734:
683:
717:
751:
700:
31:
53:
550:
characterized by predominantly dorso-ventrally compressed bodies, head and pronotal discs bordered by ridges or grooves, and inverted male genitalia. Size range of adults is 1–5 mm (0.04–0.2 in) in length. Currently, it contains 40 genera and about 450 species, and is represented on all
671:, a discovery has been made regarding the larvae of Laemophloeidae. Studies show that when the larvae are in the open, the larvae will straighten themselves out and jump curling up into a ball in the process. It has also been discovered that the larvae can bounce when they are curled up.
559:
Historically, Laemophloeidae was treated as a subfamily of
Cucujidae, but starting in the middle of the 20th century, most of what had been treated as subfamilies of the Cucujidae were considered to be families.
658:
are atypical in that the adults are rostrate to varying degrees. Photographs of most world genera are available at the
Florida State Collection of Arthropod website, with most North American species pictured.
568:
Most laemophloeids, adults and larvae, are found under bark of dead trees, where they apparently are primarily fungivores, although some genera with adults having subcylindrical bodies (e.g.,
814:
978:
Thomas MC (March 1984). "A new
Neotropical genus and species of rostrate Laemophloeinae (Coleoptera: Cucujidae) with discussion of the systematic position of the subfamily".
1033:
3219:
3258:
3040:
3193:
901:
Halstead DG (June 1993). "Keys for the identification of beetles associated with stored products—II. Laemophloeidae, Passandridae and
Silvanidae".
3232:
1006:
822:
1153:
733:
928:
Lefkovitch LP (July 1958). "Unusual antennal characters in some
Laemophloeinae (Coleoptera: Cucujidae) and their taxonomic importance".
716:
682:
801:
Handbook of
Zoology. Coleoptera, Beetles. Volume 2: Morphology and Systematics (Elateroidea, Bostrichiformia, Cucujiformia partim)
750:
890:. Vol. 15. Florida Department of Agriculture and Consumer Services, Division of Plant Industry. pp. i-viii and 1–93.
699:
1037:
3237:
956:
3245:
52:
3340:
783:
Thomas MC (2002). "Family 83. Laemophloeidae
Ganglbauer 1899.". In Arnett Jr R, Thomas MC, Skelley PE, Frank JH (eds.).
3115:
3102:
3263:
3172:
859:
Lawrence JF (December 1977). "Coleoptera associated with an
Hypoxylon species (Ascomycetes: Xylariaceae) on oak".
3185:
799:
Thomas MC, Leschen RA (2010). "Laemophloeidae, Ganglbauer, 1899.". In
Leschen RA, Beutel RG, Lawrence JF (eds.).
1146:
757:
596:
3335:
740:
706:
39:
3055:
3120:
3294:
723:
608:
602:
3312:
3273:
3154:
2292:
3093:
2511:
1139:
1010:
886:
Thomas MC (1993). "The flat bark beetles of
Florida (Laemophloeidae, Passandridae, Silvanidae).".
987:
868:
47:
594:, contain some species that are pests of stored grain products. The most important of these are
30:
3299:
3281:
3141:
2780:
2197:
1117:
149:
3224:
3286:
2877:
2656:
2651:
2574:
2332:
1348:
1107:
1097:
937:
910:
418:
409:
2566:
2519:
2364:
2348:
2024:
1784:
1753:
1668:
1586:
1477:
1445:
1060:
634:), with the scape expanded into hook-like or blade-like structures. Several other genera (
1086:"A novel power-amplified jumping behavior in larval beetles (Coleoptera: Laemophloeidae)"
3180:
3167:
2969:
2920:
2775:
2582:
2250:
2234:
2173:
2068:
2039:
1931:
1915:
1831:
1776:
1681:
1578:
1427:
1389:
1369:
1265:
1112:
1085:
941:
508:
373:
256:
166:
3107:
3329:
2951:
2755:
2746:
2487:
2458:
2450:
2442:
2416:
2391:
2287:
2274:
2213:
2131:
2034:
2029:
1910:
1844:
1815:
1799:
1763:
1733:
1686:
1640:
1607:
1523:
1515:
1486:
1324:
914:
689:
648:
585:
472:
463:
391:
337:
292:
193:
2928:
2852:
2598:
2590:
2545:
2529:
2479:
2400:
2356:
2308:
2226:
2139:
2008:
1995:
1975:
1936:
1836:
1823:
1794:
1728:
1716:
1612:
1569:
1552:
1544:
1494:
1332:
1302:
1233:
1222:
1206:
654:
642:
630:
590:
581:
570:
499:
436:
346:
301:
274:
238:
220:
211:
184:
124:
3146:
1102:
3250:
3206:
3087:
3014:
2985:
2959:
2946:
2886:
2844:
2828:
2820:
2812:
2796:
2788:
2724:
2682:
2674:
2643:
2558:
2537:
2503:
2495:
2408:
2340:
2324:
2316:
2279:
2266:
2242:
2160:
2152:
2102:
2094:
2052:
2016:
1987:
1959:
1954:
1870:
1865:
1857:
1807:
1743:
1738:
1652:
1617:
1602:
1594:
1510:
1502:
1319:
526:
481:
454:
427:
355:
328:
319:
310:
265:
3078:
2941:
2902:
2894:
2836:
2699:
2661:
2614:
2606:
2426:
2374:
2258:
2189:
2060:
2044:
2000:
1941:
1905:
1885:
1849:
1758:
1748:
1622:
1356:
1340:
636:
547:
517:
490:
445:
400:
382:
364:
202:
134:
1131:
588:: Scolytinae), upon which they may feed. A few genera, but most particularly
3006:
2998:
2990:
2977:
2933:
2732:
2707:
2635:
2463:
2434:
2118:
1949:
1923:
1875:
1789:
1691:
1660:
1632:
1465:
1440:
1435:
1416:
1381:
1270:
1253:
1245:
1194:
1182:
930:
Proceedings of the Royal Entomological Society of London. Series B, Taxonomy
283:
229:
114:
84:
64:
1121:
785:
American Beetles. Volume 2: Polyphaga: Scarabaeoidea through Curculionoidea
551:
continents except Antarctica; species richness is greatest in the tropics.
