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Listriolobus pelodes

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and other micro-organisms form part of the diet. Other movements of the body wall draw water into and out of the anal vesicles and respiratory gases are exchanged. The worm periodically reverses its position in its burrow. The proboscis can extend to a length of 20 centimetres (7.9 in) and,
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movements and this washes the pellets out of the second aperture of the tube. They accumulate nearby in little piles, the presence of which may show where these spoon worms are buried. The faecal pellets get ingested during further feeding activities and it is probable that rapidly growing
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is found along the coast of North America ranging from northern California as far south as Baja California at depths between about 18 to 155 metres (59 to 509 ft). It lives in a burrow in fine grained sands and mud with a high content of organic matter.
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feeding from the two burrow apertures, each worm can sweep an area of sediment of 0.25 square metres (2.7 sq ft). The feeding activities of this worm, occurring as it does in large numbers, has a considerable effect on the seabed
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into juvenile spoon worms. They seem to be attracted to settle in areas of fine, nutrient-rich sediment but the mechanism for this discrimination is unclear. The juveniles become sexually mature in another six to twelve months.
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at densities of up to 1,500 individuals per square metre (11 square feet). The burrowing and feeding activities of these worms redistributed and aerated the sediment and promoted a more diverse community of
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has a plump, sausage-shaped body about 4 centimetres (1.6 in) long and 2 to 3 centimetres (0.79 to 1.18 in) wide. An extensible spoon-shaped
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with the ventral side upwards. Sediment is scooped up by the proboscis and is wafted along a central groove to the mouth by the action of
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lives in a U-shaped burrow in the sediment. Its body remains below the surface while it extends its flexible proboscis across the
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projects from the anterior (front) end of the body and the mouth is at its base on the ventral side. There is a pair of hooked
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is at the posterior end of the body. There are two anal vesicles on either side of the rear end of the gut which are used for
189:. It is found in shallow seas in the North East Pacific off the coast of California. It lives in a burrow in soft sediments. 654: 36: 302:. After several developmental stages over a period of about six months, the larvae settle on the seabed and undergo 560:(Echiura) on coastal shelf benthic communities and sediments modified by a major California wastewater discharge". 667: 220:. The body is a translucent green colour and there are eight longitudinal bundles of muscle in the body wall. 397:
area. The worms flourished for several years before disappearing from the area but their legacy of improved
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A number of other invertebrates take up occupation in this spoon worm's burrow and live there as
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that the burrow provides and the nutrient rich stream of water passing through the burrow.
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before being liberated into the sea in the late winter or spring. After fertilisation, the
208:(bristles) projecting from the ventral surface of the body behind the mouth and a pair of 81: 736: 575: 303: 289: 209: 401: 134: 701: 500:"The Annual Cycle of Oogenesis, Spawning, and Larval Settlement of the Echiuran 385: 296: 252: 114: 614: 381: 338: 124: 94: 394: 354: 347: 316: 285: 262: 248: 201: 48: 556:
Stull, Janet K.; Haydock, C.Irwin; Montagne, David E. (1986). "Effects of
