
1320:
1314:
1304:
1301:
1294:
1292:
1290:
1286:
1278:
1276:
1274:
1270:
1269:
1264:
1263:
1262:Akodon azarae
1258:
1257:
1252:
1251:
1246:
1245:
1240:
1236:
1235:
1230:
1229:
1225:
1221:
1217:
1213:
1209:
1204:
1200:
1196:
1189:
1187:
1185:
1181:
1177:
1176:
1171:
1166:
1162:
1158:
1154:
1153:Bajo San José
1150:
1146:
1141:
1139:
1135:
1131:
1127:
1123:
1122:ground sloths
1119:
1115:
1111:
1110:type locality
1107:
1106:
1101:
1100:
1095:
1094:
1089:
1088:
1083:
1079:
1075:
1071:
1067:
1065:
1061:
1057:
1050:
1048:
1046:
1042:
1038:
1034:
1030:
1026:
1022:
1018:
1013:
1011:
1010:Pseudoryzomys
1007:
1003:
999:
995:
994:Pseudoryzomys
991:
987:
983:
979:
978:Pseudoryzomys
975:
971:
967:
966:Pseudoryzomys
963:
959:
955:
951:
946:
944:
940:
936:
932:
928:
924:
920:
916:
912:
908:
903:
899:
895:
891:
887:
883:
879:
875:
870:
867:
863:
859:
855:
851:
847:
846:tufts of hair
843:
839:
838:
833:
832:
827:
826:
821:
817:
816:
811:
810:
805:
798:
796:
794:
790:
786:
785:
780:
776:
772:
771:morphological
768:
764:
763:Pseudoryzomys
760:
756:
752:
751:Pseudoryzomys
748:
744:
740:
736:
732:
728:
727:
722:
721:
716:
715:Pseudoryzomys
712:
708:
703:
701:
700:
695:
694:Pseudoryzomys
691:
687:
683:
679:
675:
671:
667:
663:
659:
655:
651:
647:
642:
640:
636:
632:
628:
624:
620:
616:
611:
609:
605:
601:
597:
593:
589:
588:
583:
582:
577:
576:
571:
567:
563:
559:
555:
551:
547:
543:
539:
534:
532:
528:
527:
522:
521:
516:
515:
510:
509:
504:
500:
499:
494:
490:
486:
482:
478:
474:
467:
465:
463:
462:
461:Pseudoryzomys
457:
456:
451:
450:
445:
444:
439:
435:
434:Sigmodontinae
431:
427:
426:
421:
417:
413:
412:
407:
404:Its external
402:
400:
396:
392:
388:
384:
380:
376:
371:
369:
368:South America
365:
361:
357:
353:
349:
348:
333:
330:
327:Massoia, 1980
324:
321:
315:
312:
308:
304:
301:
295:
292:
287:
284:
283:
281:
278:
274:
270:
265:
261:
256:
251:
245:
243:
237:
234:
233:Binomial name
230:
226:
225:
220:
217:
216:
211:
205:
204:
200:
197:
196:
193:
192:Sigmodontinae
190:
187:
186:
183:
180:
177:
176:
173:
170:
167:
166:
163:
160:
157:
156:
153:
150:
147:
146:
143:
140:
137:
136:
133:
130:
127:
126:
121:
116:
112:
106:
101:
100:Least Concern
90:
86:
81:
77:
73:
69:
65:
61:
57:
52:
47:
43:
37:
33:
30:
19:
4068:
4026:
3830:
3801:
3780:
3773:
3765:Zygodontomys
3763:
3751:
3746:T. bolivaris
3744:
3734:
3722:
3712:
3702:S. aphrastus
3700:
3693:
3683:
3671:
3664:
3654:
3642:
3631:
3618:
3608:
3596:
3585:
3572:
3566:
3558:
3550:
3543:
3536:
3529:
3521:
3511:
3501:O. balneator
3499:
3489:
3477:
3469:
3462:
3457:O. rupestris
3455:
3448:
3441:
3434:
3427:
3420:
3415:O. griseolus
3413:
3406:
3399:
3392:
3385:
3378:
3371:
3364:
3357:
3349:Oligoryzomys
3347:
3335:
3328:
3321:
3316:O. speciosus
3314:
3307:
3300:
3293:
3286:
3279:
3272:
3267:O. flavicans
3265:
3258:
3251:
3244:
3237:
3230:
3220:
3210:N. vespuccii
3208:
3197:
3184:
3177:
3170:
3163:
3155:
3144:
3134:N. pirrensis
3132:
3125:
3118:
3111:
3104:
3097:
3090:
3083:
3076:
3069:
3064:N. caracolus
3062:
3055:
3048:
3038:
3028:N. squamipes
3026:
3019:
3012:
3005:
2998:
2988:
2976:
2969:
2962:
2955:
2948:
2941:
2934:
2927:
2917:
2905:
2895:
2883:
2876:
2868:Microryzomys
2866:
2854:
2844:
2832:
2825:
2818:
2808:
2796:
2790:M. georginae
2788:
2780:
2772:
2764:
2753:
2741:
2740:
2731:
2730:
2718:
2711:
2704:
2697:
2690:
2683:
2676:
2666:
2654:
2649:H. chacarius
2647:
2640:
2630:
2618:
2613:H. rostratus
2611:
2604:
2599:H. melanotis
2597:
2590:
2583:
2576:
2569:
2559:
2547:
2540:
2533:
2526:
2519:
2512:
2504:Euryoryzomys
2502:
2490:
2482:Eremoryzomys
2480:
2468:
2458:
2446:
2441:C. subflavus
2439:
2432:
2425:
2418:
2413:C. langguthi
2411:
2404:
2394:
2382:
2373:Carletonomys
2371:
2358:
2348:
2336:
2328:
2319:Agathaeromys
2317:
2304:
2297:
2287:
2194:
2186:206:414–432.
2183:
2177:
2169:
2163:
2159:
2153:114:129–142.
2150:
2139:
2135:
2128:
2122:
2110:
2106:
2099:
2088:
2077:
2066:
2055:
2049:
2016:
1996:
1986:
1980:
1960:
1954:
1948:
1938:
1932:
1915:
1908:
1897:
1876:
1867:
1858:
1849:
1840:
1831:
1822:
1801:
1792:
1783:
1774:
1765:
1756:
1747:
1738:
1729:
1720:
1711:
1702:
1693:
1684:
1675:
1666:
1657:
1636:
1627:
1618:
1609:
1600:
1579:
1570:
1561:
1552:
1543:
1520:
1511:
1502:
1493:
1484:
1475:
1466:
1457:
1448:
1427:
1418:
1409:
1376:
1367:
1358:
1303:
1282:
1272:
1266:
1260:
1254:
1248:
1242:
1238:
1232:
1226:
1219:
1207:
1198:
1194:
1193:
1183:
1179:
1173:
1169:
1164:
1160:
1144:
1142:
1130:gomphotheres
1126:litopternans
1113:
1103:
1099:Macrauchenia
1097:
1091:
1085:
1081:
1068:
1064:Minas Gerais
1055:
1054:
1045:X chromosome
1041:Y chromosome
1014:
1009:
1005:
993:
989:
985:
977:
973:
969:
965:
961:
957:
947:
934:
922:
889:
873:
871:
866:gall bladder
835:
834:. Unlike in
829:
823:
813:
807:
803:
802:
792:
788:
782:
778:
774:
773:data placed
766:
762:
758:
750:
746:
742:
738:
734:
730:
724:
718:
714:
710:
706:
704:
699:Zygodontomys
697:
693:
689:
685:
681:
673:
669:
645:
643:
638:
634:
630:
626:
622:
618:
614:
612:
599:
594:, which are
585:
579:
573:
569:
565:
561:
557:
545:
541:
535:
530:
524:
518:
512:
506:
502:
496:
477:Herluf Winge
472:
471:
459:
455:Carletonomys
453:
447:
441:
424:
423:
409:
403:
394:
387:Minas Gerais
372:
355:
351:
346:
345:
344:
331:
322:
313:
302:
293:
285:
268:
241:
239:
223:
222:
202:
201:
71:
35:
29:
3909:iNaturalist
3855:Wikispecies
3782:Z. brunneus
3666:S. melanops
3491:Oreoryzomys
3450:O. nigripes
3436:O. microtis
3366:O. arenalis
3323:O. superans
3288:O. phaeotis
3281:O. paricola
3260:O. concolor
3146:Nesoryzomys
3120:N. nimbosus
3014:N. palmipes
3000:N. apicalis
2978:N. tenuipes
2971:N. spinosus
2907:M. hammondi
2766:M. audreyae
2720:H. yunganus
2685:H. laticeps
2656:H. sciureus
2606:H. rhabdops
2592:H. intectus
2585:H. fuscatus
2578:H. chapmani
2561:Handleyomys
2549:E. russatus
2514:E. emmonsae
2460:Drymoreomys
2427:C. marinhus
2330:A. donovani
2276:Subfamily:
2235:(rice rats)
2233:Oryzomyini
2142:23:179–181.