620:
Several genera exhibit unusual modifications to male antennae (especially
3128:
3072:
2910:
2860:
2804:
2763:
2712:
2369:
2300:
2181:
1893:
1699:
1673:
1531:
1364:
1311:
1290:
1200:
576:
247:
3133:
2627:
2619:
2471:
2205:
2126:
2110:
991:
872:
3211:
2218:
2078:
1711:
1188:
1176:
1163:
543:
175:
104:
94:
74:
3198:
3049:
1084:
Bertone MA, Gibson JC, Seago AE, Yoshida T, Smith AA (2022-01-19).
957:"A review of the genus Dysmerus Casey (Coleoptera: Laemophloeidae)"
3307:
157:
3053:
1462:
1413:
1287:
1219:
1135:
3159:
1534:(furniture beetles, death watch beetles, spider beetles)
1036:. Florida State Collection of Arthropods. Archived from
1009:. Florida State Collection of Arthropods. Archived from
846:
The natural classification of the families of Coleoptera
936:(5–6). Oxford, UK: Blackwell Publishing Ltd.: 93–100.
3062:
2968:
2919:
2885:
2876:
2754:
2745:
2698:
2673:
2557:
2528:
2425:
2399:
2390:
2172:
2151:
2077:
1884:
1775:
1631:
1577:
1568:
1543:
1485:
1476:
1301:
2411:(jewel beetles, or metallic wood-boring beetles)
888:Arthropods of Florida and Neighboring Land Areas
2134:(true weevils, bark beetles, ambrosia beetles)
803:. Berlin: Walter de Gruyter. pp. 376–380.
1147:
8:
2377:(ironclad beetles, cylindrical bark beetles)
1007:"Photographic Atlas of Laemophloeid Genera"
3050:
2882:
2751:
2396:
1574:
1482:
1473:
1459:
1410:
1298:
1284:
1216:
1154:
1140:
1132:
821:. Texas A&M University. Archived from
787:. Boca Raton: CRC Press. pp. 331–334.
29:
20:
3041:List of subgroups of the order Coleoptera
2121:(straight snout weevils, New York weevil)
1111:
1101:
2548:(cicada beetle, cicada parasite beetles)
848:. London: Nathaniel Lloyd. p. 187.
775:
678:
2638:(false fireflies, long-lipped beetles)
667:According to an article published by
7:
3313:3F66C508-1514-9F3A-7CB6-C65AD3FAF808
3274:a88b35d2-f899-44ca-9dc4-a7b8391b4585
3186:6e88d022-160b-4451-a1e7-e34d54505822
2664:(false metallic wood-boring beetles)
903:Journal of Stored Products Research
942:10.1111/j.1365-3113.1958.tb00419.x
546:," is a family in the superfamily
14:
270:Sengupta & Mukhopadhyay, 1978
2275:Perimylopidae, or Promecheilidae
1860:(minute brown scavenger beetles)
749:
732:
715:
698:
681:
51:
2490:(variegated mud-loving beetles)
1:
2319:(narrow-waisted bark beetles)
2011:(parasitic flat bark beetles)
1962:(short-winged flower beetles)
1818:(lady beetles, or God's cows)
1555:(tooth-necked fungus beetles)
819:Joel Hallan’s Biology Catalog
815:"Laemophloeidae Species List"
2831:(sand-loving scarab beetles)
2221:(minute tree-fungus beetles)
2047:(silvanid flat bark beetles)
1103:10.1371/journal.pone.0256509
915:10.1016/0022-474X(93)90030-8
580:) occur in the galleries of
2980:(primitive carrion beetles)
2936:(minute mud-loving beetles)
2799:(bumble bee scarab beetles)
2569:(soft-bodied plant beetles)
2540:(soft bodied plant beetles)
1343:(predaceous diving beetles)
1034:"Laemophloeidae of Florida"
980:The Coleopterists' Bulletin
861:The Coleopterists' Bulletin
3357:
2815:(scavenger scarab beetles)
2807:(enigmatic scarab beetles)
1980:Lamingtonium binnaberrense
1694:(soft-wing flower beetles)
405:J.LeConte & Horn, 1880
279:Laporte de Castelnau, 1840
3038:
2954:(water scavenger beetles)
2687:Rhinorhipus tamborinensis
2466:(long-toed water beetles)
2359:(polypore fungus beetles)
2261:(palm and flower beetles)
2245:(tumbling flower beetles)
2192:(ant-like flower beetles)
1970:(lined flat bark beetles)
1944:(pleasing fungus beetles)
1839:(handsome fungus beetles)
1472:
1458:
1423:
1409:
1384:(burrowing water beetles)
1297:
1283:
1229:
1215:
1171:
163:
156:
48:Scientific classification
46:
37:
28:
23:
3001:(feather-winged beetles)
2335:(false longhorn beetles)
2282:(jugular-horned beetles)
2229:(false darkling beetles)
2200:(cryptic fungus beetles)
2019:(shining flower beetles)
1359:(crawling water beetles)
1258:Sikhotealinia zhiltzovae
1061:"These Beetles Go Boing"
758:Lepidophloeus exquisitus
597:Cryptolestes ferrugineus
468:Yablokov-Khnzoryan, 1977
2482:(forest stream beetles)
2343:(synchroa bark beetles)
2303:(dead log bark beetles)
2295:(fire-coloured beetles)
2184:(ant-like leaf beetles)
1918:(silken fungus beetles)
1852:(well polished beetles)
1826:(minute fungus beetles)
1645:Acanthocnemus nigricans
1059:Bergquist C, Flatow I.
