31:
51:
145:
499:
Yoshizawa, Kazunori, Charles
Lienhard, and Kevin P. Johnson. "Molecular systematics of the suborder Trogiomorpha (Insecta: Psocodea:‘Psocoptera’)." Zoological Journal of the Linnean Society 146.2 (2006):
509:
Enderlein, Gunther. "The Scaly Winged
Copeognatha (Monograph of the Amphientomidae, Lepidopsocidae, and Lepidillidae in relation to their Morphology and Taxonomy." Spolia Zeylandica 4 (1906): 39-122.
1622:
1661:
1596:
1635:
206:
The Scaly Winged
Copeognatha (Monograph of the Amphientomidae, Lepidopsocidae, and Lepidillidae in relation to their Morphology and Taxonomy)
619:
169:. Colloquially, Lepidopsocidae are referred to as the Scaly-winged Barklice. There are more than 220 species described worldwide.
30:
1640:
1648:
50:
1748:
1518:
1505:
1666:
1575:
1588:
1725:
523:
1458:
612:
1132:
1753:
1697:
1523:
1109:
935:
1715:
1031:
1026:
1676:
1557:
1000:
907:
333:
182:
413:
303:
224:
216:
1496:
945:
897:
353:
253:
245:
232:
1099:
1036:
1013:
955:
912:
313:
283:
1094:
393:
373:
273:
45:
1702:
1342:
1114:
1104:
1070:
403:
363:
237:
433:
323:
1684:
1544:
1380:
1294:
979:
902:
823:
605:
173:
DNA analysis of relationships between families in
Trogiomorpha propose that Lepidopsocidae is
1627:
383:
1309:
987:
736:
728:
475:
464:
158:
1689:
1266:
922:
860:
178:
1583:
1570:
1385:
1357:
1319:
1289:
1261:
1127:
1005:
950:
855:
850:
828:
813:
803:
795:
759:
751:
443:
1510:
144:
1742:
1413:
1375:
1367:
1337:
1304:
1271:
1256:
1246:
1238:
1228:
1220:
1119:
1018:
965:
1549:
1403:
1352:
1347:
1327:
1251:
1177:
1150:
1142:
1054:
992:
838:
818:
783:
713:
688:
672:
343:
198:
190:
174:
170:
166:
112:
1653:
1609:
1490:
1408:
1332:
1299:
1281:
1208:
927:
808:
423:
293:
209:
1481:
1438:
1185:
1173:
1080:
453:
1181:
1046:
1041:
940:
917:
889:
718:
700:
660:
648:
194:
162:
122:
82:
62:
597:
1531:
1475:
1418:
1395:
666:
629:
102:
1536:
577:
1062:
1614:
654:
642:
92:
72:
1601:
1452:
550:
1710:
1442:
874:
186:
143:
1456:
1205:
780:
685:
601:
1562:
267:
204:
Lepidopsocidae was described by
Gunther Enderlein in
1465:
1394:
1366:
1318:
1280:
1237:
1219:
1141:
1079:
964:
888:
873:
837:
794:
750:
727:
699:
189:covering the body and forewings of Lepodopsocids.
613:
522:Johnson, Kevin P.; Smith, Vincent S. (2019).
517:
515:
8:
1437:Order Psocodea is comprised of paraphyletic
177:, meaning that the taxa within all share a
1453:
1216:
1202:
885:
870:
843:
791:
777:
696:
682:
620:
606:
598:
29:
20:
528:Psocodea species file online, Version 5.0
572:
570:
1445:, both previously classified as orders.
492:
545:
543:
7:
1716:C64E87D9-EF21-152F-FFC7-FBF15E3095E0
1677:a4b79977-c80b-4402-b5f9-372d3453589b
1589:10a45696-9d43-4f2f-839e-a401957953be
1439:Psocoptera (book lice or bark lice)
578:"Lepidopsocidae Family Information"
193:families of Lepidoposcidae include
14:
49:
1184:comprise paraphyletic suborder
185:by the presence of scales and
1:
1723:Psocodea Species File (old):
212:of Lepidopsocidae include:
1770:
165:belonging to the suborder
1435:
1215:
1201:
1186:Mallophaga (chewing lice)
1171:
1066:(crab lice or pubic lice)
869:
846:
790:
776:
695:
681:
637:
46:Scientific classification
44:
37:
28:
23:
524:"family Lepidopsocidae"
1050:(body lice, head lice)
151:
1698:Paleobiology Database
208:, published in 1906.