2100:Ameghiniana
1989:37:619–673.
1935:from Brazil
1216:herbivorous
1143:Remains of
1134:glyptodonts
1070:Pleistocene
1037:metacentric
1033:telocentric
1029:acrocentric
1021:chromosomes
998:anteroconid
822:, such as "
799:Description
556:, reviewed
544:(currently
397:is a large
377:and nearby
360:semiaquatic
307:Hershkovitz
289:Winge, 1888
208:Voss &
188:Subfamily:
44:to Recent (
42:Pleistocene
4106:Oryzomyini
4075:Categories
4042:Q109647311
3724:S. angouya
3714:Sooretamys
3620:P. simplex
3598:P. nivalis
3587:Pennatomys
3560:O. nelsoni
3552:O. gorgasi
3471:O. vegetus
3443:O. moojeni
3401:O. fornesi
3373:O. brendae
3359:O. andinus
3309:O. rutilus
3302:O. roberti
3274:O. mamorae
3253:O. cleberi
3239:O. bicolor
3199:Noronhomys
3186:N. swarthi
3157:N. darwini
3092:N. levipes
3040:Nephelomys
3007:N. grandis
2957:N. paracou
2950:N. musseri
2943:N. minutus
2936:N. guianae
2929:N. dubosti
2885:M. minutus
2834:M. zunigae
2742:L. molitor
2699:H. oniscus
2678:H. acritus
2632:Holochilus
2571:H. alfaroi
2542:E. nitidus
2528:E. legatus
2406:C. goytaca
2396:Cerradomys
2360:A. savamis
2289:Aegialomys
2272:Cricetidae
2197:296:1–149.
2172:3085:1–39.
2162:Winge and
2123:Holochilus
1981:Holochilus
1911:3256:1–59.
1315:References
1220:Holochilus
1208:Holochilus
1199:Holochilus
1155:, date to
1120:, such as
1114:L. molitor
1082:L. molitor
1006:Holochilus
990:Holochilus
982:anteroloph
974:Holochilus
962:Holochilus
958:Holochilus
935:Holochilus
890:Holochilus
882:cheek bone
874:Holochilus
864:, and the
809:Holochilus
789:Holochilus
775:Holochilus
759:Holochilus
747:Holochilus
735:Holochilus
707:Holochilus
690:Holochilus
670:Holochilus
666:mesolophid
650:Oryzomyini
631:Holochilus
623:Holochilus
581:Reithrodon
570:Holochilus
562:Holochilus
558:Holochilus
508:Hesperomys
449:Noronhomys
443:Holochilus
438:oryzomyine
430:Cricetidae
406:morphology
182:Cricetidae
72:L. molitor
3695:S. alfari
3538:O. couesi
3479:O. victus
3113:N. moerex
3085:N. keaysi
3078:N. devius
3071:N. childi
3021:N. rattus
2964:N. pictus
2810:Melanomys
2798:M. luciae
2755:Megalomys
2668:Hylaeamys
2492:E. polius
2434:C. scotti
2384:C. cailoi
2246:Kingdom:
1941:61:13–20.