2993:(round fungus beetles)
2327:(false flower beetles)
2311:(wedge-shaped beetles)
2269:(false blister beetle)
2253:(hairy fungus beetles)
2208:(conifer bark beetles)
2105:(leaf-rolling weevils)
1766:(bark-gnawing beetles)
1526:(wounded-tree beetles)
1392:(false ground beetles)
1314:(trout-stream beetles)
955:Thomas MC (May 2009).
741:Metaxyphloeus germaini
3295:Paleobiology Database
2905:(false clown beetles)
2727:(plate-thigh beetles)
2609:(false click beetles)
2514:(water-penny beetles)
2163:(ship-timber beetles)
2142:(pine flower weevils)
1990:(root-eating beetles)
1810:(minute bark beetles)
1721:Phloiophilus edwardsi
1248:(reticulated beetles)
707:Phloeolaemus hoplites
609:Cryptolestes turcicus
603:Cryptolestes pusillus
564:Habitat and behaviour
40:Laemophloeus germaini
3181:Fauna Europaea (new)
2783:(false stag beetles)
2768:Belohina inexpectata
2630:(net-winged beetles)
2593:(rare click beetles)
2506:(travertine beetles)
2498:(minute mud beetles)
2063:(dry-fungus beetles)
1898:Agapytho foveicollis
1655:(false skin beetles)
1518:(Jacobson's beetles)
1238:Crowsoniella relicta
3341:Cucujoidea families
2781:Diphyllostomatidae
2113:(primitive weevils)
1926:(flat bark beetles)
1676:(checkered beetles)
1663:(fruitworm beetles)
1351:(whirligig beetles)
844:Crowson RA (1955).
542:, "lined flat bark
2646:(glowworm beetles)
2351:(darkling beetles)
2198:Archeocrypticidae
2055:(palmetto beetles)
1802:(dry bark beetles)
1589:(longhorn beetles)
1040:on 1 February 2014
1013:on 1 February 2014
825:on 23 October 2012
724:Dysmerus symphilus
3323:
3322:
3282:Open Tree of Life
3056:Taxon identifiers
3047:
3046:
3034:
3033:
3030:
3029:
3026:
3025:
3018:
3010:
3009:(carrion beetles)
3002:
2994:
2981:
2955:
2937:
2906:
2898:
2872:
2871:
2864:
2856:
2848:
2840:
2832:
2824:
2816:
2808:
2800:
2792:
2784:
2771:
2741:
2740:
2728:
2720:
2690:
2665:
2647:
2639:
2631:
2623:
2610:
2602:
2594:
2586:
2585:(soldier beetles)
2578:
2575:Brachypsectridae
2570:
2549:
2541:
2515:
2507:
2499:
2491:
2483:
2475:
2467:
2454:
2446:
2438:
2412:
2386:
2385:
2378:
2360:
2352:
2344:
2336:
2333:Stenotrachelidae
2328:
2320:
2312:
2304:
2296:
2283:
2270:
2262:
2254:
2246:
2238:
2237:(blister beetles)
2230:
2222:
2209:
2201:
2193:
2185:
2164:
2143:
2135:
2122:
2114:
2106:
2098:
2088:
2064:
2056:
2048:
2020:
2012:
2004:
1991:
1983:
1971:
1963:
1945:
1927:
1919:
1901:
1861:
1853:
1840:
1827:
1819:
1811:
1803:
1785:Akalyptoischiidae
1767:
1724:
1707:
1695:
1677:
1664:
1656:
1648:
1598:
1590:
1564:
1563:
1556:
1535:
1527:
1519:
1506:
1498:
1454:
1453:
1431:
1405:
1404:
1401:
1400:
1393:
1385:
1377:
1360:
1352:
1344:
1336:
1328:
1315:
1279:
1278:
1261:
1249:
1241:
606:(Schönherr), and
537:
536:
532:
523:
514:
505:
496:
487:
478:
469:
460:
451:
442:
433:
424:
415:
406:
397:
388:
379:
370:
361:
352:
343:
334:
325:
316:
307:
298:
289:
280:
271:
262:
253:
244:
235:
226:
217:
208:
199:
190:
181:
172:
152:
16:Family of beetles
3348:
3316:
3315:
3303:
3302:
3290:
3289:
3277:
3276:
3267:
3266:
3254:
3253:
3251:NHMSYS0020152353
3241:
3240:
3228:
3227:
3215:
3214:
3202:
3201:
3189:
3188:
3176:
3175:
3163:
3162:
3150:
3149:
3137:
3136:
3124:
3123:
3111:
3110:
3098:
3097:
3096:
3083:
3082:
3081:
3051:
3016:
3008:
3000:
2992:
2979:
2953:
2935:
2904:
2896:
2883:
2878:Staphyliniformia
2862:
2854:
2846:
2838:
2830:
2822:
2814:
2806:
2798:
2790:
2782:
2765:
2752:
2726:
2717:Declinia relicta