149:Echmepteryx hamiltoni
147:
1584:Fauna Europaea (new)
181:. This is supported
1749:Psocoptera families
1443:Phthiraptera (lice)
1133:Trichophilopteridae
931:(chicken body lice)
1110:Heptapsogasteridae
152:
39:Echmepteryx hageni
16:Family of booklice
1736:
1735:
1685:Open Tree of Life
1459:Taxon identifiers
1450:
1449:
1441:and monophyletic
1431:
1430:
1427:
1426:
1381:Pseudocaeciliidae
1295:Dolabellopsocidae
1197:
1196:
1193:
1192:
1167:
1166:
1163:
1162:
1159:
1158:
980:Echinophthiriidae
936:Pseudomenoponidae
824:Protroctopsocidae
772:
771:
768:
767:
481:
470:
459:
449:
439:
429:
419:
409:
399:
389:
379:
369:
359:
349:
339:
329:
319:
309:
299:
289:
279:
258:
250:
242:
229:
221:
142:
141:
138:
1761:
1729:
1728:
1719:
1718:
1706:
1705:
1693:
1692:
1680:
1679:
1670:
1669:
1657:
1656:
1654:NBNSYS0000160291
1644:
1643:
1631:
1630:
1618:
1617:
1605:
1604:
1592:
1591:
1579:
1578:
1566:
1565:
1553:
1552:
1540:
1539:
1527:
1526:
1514:
1513:
1501:
1500:
1499:
1486:
1485:
1484:
1454:
1310:Spurostigmatidae
1217:
1203:
1124:
1088:
1087:
1067:
1059:
1058:(spiny rat lice)
1051:
1032:Neolinognathidae
1027:Microthoraciidae
1023:
1010:
997:
988:Enderleinellidae
984:
973:
972:
932:
886:
882:
881:
871:
844:
792:
778:
743:
742:
737:Prionoglarididae
729:Prionoglaridetae
697:
683:
622:
615:
608:
599:
592:
591:
589:
588:
574:
565:
564:
562:
561:
551:"Lepidopsocidae"
547:
538:
537:
535:
534:
519:
510:
507:
501:
497:
479:
468:
457:
447:
437:
427:
417:
407:
397:
387:
377:
367:
357:
347:
337:
327:
317:
307:
297:
287:
277:
256:
248:
240:
227:
219:
136:
54:
53:
33:
21:
1769:
1768:
1764:
1763:
1762:
1760:
1759:
1758:
1739:
1738:
1737:
1732:
1724:
1722:
1714:
1709:
1701:
1696:
1688:
1683:
1675:
1673:
1665:
1660:
1652:
1647:
1639:
1634:
1626:
1621:
1613:
1608:
1600:
1595:
1587:
1582:
1574:
1569:
1561:
1556:
1548:
1543:
1535:
1530:
1522:
1517:
1509:
1504:
1495:
1494:
1489:
1480:
1479:
1474:
1461:
1451:
1446:
1423:
1390:
1362:
1314:
1276:
1267:Paracaeciliidae
1233:
1211:
1189:
1178:Rhyncophthirina
1155:
1143:Rhyncophthirina
1137:
1122:
1085:
1084:
1083:
1075:
1065:
1057:
1049:
1021:
1009:(armoured lice)
1008:
1001:Hamophthiriidae
996:(ungulate lice)
995:
982:
970:
969:
968:
960:
930:
923:Laemobothriidae
908:Colpocephalidae
879:
878:
877:
865:
861:Sphaeropsocidae
833:
786:
764:
746:
740:
739:
723:
691:
677:
633:
626:
596:
595:
586:
584:
576:
575:
568:
559:
557:
549:
548:
541:
532:
530:
521:
520:
513:
508:
504:
498:
494:
489:
484:
469:Enderlein, 1906
458:Enderlein, 1911
448:Enderlein, 1904
438:Enderlein, 1906
428:Enderlein, 1922
368:Enderlein, 1910
358:Enderlein, 1906
334:Neolepidopsocus
328:Enderlein, 1906
318:Enderlein, 1903
288:Enderlein, 1903
265:
249:Enderlein, 1903
228:Enderlein, 1903
220:Enderlein, 1906
183:morphologically