1295:Footnotes
1118:megafauna
1093:Glyptodon
1017:karyotype
950:hypsodont
911:squamosal
600:H. magnus
596:lophodont
416:taxonomic
391:Argentina
218:Species:
138:Kingdom:
132:Eukaryota
64:Lectotype
4036:Wikidata
3979:13000692
3927:10542062
3846:Q1321166
3840:Wikidata
3656:Scolomys
3633:Reigomys
3513:Oryzomys
2990:Nectomys
2919:Neacomys
2897:Mindomys
2732:Lundomys
2713:H. tatei
2521:E. lamia
2448:C. vivoi
2270:Family:
2266:Rodentia
2260:Mammalia
2254:Chordata
2252:Phylum:
2248:Animalia
2136:Lundomys
2040:62265494
1939:Genetica
1212:stomachs
1184:Lundomys
1180:Lundomys
1170:Lundomys
1165:Lundomys
1161:Lundomys
1145:Lundomys
1138:cricetid
1074:Lujanian
986:Lundomys
970:Lundomys
923:Lundomys
902:incisors
815:Nectomys
793:Lundomys
779:Lundomys
767:Lundomys
743:Lundomys
739:Lundomys
731:Lundomys
711:Lundomys
686:Lundomys
682:Lundomys
674:Lundomys
662:mesoloph
646:Lundomys
635:Lundomys
587:Neotomys
575:Sigmodon
526:Oryzomys
493:mandible
468:Taxonomy
425:Lundomys
277:Synonyms
210:Carleton
203:Lundomys
178:Family:
172:Rodentia
162:Mammalia
152:Chordata
148:Phylum:
142:Animalia
128:Domain:
105:IUCN 3.1
76:mandible
66:partial
46:Lujanian
36:Lundomys
4056:5786882
4015:6363585
3966:1002561
3901:4265127
3222:Oecomys
2264:Order:
2258:Class:
1218:, like
1105:Toxodon
954:derived
927:angular
858:plantar
765:, with
531:Calomys
520:Calomys
489:maxilla
375:Uruguay
364:species
358:, is a
354:or the
252:, 1887)
198:Genus:
168:Order:
158:Class:
103: (
68:cranium
4012:uBio:
4005:380006
3992:218791
3940:970677
3295:O. rex
2107:Kunsia
2038:
2028:
1922:
1271:, and
1132:, and
1108:. The
1102:, and
937:. The
921:, but
892:. The
876:, the
828:" and
749:, and
713:, and
696:, and
658:palate
592:molars
584:, and
458:, and
399:rodent
383:Brazil
309:, 1955
212:, 1993
3953:10219
3922:IRMNG
3914:74529
3888:72PZS
2010:. In
1224:mites
1002:roots
886:jugal
862:teats
652:in a
420:genus
250:Winge
4051:GBIF
3987:NCBI
3948:IUCN
3935:ITIS
3896:GBIF
2067:Gaea
2036:OCLC
2026:ISBN
1961:2016
1920:ISBN
1231:and
1203:nest
1015:The
1008:and
992:and
976:and
964:and
929:and
850:pads
812:and
791:and
777:and
761:and
755:IRBP
723:and
672:and
362:rat
3974:MSW
3961:MDD
3883:CoL
3870:ADW
1965:doi
606:of
529:or
70:of
4077::
4053::
4038::
4002::
3989::
3976::
3963::
3950::
3937::
3924::
3911::
3898::
3885::
3872::
3857::
3842::
2193:.
2182:.
2168:.
2098:.
2087:.
2065:.
2034:.
1985:.
1959:.
1953:.
1907:.
1896:.
1810:^
1645:^
1588:^
1529:^
1436:^
1397:^
1385:^
1346:^
1334:^
1322:^
1275:.
1265:,
1259:,
1253:,
1247:,
1128:,
1124:,
1096:,
1025:FN
795:.
745:,
709:,
692:,
641:.
625:,
578:,
552:,
533:.
464:.
452:,
446:,
422:,
381:,
370:.
334::
325::
316::
296::
3806:"
3802:"
3800:†
3641:†
3630:†
3595:†
3584:†
3557:†
3528:†
3476:†
3207:†
3196:†
3162:†
3154:†
2795:†
2787:†
2779:†
2771:†
2763:†
2752:†
2381:†
2370:†
2335:†
2327:†
2316:†
2224:e
2217:t
2210:v
2042:.
1973:.
1967::
1951:"
1947:"
1926:.
248:(
107:)
48:)
20:)
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