2714:
2684:
2663:
2657:Sinopyrophoridae
2652:Rhagophthalmidae
2645:
2637:
2629:
2621:
2608:
2600:
2592:
2584:
2576:
2568:
2567:Artematopodidae
2547:
2539:
2513:
2505:
2497:
2489:
2481:
2474:(riffle beetles)
2473:
2465:
2453:(turtle beetles)
2452:
2444:
2436:
2410:
2397:
2376:
2358:
2350:
2342:
2334:
2326:
2318:
2310:
2302:
2294:
2281:
2268:
2260:
2252:
2244:
2236:
2228:
2220:
2207:
2199:
2191:
2183:
2162:
2141:
2133:
2120:
2112:
2104:
2097:(fungus weevils)
2096:
2087:
2086:
2082:
2069:Tasmosalpingidae
2062:
2054:
2046:
2018:
2010:
2002:
1989:
1977:
1969:
1961:
1943:
1925:
1917:
1895:
1859:
1851:
1838:
1825:
1817:
1809:
1801:
1765:
1718:
1701:
1693:
1675:
1662:
1654:
1642:
1641:Acanthocnemidae
1596:
1588:
1575:
1554:
1533:
1525:
1517:
1504:
1496:
1483:
1474:
1460:
1429:
1411:
1391:
1383:
1371:
1358:
1350:
1342:
1334:
1327:(ground beetles)
1326:
1313:
1299:
1285:
1255:
1247:
1235:
1234:Crowsoniellidae
1217:
1156:
1149:
1142:
1133:
1126:
1125:
1115:
1105:
1081:
1075:
1074:
1072:
1071:
1056:
1050:
1049:
1047:
1045:
1029:
1023:
1022:
1020:
1018:
1002:
996:
995:
975:
969:
968:
952:
946:
945:
925:
919:
918:
898:
892:
891:
883:
877:
876:
856:
850:
849:
841:
835:
834:
832:
830:
811:
805:
804:
796:
790:
788:
780:
753:
736:
719:
702:
685:
555:Classicification
531:Lefkovitch, 1962
530:
522:Lefkovitch, 1962
521:
512:
503:
494:
485:
476:
467:
459:Lefkovitch, 1958
458:
449:
440:
431:
422:
419:Passandrophloeus
413:
410:Paraphloeolaemus
404:
395:
387:Lefkovitch, 1959
386:
377:
368:
359:
350:
341:
333:Lefkovitch, 1962
332:
324:Lefkovitch, 1962
323:
315:Lefkovitch, 1962
314:
305:
296:
287:
278:
269:
261:Lefkovitch, 1962
260:
251:
242:
233:
225:Ganglbauer, 1899
224:
215:
206:
198:Lefkovitch, 1961
197:
188:
180:Lefkovitch, 1962
179:
170:
150:Ganglbauer, 1899
148:
56:
55:
33:
21:
3356:
3355:
3351:
3350:
3349:
3347:
3346:
3345:
3326:
3325:
3324:
3319:
3311:
3306:
3298:
3293:
3285:
3280:
3272:
3270:
3262:
3257:
3249:
3244:
3236:
3231:
3223:
3218:
3210:
3205:
3197:
3192:
3184:
3179:
3171:
3166:
3158:
3153:
3145:
3140:
3132:
3127:
3119:
3114:
3106:
3101:
3092:
3091:
3086:
3077:
3076:
3071:
3058:
3048:
3043:
3022:
2964:
2915:
2897:(clown beetles)
2868:
2839:(betsy beetles)
2747:Scarabaeiformia
2737:
2694:
2669:
2601:(click beetles)
2577:(Texas beetles)
2553:
2524:
2520:Ptilodactylidae
2445:(cedar beetles)
2421:
2382:
2365:Trictenotomidae
2251:Mycetophagidae
2168:
2147:
2084:
2083:
2081:
2073:
2025:Phloeostichidae
1976:Lamingtoniidae
1968:Laemophloeidae
1916:Cryptophagidae
1880:
1771:
1754:Thanerocleridae
1717:Phloiophilidae
1704:Metaxina ornata
1669:Chaetosomatidae
1627:
1560:
1539:
1497:(auger beetles)
1478:Bostrichiformia
1468:
1450:
1446:Torridincolidae
1430:(skiff beetles)
1428:Hydroscaphidae
1419:
1397:
1390:Trachypachidae
1335:(tiger beetles)
1293:
1275:
1225:
1211:
1167:
1160:
1130:
1129:
1096:(1): e0256509.
1083:
1082:
1078:
1069:
1067:
1058:
1057:
1053:
1043:
1041:
1031:
1030:
1026:
1016:
1014:
1004:
1003:
999:
977:
976:
972:
954:
953:
949:
927:
926:
922:
900:
899:
895:
885:
884:
880:
858:
857:
853:
843:
842:
838:
828:
826:
813:
812:
808:
798:
797:
793:
782:
781:
777:
772:
766:
762:
754:
745:
737:
728:
720:
711:
703:
694:
686:
677:
665:
618:
616:Characteristics
566:
557:
524:
515:
506:
497:
488:
479:
470:
461:
452:
443:
434:
425:
416:
407:
398:
389:
380:
371:
369:J.LeConte, 1861
362:
360:Lefkovich, 1962
353:
344:
335:
326:
317:
308:
299:
290:
281:
272:
263:
254:
245:
236:
227:
218:
209:
207:Wollaston, 1862
200:
191:
182:
173:
147:
50:
24:Laemophloeidae
17:
12:
11:
5:
3354:
3352:
3344:
3343:
3338:
3336:Laemophloeidae
3328:
3327:
3321:
3320:
3318:
3317:
3304:
3291:
3278:
3268:
3255:
3242:
3229:
3216:
3203:
3190:
3177:
3168:Fauna Europaea
3164:
3151:
3138:
3125:
3112:
3108:Laemophloeidae
3099:
3094:Laemophloeidae
3084:
3068:
3066:
3064:Laemophloeidae
3060:
3059:
3054:
3045:
3044:
3039:
3036:
3035:
3032:
3031:
3028:
3027:
3024:
3023:
3021:
3020:
3017:(rove beetles)
3015:Staphylinidae
3012:
3004:
2996:
2988:
2983:
2974:
2972:
2970:Staphylinoidea
2966:
2965:
2963:
2962:
2957:
2952:Hydrophilidae
2949:
2944:
2939:
2931:
2925:
2923:
2921:Hydrophiloidea
2917:
2916:
2914:
2913:
2908:
2900:
2891:
2889:
2880:
2874:
2873:
2870:
2869:
2867:
2866:
2863:(hide beetles)
2858:
2850:
2847:(rain beetles)
2842:
2834:
2826:
2823:(stag beetles)
2818:
2810:
2802:
2794:
2786:
2778:
2776:Bolboceratidae
2773:
2760:
2758:
2749:
2743:
2742:
2739:
2738:
2736:
2735:
2730:
2722:
2710:
2704:
2702:
2696:
2695:
2693:
2692:
2683:Rhinorhipidae
2679:
2677:
2675:Rhinorhipoidea
2671:
2670:
2668:
2667:
2659:
2654:
2649:
2641:
2633:
2625:
2617:
2612:
2604:
2596:
2588:
2580:
2572:
2563:
2561:
2555:
2554:
2552:
2551:
2543:
2534:
2532:
2526:
2525:
2523:
2522:
2517:
2509:
2501:
2493:
2488:Heteroceridae
2485:
2477:
2469:
2461:
2456:
2451:Chelonariidae
2448:
2443:Callirhipidae
2440:
2437:(pill beetles)
2431:
2429:
2423:
2422:
2420:
2419:
2414:
2405:
2403:
2394:
2388:
2387:
2384:
2383:
2381:
2380:
2372:
2367:
2362:
2354:
2349:Tenebrionidae
2346:
2338:
2330:
2322:
2314:
2306:
2298:
2290:
2285:
2277:
2272:
2264:
2256:
2248:
2240:
2232:
2224:
2216:
2211:
2203:
2195:
2187:
2178:
2176:
2174:Tenebrionoidea
2170:
2169:
2167:
2166:
2157:
2155:
2149:
2148:
2146:
2145:
2137:
2132:Curculionidae
2129:
2124:
2116:
2108:
2100:
2091:
2089:
2079:Curculionoidea
2075:
2074:
2072:
2071:
2066:
2058:
2050:
2042:
2040:Protocucujidae
2037:
2032:
2027:
2022:
2014:
2006:
1998:
1993:
1985:
1973:
1965:
1957:
1952:
1947:
1939:
1934:
1932:Cybocephalidae
1929:
1921:
1913:
1908:
1903:
1890:
1888:
1882:
1881:
1879:
1878:
1873:
1868:
1863:
1855:
1847:
1842:
1834:
1832:Discolomatidae
1829:
1821:
1816:Coccinellidae
1813:
1805:
1800:Bothrideridae
1797:
1792:
1787:
1781:
1779:
1777:Coccinelloidea
1773:
1772:
1770:
1769:
1764:Trogossitidae
1761:
1756:
1751:
1746:
1741:
1736:
1731:
1726:
1714:
1709:
1697:
1689:
1684:
1682:Lophocateridae
1679:
1671:
1666:
1658:
1650:
1637:
1635:
1629:
1628:
1626:
1625:
1620:
1615:
1610:
1605:
1600:
1597:(leaf beetles)
1595:Chrysomelidae
1592:
1583:
1581:
1579:Chrysomeloidea
1572:
1566:
1565:
1562:
1561:
1559:
1558:
1549:
1547:
1541:
1540:
1538:
1537:
1529:
1524:Nosodendridae
1521:
1516:Jacobsoniidae
1513:
1508:
1505:(skin beetles)
1500:
1491:
1489:
1480:
1470:
1469:
1463:
1456:
1455:
1452:
1451:
1449:
1448:
1443:
1438:
1433:
1424:
1421:
1420:
1414:
1407:
1406:
1403:
1402:
1399:
1398:
1396:
1395:
1387:
1379:
1374:Meru phyllisae
1367:
1362:
1354:
1346:
1338:
1330:
1322:
1317:
1308:
1306:
1295:
1294:
1288:
1281:
1280:
1277:
1276:
1274:
1273:
1268:
1266:Micromalthidae
1263:
1251:
1243:
1230:
1227:
1226:
1220:
1213:
1212:
1210:
1209:
1203:
1197:
1191:
1185:
1179:
1172:
1169:
1168:
1161:
1159:
1158:
1151:
1144:
1136:
1128:
1127:
1076:
1065:Science Friday
1051:
1024:
997:
970:
947:
920:
893:
878:
867:(4): 309–312.