179:common ancestor
137:Enderlein, 1903
135:
48:
24:Lepidopsocidae
17:
12:
11:
5:
1767:
1765:
1757:
1756:
1751:
1741:
1740:
1734:
1733:
1731:
1730:
1720:
1707:
1694:
1681:
1671:
1658:
1645:
1632:
1619:
1606:
1593:
1580:
1571:Fauna Europaea
1567:
1554:
1541:
1528:
1515:
1511:Lepidopsocidae
1502:
1497:Lepidopsocidae
1487:
1471:
1469:
1467:Lepidopsocidae
1463:
1462:
1457:
1448:
1447:
1436:
1433:
1432:
1429:
1428:
1425:
1424:
1422:
1421:
1416:
1411:
1406:
1400:
1398:
1392:
1391:
1389:
1388:
1386:Trichopsocidae
1383:
1378:
1372:
1370:
1364:
1363:
1361:
1360:
1358:Sabulopsocidae
1355:
1350:
1345:
1340:
1335:
1330:
1324:
1322:
1320:Homilopsocidea
1316:
1315:
1313:
1312:
1307:
1302:
1297:
1292:
1290:Cladiopsocidae
1286:
1284:
1278:
1277:
1275:
1274:
1269:
1264:
1262:Dasydemellidae
1259:
1254:
1249:
1243:
1241:
1235:
1234:
1232:
1231:
1225:
1223:
1213:
1212:
1206:
1199:
1198:
1195:
1194:
1191:
1190:
1172:
1169:
1168:
1165:
1164:
1161:
1160:
1157:
1156:
1154:
1153:
1151:Haematomyzidae
1147:
1145:
1139:
1138:
1136:
1135:
1130:
1128:Trichodectidae
1125:
1123:(paraphyletic)
1117:
1112:
1107:
1102:
1097:
1091:
1089:
1086:(paraphyletic)
1077:
1076:
1074:
1073:
1068:
1060:
1052:
1044:
1039:
1034:
1029:
1024:
1016:
1011:
1006:Hoplopleuridae
1003:
998:
993:Haematopinidae
990:
985:
976:
974:
971:(sucking lice)
962:
961:
959:
958:
953:
951:Trimenoponidae
948:
943:
938:
933:
925:
920:
915:
910:
905:
900:
894:
892:
883:
867:
866:
864:
863:
858:
856:Pachytroctidae
853:
851:Liposcelididae
847:
841:
835:
834:
832:
831:
829:Troctopsocidae
826:
821:
816:
814:Manicapsocidae
811:
806:
804:Amphientomidae
800:
798:
796:Amphientometae
788:
787:
781:
774:
773:
770:
769:
766:
765:
763:
762:
760:Psyllipsocidae
756:
754:
752:Psyllipsocetae
748:
747:
745:
744:
741:(paraphyletic)
733:
731:
725:
724:
722:
721:
716:
711:
709:Lepidopsocidae
705:
703:
693:
692:
686:
679:
678:
676:
675:
669:
663:
657:
651:
645:
638:
635:
634:
627:
625:
624:
617:
610:
602:
594:
593:
566:
539:
511:
502:
491:
490:
488:
485:
483:
482:
471:
460:
450:
440:
430:
420:
414:Pseudothylacus
410:
408:Badonnel, 1949
400:
390:
388:Thornton, 1962
380:
378:Mockford, 2005
370:
360:
350:
348:Mockford, 1993
340:
330:
320:
310:
304:Illepidopsocus
300:
290:
280:
269:
264:
261:
260:
259:
251:
243:
241:Mockford, 2005
235:
230:
225:Lepidopsocinae
222:
217:Echinopsocinae
157:are an insect
155:Lepidopsocidae
140:
139:
133:Lepidopsocidae
130:
126:
125:
120:
116:
115:
110:
106:
105:
100:
96:
95:
90:
86:
85:
80:
76:
75:
70:
66:
65:
60:
56:
55:
42:
41:
35:
34:
26:
25:
15:
13:
10:
9:
6:
4:
3:
2:
1766:
1755:
1752:
1750:
1747:
1746:
1744:
1727:
1721:
1717:
1712:
1708:
1704:
1699:
1695:
1691:
1686:
1682:
1678:
1672:
1668:
1663:
1659:
1655:
1650:
1646:
1642:
1637:
1633:
1629:
1624:
1620:
1616:
1611:
1607:
1603:
1598:
1594:
1590:
1585:
1581:
1577:
1572:
1568:
1564:
1559:
1555:
1551:
1546:
1542:
1538:
1533:
1529:
1525:
1520:
1516:
1512:
1507:
1503:
1498:
1492:
1488:
1483:
1477:
1473:
1472:
1470:
1468:
1464:
1460:
1455:
1444:
1440:
1434:
1420:
1417:
1415:
1414:Psilopsocidae
1412:
1410:
1407:
1405:
1402:
1401:
1399:
1397:
1393:
1387:
1384:
1382:
1379:
1377:
1376:Philotarsidae
1374:
1373:
1371:
1369:
1368:Philotarsetae
1365:
1359:
1356:
1354:
1351:
1349:
1346:
1344:
1341:
1339:
1338:Lachesillidae
1336:
1334:
1331:
1329:
1326:
1325:
1323:
1321:
1317:
1311:
1308:
1306:
1305:Ptiloneuridae
1303:
1301:
1298:
1296:
1293:
1291:
1288:
1287:
1285:
1283:
1279:
1273:
1272:Stenopsocidae
1270:
1268:
1265:
1263:
1260:
1258:
1257:Caeciliusidae
1255:
1253:
1250:
1248:
1247:Amphipsocidae
1245:
1244:
1242:
1240:
1239:Caeciliusetae
1236:
1230:
1229:Archipsocidae
1227:
1226:
1224:
1222:
1221:Archipsocetae
1218:
1214:
1210:
1204:
1200:
1187:
1183:
1179:
1175:
1170:
1152:
1149:
1148:
1146:
1144:
1140:
1134:
1131:
1129:
1126:
1121:
1120:Philopteridae
1118:
1116:
1113:
1111:
1108:
1106:
1103:
1101:
1098:
1096:
1093:
1092:
1090:
1082:
1078:
1072:
1069:
1064:
1061:
1056:
1053:
1048:
1045:
1043:
1040:
1038:
1035:
1033:
1030:
1028:
1025:
1020:
1019:Linognathidae
1017:
1015:
1012:
1007:
1004:
1002:
999:
994:
991:
989:
986:
981:
978:
977:
975:
967:
963:
957:
954:
952:
949:
947:
946:Somaphantidae
944:
942:
939:
937:
934:
929:
926:
924:
921:
919:
916:
914:
911:
909:
906:
904:
901:
899:
898:Ancistronidae
896:
895:
893:
891:
887:
884:
876:
872:
868:
862:
859:
857:
854:
852:
849:
848:
845:
842:
840:
836:
830:
827:
825:
822:
820:
817:
815:
812:
810:
807:
805:
802:
801:
799:
797:
793:
789:
785:
779:
775:
761:
758:
757:
755:
753:
749:
738:
735:
734:
732:
730:
726:
720:
717:
715:
712:
710:
707:
706:
704:
702:
698:
694:
690:
684:
680:
674:
670:
668:
664:
662:
658:
656:
652:
650:
646:
644:
640:
639:
636:
631:
623:
618:
616:
611:
609:
604:
603:
600:
583:
579:
573:
571:
567:
556:
552:
546:
544:
540:
529:
525:
518:
516:
512:
506:
503:
496:
493:
486:
478:
477:
472:
467:
466:
461:
456:
455:
451:
446:
445:
441:
436:
435:
431:
426:
425:
421:
416:
415:
411:
406:
405:
401:
396:
395:
391:
386:
385:
381:
376:
375:
371:
366:
365:
361:
356:
355:
354:Nepticulomima
351:
346:
345:
341:
336:
335:
331:
326:
325:
321:
316:
315:
311:
306:
305:
301:
296:
295:
291:
286:
285:
281:
276:
275:
271:
270:
268:
262:
257:Roesler, 1944
255:
254:Thylacellinae
252:
247:
246:Perientominae
244:
239:
236:
234:
233:Lepolepidinae
231:
226:
223:
218:
215:
214:
213:
211:
207:
202:
200:
196:
192:
188:
184:
180:
176:
172:
168:
164:
160:
156:
150:
146:
134:
131:
128:
127:
124:
121:
118:
117:
114:
111:
108:
107:
104:
101:
98:
97:
94:
91:
88:
87:
84:
81:
78:
77:
74:
71:
68:
67:
64:
61:
58:
57:
52:
47:
43:
40:
36:
32:
27:
22:
19:
1754:Trogiomorpha
1466:
1404:Hemipsocidae
1353:Peripsocidae
1348:Mesopsocidae
1328:Ectopsocidae
1252:Asiopsocidae
1100:Dasyonygidae
1055:Polyplacidae
1037:Pecaroecidae
1014:Hybothiridae
956:Trinotonidae
913:Gliricolidae
875:Phthiraptera
839:Nanopsocetae
819:Musapsocidae
784:Troctomorpha
714:Psoquillidae
708:
689:Trogiomorpha
673:Paraneoptera
671:Superorder:
665:Infraclass:
585:. Retrieved
582:BugGuide.net
581:
558:. Retrieved
554:
531:. Retrieved
527:
505:
495:
474:
463:
452:
442:
432:
422:
412:
402:
392:
382:
372:
362:
352:
344:Neolepolepis
342:
332:
322:
314:Lepidopsocus
312:
302:
292:
284:Echinopsocus
282:
272:
266:
205:
203:
199:Psoquillidae
175:monophyletic
171:Phylogenetic
167:Trogiomorpha
154:
153:
148:
132:
119:Infraorder:
113:Trogiomorpha
38:
18:
1610:iNaturalist
1491:Wikispecies
1409:Myopsocidae
1333:Elipsocidae
1300:Epipsocidae
1282:Epipsocetae
1209:Psocomorpha
1095:Bovicolidae
1047:Pediculidae
1042:Pedicinidae
1022:(pale lice)
983:(seal lice)
928:Menoponidae
809:Compsocidae
480:Hagen, 1866
424:Pteroxanium
398:Hagen, 1865
394:Perientomum
374:Papuapsocus
298:Aaron, 1886
294:Echmepteryx
278:Banks, 1931
274:Cyptophania
210:Subfamilies
1743:Categories
1343:Lesneiidae
1174:Ischnocera
1115:Lipeuridae
1105:Goniodidae
1081:Ischnocera
1071:Ratemiidae
659:Subclass:
649:Arthropoda
587:2018-04-21
560:2018-04-21
533:2019-05-04
487:References
454:Thylacella
404:Proentomum
364:Notolepium
238:Parasoinae
109:Suborder:
83:Arthropoda
1207:Suborder
1182:Amblycera
1063:Pthiridae
941:Ricinidae
918:Gyropidae
890:Amblycera
782:Suborder
719:Trogiidae
701:Atropetae
687:Suborder
661:Pterygota
641:Kingdom:
434:Scolopama
324:Lepolepis
195:Trogiidae
163:bark lice
123:Atropetae
69:Kingdom:
63:Eukaryota
1532:BugGuide
1482:Q4042775
1476:Wikidata
1419:Psocidae
1396:Psocetae
966:Anoplura
903:Boopidae
667:Neoptera
647:Phylum:
643:Animalia
632:families
630:Psocodea
500:287-299.
418:Li, 2002
338:Li, 1992
308:Li, 1994
129:Family:
103:Psocodea
79:Phylum:
73:Animalia
59:Domain:
1726:1191766
655:Insecta
653:Class:
628:Extant
384:Parasoa
99:Order:
93:Insecta
89:Class:
1703:217783
1674:NZOR:
1667:159970
1641:103323
1628:100849
1615:245136
1537:145677
880:(lice)
476:Thylax
465:Lepium
263:Genera
191:Sister
159:family
1711:Plazi
1690:34121
1623:IRMNG
1576:12131
1550:8MPF3
1524:27248
187:setae
1662:NCBI
1636:ITIS
1602:4378
1597:GBIF
1563:1043
1519:BOLD
1180:and
555:GBIF
197:and
1649:NBN
1558:EoL
1545:CoL
1506:AFD
444:Soa
161:of
1745::
1713::
1700::
1687::
1664::
1651::
1638::
1625::
1612::
1599::
1586::
1573::
1560::
1547::
1534::
1521::
1508::
1493::
1478::
1176:,
580:.
569:^
553:.
542:^
526:.
514:^
473:†
462:†
201:.
1188:.
621:e
614:t
607:v
590:.
563:.
536:.
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