851:
836:
806:
791:
774:
773:
771:
768:
764:
763:
755:
748:
746:
738:
731:
729:
721:
714:
712:
704:
697:
695:
687:
680:
676:
673:
664:
663:Leaping larvae
661:
617:
614:
565:
562:
556:
553:
540:Laemophloeidae
535:
534:
509:Sinuatophloeus
374:Nipponophloeus
288:Erichson, 1845
167:Acompsophloeus
161:
160:
154:
153:
145:Laemophloeidae
142:
138:
137:
132:
128:
127:
122:
118:
117:
112:
108:
107:
102:
98:
97:
92:
88:
87:
82:
78:
77:
72:
68:
67:
62:
58:
57:
44:
43:
35:
34:
26:
25:
15:
13:
10:
9:
6:
4:
3:
2:
3353:
3342:
3339:
3337:
3334:
3333:
3331:
3314:
3309:
3305:
3301:
3296:
3292:
3288:
3283:
3279:
3275:
3269:
3265:
3260:
3256:
3252:
3247:
3243:
3239:
3234:
3230:
3226:
3221:
3217:
3213:
3208:
3204:
3200:
3195:
3191:
3187:
3182:
3178:
3174:
3169:
3165:
3161:
3156:
3152:
3148:
3143:
3139:
3135:
3130:
3126:
3122:
3117:
3113:
3109:
3104:
3100:
3095:
3089:
3085:
3080:
3074:
3070:
3069:
3067:
3065:
3061:
3057:
3052:
3042:
3037:
3019:
3013:
3011:
3005:
3003:
2997:
2995:
2989:
2987:
2984:
2982:
2976:
2975:
2973:
2971:
2967:
2961:
2958:
2956:
2950:
2948:
2945:
2943:
2940:
2938:
2932:
2930:
2927:
2926:
2924:
2922:
2918:
2912:
2909:
2907:
2903:Sphaeritidae
2901:
2899:
2893:
2892:
2890:
2888:
2884:
2881:
2879:
2875:
2865:
2859:
2857:
2853:Scarabaeidae
2851:
2849:
2843:
2841:
2835:
2833:
2827:
2825:
2819:
2817:
2811:
2809:
2803:
2801:
2795:
2793:
2791:(dor beetles)
2787:
2785:
2779:
2777:
2774:
2772:
2769:
2762:
2761:
2759:
2757:
2756:Scarabaeoidea
2753:
2750:
2748:
2744:
2734:
2731:
2729:
2723:
2721:
2718:
2711:
2709:
2706:
2705:
2703:
2701:
2697:
2691:
2688:
2681:
2680:
2678:
2676:
2672:
2666:
2660:
2658:
2655:
2653:
2650:
2648:
2642:
2640:
2634:
2632:
2626:
2624:
2618:
2616:
2613:
2611:
2605:
2603:
2597:
2595:
2591:Cerophytidae
2589:
2587:
2581:
2579:
2573:
2571:
2565:
2564:
2562:
2560:
2556:
2550:
2546:Rhipiceridae
2544:
2542:
2536:
2535:
2533:
2531:
2527:
2521:
2518:
2516:
2510:
2508:
2502:
2500:
2494:
2492:
2486:
2484:
2480:Eulichadidae
2478:
2476:
2470:
2468:
2462:
2460:
2459:Cneoglossidae
2457:
2455:
2449:
2447:
2441:
2439:
2433:
2432:
2430:
2428:
2424:
2418:
2417:Schizopodidae
2415:
2413:
2407:
2406:
2404:
2402:
2398:
2395:
2393:
2392:Elateriformia
2389:
2379:
2373:
2371:
2368:
2366:
2363:
2361:
2357:Tetratomidae
2355:
2353:
2347:
2345:
2339:
2337:
2331:
2329:
2323:
2321:
2315:
2313:
2309:Ripiphoridae
2307:
2305:
2299:
2297:
2293:Pyrochroidae
2291:
2289:
2288:Pterogeniidae
2286:
2284:
2278:
2276:
2273:
2271:
2265:
2263:
2257:
2255:
2249:
2247:
2241:
2239:
2233:
2231:
2227:Melandryidae
2225:
2223:
2217:
2215:
2214:Chalcodryidae
2212:
2210:
2204:
2202:
2196:
2194:
2188:
2186:
2180:
2179:
2177:
2175:
2171:
2165:
2159:
2158:
2156:
2154:
2150:
2144:
2140:Nemonychidae
2138:
2136:
2130:
2128:
2125:
2123:
2117:
2115:
2109:
2107:
2101:
2099:
2093:
2092:
2090:
2080:
2076:
2070:
2067:
2065:
2059:
2057:
2051:
2049:
2043:
2041:
2038:
2036:
2035:Propalticidae
2033:
2031:
2030:Priasilphidae
2028:
2026:
2023:
2021:
2015:
2013:
2009:Passandridae
2007:
2005:
2003:(sap beetles)
1999:
1997:
1994:
1992:
1986:
1984:
1981:
1974:
1972:
1966:
1964:
1958:
1956:
1953:
1951:
1948:
1946:
1940:
1938:
1935:
1933:
1930:
1928:
1922:
1920:
1914:
1912:
1911:Cavognathidae
1909:
1907:
1904:
1902:
1899:
1892:
1891:
1889:
1887:
1883:
1877:
1874:
1872:
1869:
1867:
1864:
1862:
1856:
1854:
1848:
1846:
1845:Eupsilobiidae
1843:
1841:
1837:Endomychidae
1835:
1833:
1830:
1828:
1824:Corylophidae
1822:
1820:
1814:
1812:
1806:
1804:
1798:
1796:
1793:
1791:
1788:
1786:
1783:
1782:
1780:
1778:
1774:
1768:
1762:
1760:
1757:
1755:
1752:
1750:
1747:
1745:
1742:
1740:
1739:Protopeltidae
1737:
1735:
1734:Prionoceridae
1732:
1730:
1727:
1725:
1722:
1715:
1713:
1710:
1708:
1705:
1698:
1696:
1690:
1688:
1687:Mauroniscidae
1685:
1683:
1680:
1678:
1672:
1670:
1667:
1665:
1659:
1657:
1651:
1649:
1646:
1639:
1638:
1636:
1634:
1630:
1624:
1621:
1619:
1616:
1614:
1611:
1609:
1608:Megalopodidae
1606:
1604:
1601:
1599:
1593:
1591:
1587:Cerambycidae
1585:
1584:
1582:
1580:
1576:
1573:
1571:
1567:
1557:
1553:Derodontidae
1551:
1550:
1548:
1546:
1545:Derodontoidea
1542:
1536:
1530:
1528:
1522:
1520:
1514:
1512:
1511:Endecatomidae
1509:
1507:
1501:
1499:
1495:Bostrichidae
1493:
1492:
1490:
1488:
1487:Bostrichoidea
1484:
1481:
1479:
1475:
1471:
1467:
1461:
1457:
1447:
1444:
1442:
1441:Sphaeriusidae
1439:
1437:
1434:
1432:
1426:
1425:
1422:
1418:
1412:
1408:
1394:
1388:
1386:
1380:
1378:
1375:
1368:
1366:
1363:
1361:
1355:
1353:
1347:
1345:
1339:
1337:
1333:Cicindelidae
1331:
1329:
1323:
1321:
1318:
1316:
1310:
1309:
1307:
1304:
1300:
1296:
1292:
1286:
1282:
1272:
1269:
1267:
1264:
1262:
1259:
1252:
1250:
1244:
1242:
1239:
1232:
1231:
1228:
1224:
1218:
1214:
1208:
1204:
1202:
1198:
1196:
1192:
1190:
1186:
1184:
1180:
1178:
1174:
1173:
1170:
1165:
1157:
1152:
1150:
1145:
1143:
1138:
1137:
1134:
1123:
1119:
1114:
1109:
1104:
1099:
1095:
1091:
1087:
1080:
1077:
1066:
1062:
1055:
1052:
1039:
1035:
1028:
1025:
1012:
1008:
1001:
998:
993:
989:
985:
981:
974:
971:
966:
962:
961:Insecta Mundi
958:
951:
948:
943:
939:
935:
931:
924:
921:
916:
912:
909:(2): 99–197.
908:
904:
897:
894:
889:
882:
879:
874:
870:
866:
862:
855:
852:
847:
840:
837:
824:
820:
816:
810:
807:
802:
795:
792:
786:
779:
776:
769:
767:
760:
759:
752:
747:
743:
742:
735:
730:
726:
725:
718:
713:
709:
708:
701:
696:
692:
691:
690:Odontophloeus
684:
679:
674:
672:
670:
662:
660:
657:
656:
652:) related to
651:
650:
649:Metaxyphloeus
645:
644:
639:
638:
633:
632:
627:
623:
615:
613:
612:(Grouvelle).
611:
610:
605:
604:
599:
598:
593:
592:
587:
586:Curculionidae
583:
579:
578:
573:
572:
563:
561:
554:
552:
549:
545:
541:
533:
529:
528:
520:
519:
511:
510:
502:
501:
493:
492:
484:
483:
475:
474:
473:Rhabdophloeus
466:
465:
464:Pseudophloeus
457:
456:
450:MacLeay, 1871
448:
447:
439:
438:
430:
429:
421:
420:
412:
411:
403:
402:
394:
393:
392:Odontophloeus
385:
384:
376:
375:
367:
366:
358:
357:
351:Reitter, 1874
349:
348:
340:
339:
338:Metaxyphloeus
331:
330:
322:
321:
313:
312:
304:
303:
295:
294:
293:Lepidophloeus
286:
285:
277:
276:
268:
267:
259:
258:
250:
249:
241:
240:
232:
231:
223:
222:
214:
213:
205:
204:
196:
195:
194:Carinophloeus
187:
186:
178:
177:
169:
168:
162:
159:
155:
151:
146:
143:
140:
139:
136:
133:
131:Superfamily:
130:
129:
126:
123:
120:
119:
116:
113:
110:
109:
106:
103:
100:
99:
96:
93:
90:
89:
86:
83:
80:
79:
76:
73:
70:
69:
66:
63:
60:
59:
54:
49:
45:
42:
41:
36:
32:
27:
22:
19:
3063:
2942:Helophoridae
2934:Georissidae
2929:Epimetopidae
2845:Pleocomidae
2829:Ochodaeidae
2813:Hybosoridae
2797:Glaphyridae
2789:Geotrupidae
2767:
2764:Belohinidae
2725:Eucinetidae
2716:
2713:Decliniidae
2686:
2644:Phengodidae
2583:Cantharidae
2538:Dascillidae
2530:Dascilloidea
2512:Psephenidae
2504:Lutrochidae
2496:Limnichidae
2409:Buprestidae
2401:Buprestoidea
2341:Synchroidae
2325:Scraptiidae
2317:Salpingidae
2280:Prostomidae
2267:Oedemeridae
2243:Mordellidae
2161:Lymexylidae
2153:Lymexyloidea
2103:Attelabidae
2095:Anthribidae
2053:Smicripidae
2017:Phalacridae
2001:Nitidulidae
1996:Myraboliidae
1988:Monotomidae
1979:
1967:
1960:Kateretidae
1937:Cyclaxyridae
1897:
1894:Agapythidae
1858:Latridiidae
1808:Cerylonidae
1795:Anamorphidae
1729:Phycosecidae
1720:
1703:
1700:Metaxinidae
1653:Biphyllidae
1644:
1613:Orsodacnidae
1570:Cucujiformia
1503:Dermestidae
1373:
1312:Amphizoidae
1257:
1237:
1223:Archostemata
1207:Holometabola
1205:Superorder:
1199:Infraclass:
1093:
1089:
1079:
1068:. Retrieved
1064:
1054:
1042:. Retrieved
1038:the original
1027:
1015:. Retrieved
1011:the original
1000:
986:(1): 67–83.
983:
979:
973:
964:
960:
950:
933:
929:
923:
906:
902:
896:
887:
881:
864:
860:
854:
845:
839:
827:. Retrieved
823:the original
818:
809:
800:
794:
789:xiv + 861pp.
784:
778:
765:
756:
739:
722:
705:
688:
668:
666:
655:Laemophloeus
653:
647:
643:Rhinophloeus
641:
635:
631:Microbrontes
629:
625:
622:Cryptolestes
621:
619:
607:
601:
600:(Stephens),
595:
591:Cryptolestes
589:
582:bark beetles
575:
571:Leptophloeus
569:
567:
558:
539:
538:
525:
516:
513:Kessel, 1921
507:
500:Rhinophloeus
498:
489:
480:
471:
462:
453:
444:
437:Phloeolaemus
435:
426:
423:Kessel, 1921
417:
414:Thomas, 2018
408:
399:
396:Thomas, 1984
390:
381:
378:Sasaji, 1983
372:
363:
354:
347:Microbrontes
345:
342:Thomas, 1984
336:
327:
318:
309:
302:Leptophloeus
300:
297:Thomas, 1984
291:
282:
275:Laemophloeus
273:
264:
255:
246:
239:Deinophloeus
237:
228:
221:Cryptolestes
219:
212:Charaphloeus
210:
201:
192:
185:Brontolaemus
183:
174:
171:Thomas, 2010
165:
164:
144:
125:Cucujiformia
121:Infraorder:
38:
18:
3207:iNaturalist
3088:Wikispecies
2986:Hydraenidae
2960:Spercheidae
2947:Hydrochidae
2911:Synteliidae
2895:Histeridae
2887:Histeroidea
2837:Passalidae
2805:Glaresidae
2662:Throscidae
2622:(fireflies)
2620:Lampyridae
2607:Eucnemidae
2599:Elateridae
2559:Elateroidea
2375:Zopheridae
2259:Mycteridae
2190:Anthicidae
2061:Sphindidae
2045:Silvanidae
1955:Hobartiidae
1942:Erotylidae
1871:Mycetaeidae
1866:Murmidiidae
1850:Euxestidae
1744:Rentoniidae
1618:Oxypeltidae
1603:Disteniidae
1436:Lepiceridae
1365:Hygrobiidae
1357:Haliplidae
1341:Dytiscidae
1320:Aspidytidae
1032:Thomas MC.
1005:Thomas MC.
527:Xylophloeus
504:Sharp, 1899
486:Steel, 1954
482:Rhinolaemus
477:Sharp, 1899
455:Planolestes
441:Casey, 1916
432:Casey, 1916
428:Phloeipsius
356:Microlaemus
329:Mestolaemus
320:Mariolaemus
311:Magnoleptus
306:Casey, 1916
266:Heterojinus
252:Casey, 1884
243:Sharp, 1899
234:Arrow, 1920
216:Casey, 1916
189:Sharp, 1885
3330:Categories
3007:Silphidae
2999:Ptiliidae
2991:Leiodidae
2978:Agyrtidae
2821:Lucanidae
2700:Scirtoidea
2636:Omethidae
2615:Jurasaidae
2464:Dryopidae
2435:Byrrhidae
2427:Byrrhoidea
2119:Brentidae
1924:Cucujidae
1906:Boganiidae
1886:Cucujoidea
1759:Thymalidae
1749:Rhadalidae
1692:Melyridae
1661:Byturidae
1623:Vesperidae
1532:Ptiniidae
1382:Noteridae
1349:Gyrinidae
1325:Carabidae
1254:Jurodidae
1246:Cupedidae
1193:Subclass:
1183:Arthropoda
1164:Coleoptera
1070:2022-01-22
1044:31 January
1017:31 January
770:References
637:Rhinomalus
548:Cucujoidea
518:Xylolestes
495:Gemm, 1870
491:Rhinomalus
446:Placonotus
401:Parandrita
383:Notolaemus
365:Narthecius
203:Caulonomus
135:Cucujoidea
111:Suborder:
105:Coleoptera
85:Arthropoda
2861:Trogidae
2855:(scarabs)
2733:Scirtidae
2708:Clambidae
2301:Pythidae
2235:Meloidae
2182:Aderidae
2085:(weevils)
1950:Helotidae
1876:Teredidae
1790:Alexiidae
1674:Cleridae
1633:Cleroidea
1466:Polyphaga
1464:Suborder
1417:Myxophaga
1415:Suborder
1370:Meruidae
1289:Suborder
1271:Ommatidae
1221:Suborder
1195:Pterygota
1175:Kingdom:
284:Lathropus
230:Cucujinus
115:Polyphaga
71:Kingdom:
65:Eukaryota
3129:BugGuide
3079:Q1040629
3073:Wikidata
2628:Lycidae
2472:Elmidae
2370:Ulodidae
2206:Boridae
2111:Belidae
1712:Peltidae
1305:families
1291:Adephaga
1201:Neoptera
1181:Phylum:
1177:Animalia
1166:families
1122:35045081
1090:PLOS ONE
669:PLOS ONE
626:Dysmerus
577:Dysmerus
248:Dysmerus
141:Family:
81:Phylum:
75:Animalia
61:Domain:
2219:Ciidae
2127:Caridae
1189:Insecta
1187:Class:
1162:Extant
1113:8769368
992:4008144
967:: 1–30.
873:3999883
675:Gallery
544:beetles
101:Order:
95:Insecta
91:Class:
3287:686900
3271:NZOR:
3264:196984
3238:678356
3225:117132
3212:174146
3121:312278
1303:Extant
1120:
1110:
990:
871:
829:15 May
646:, and
628:, and
257:Gannes
176:Blubos
158:Genera
3308:Plazi
3300:70402
3220:IRMNG
3173:11048
3134:15839
988:JSTOR
869:JSTOR
3259:NCBI
3233:ITIS
3199:7832
3194:GBIF
3160:7488
3116:BOLD
1118:PMID
1046:2014
1019:2014
831:2012
3246:NBN
3155:EoL
3147:BQH
3142:CoL
3103:AFD
1108:PMC
1098:doi
938:doi
911:doi
693:sp.
3332::
3310::
3297::
3284::
3261::
3248::
3235::
3222::
3209::
3196::
3183::
3170::
3157::
3144::
3131::
3118::
3105::
3090::
3075::
1116:.
1106:.
1094:17
1092:.
1088:.
1063:.
984:38
982:.
965:74
963:.
959:.
934:27
932:.
907:29
905:.
865:31
863:.
817:.
640:,
624:,
574:,
2770:)
2766:(
2719:)
2715:(
2689:)
2685:(
1982:)
1978:(
1900:)
1896:(
1723:)
1719:(
1706:)
1702:(
1647:)
1643:(
1376:)
1372:(
1260:)
1256:(
1240:)
1236:(
1155:e
1148:t
1141:v
1124:.
1100::
1073:.
1048:.
1021:.
994:.
944:.
940::
917:.
913::
875:.
833:.
761:.
744:.
727:.
710:.
584:(
Text is available under the Creative Commons Attribution-ShareAlike License. Additional terms may apply.