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or body cavity. Here they mature and are then stored in the
598: 393:than would otherwise have existed in this heavily 346:. These probably benefit from the protection from 276:are located beside the anal vesicles and liberate 532:. Holt-Saunders International. pp. 870–873. 8: 586: 20: 493: 491: 489: 487: 485: 483: 481: 551: 549: 413: 7: 563:Estuarine, Coastal and Shelf Science 457:Encyclopedia of Marine Invertebrates 357:on by bottom feeding fish such as 14: 435:World Register of Marine Species 35: 1: 576:10.1016/0272-7714(86)90020-X 460:. TFH Publications. p.  764: 528:Barnes, Robert D. (1982). 319:. These include the small 268:The sexes are separate in 748:Animals described in 1946 163: 156: 32:Scientific classification 30: 23: 504:off Southern California" 498:Pilger, John F. (1980). 454:Walls, Jerry G. (1982). 370:Myliobatis californicus 185:is a species of marine 16:Species of annelid worm 384:discharge outlets off 364:Microstomus pacificus 600:Listriolobus pelodes 558:Listriolobus pelodes 530:Invertebrate Zoology 502:Listriolobus pelodes 425:Listriolobus pelodes 378:Listriolobus pelodes 270:Listriolobus pelodes 241:Listriolobus pelodes 229:Listriolobus pelodes 198:Listriolobus pelodes 182:Listriolobus pelodes 167:Listriolobus pelodes 25:Listriolobus pelodes 421:Murina, G. (2012). 353:The spoon worm is 730: 729: 689:Open Tree of Life 592:Taxon identifiers 343:Hesperonoe laevis 178: 177: 755: 723: 722: 710: 709: 697: 696: 684: 683: 671: 670: 658: 657: 645: 644: 632: 631: 619: 618: 617: 587: 580: 579: 553: 544: 543: 525: 519: 518: 508: 495: 476: 475: 451: 445: 444: 442: 441: 418: 367:) and bat rays ( 325:Pinnixa schmitti 169: 40: 39: 21: 763: 762: 758: 757: 756: 754: 753: 752: 733: 732: 731: 726: 718: 713: 705: 700: 692: 687: 679: 674: 666: 661: 653: 648: 640: 635: 627: 622: 613: 612: 607: 594: 584: 583: 555: 554: 547: 540: 527: 526: 522: 511:Pacific Science 506: 497: 496: 479: 472: 453: 452: 448: 439: 437: 420: 419: 415: 410: 380:was found near 339:polychaete worm 330:bivalve mollusc 313: 238: 226: 195: 174: 171: 165: 152: 149:L. pelodes 82:Pleistoannelida 34: 17: 12: 11: 5: 761: 759: 751: 750: 745: 735: 734: 728: 727: 725: 724: 711: 698: 685: 672: 659: 646: 633: 620: 604: 602: 596: 595: 590: 582: 581: 545: 538: 520: 477: 470: 446: 412: 411: 409: 406: 376:In the 1970s, 361:, Dover sole ( 334:Mysella tumida 312: 309: 237: 234: 225: 222: 210:nephridiopores 194: 191: 176: 175: 172: 161: 160: 154: 153: 146: 144: 140: 139: 132: 128: 127: 122: 118: 117: 112: 108: 107: 102: 98: 97: 92: 85: 84: 79: 72: 71: 66: 62: 61: 56: 52: 51: 46: 42: 41: 28: 27: 15: 13: 10: 9: 6: 4: 3: 2: 760: 749: 746: 744: 741: 740: 738: 721: 716: 712: 708: 703: 699: 695: 690: 686: 682: 677: 673: 669: 664: 660: 656: 651: 647: 643: 638: 634: 630: 625: 621: 616: 610: 606: 605: 603: 601: 597: 593: 588: 577: 573: 569: 565: 564: 559: 552: 550: 546: 541: 539:0-03-056747-5 535: 531: 524: 521: 517:(2): 128–142. 516: 512: 505: 503: 494: 492: 490: 488: 486: 484: 482: 478: 473: 471:0-86622-141-7 467: 463: 459: 458: 450: 447: 436: 432: 428: 427:Fisher, 1946" 426: 417: 414: 407: 405: 403: 400: 396: 392: 387: 383: 379: 374: 372: 371: 366: 365: 360: 356: 351: 349: 345: 344: 340: 336: 335: 331: 327: 326: 322: 318: 310: 308: 305: 304:metamorphosis 301: 298: 295: 291: 287: 283: 279: 275: 271: 266: 264: 259: 254: 250: 246: 242: 235: 233: 230: 223: 221: 219: 215: 211: 207: 203: 199: 192: 190: 188: 184: 183: 173:Fisher, 1946 170: 168: 162: 159: 158:Binomial name 155: 151: 150: 145: 142: 141: 138: 137: 133: 130: 129: 126: 123: 120: 119: 116: 113: 110: 109: 106: 103: 100: 99: 96: 93: 90: 87: 86: 83: 80: 77: 74: 73: 70: 67: 64: 63: 60: 57: 54: 53: 50: 47: 44: 43: 38: 33: 29: 26: 22: 19: 599: 567: 561: 557: 529: 523: 514: 510: 501: 456: 449: 438:. Retrieved 430: 424: 416: 377: 375: 368: 362: 352: 341: 332: 323: 314: 269: 267: 240: 239: 228: 227: 224:Distribution 212:nearby. The 197: 196: 181: 180: 179: 166: 164: 148: 147: 136:Listriolobus 135: 88: 75: 24: 18: 702:SeaLifeBase 570:(1): 1–17. 386:Los Angeles 297:trochophore 292:hatch into 253:peristaltic 218:respiration 193:Description 115:Echiuroidea 737:Categories 440:2012-11-11 408:References 404:remained. 382:wastewater 317:commensals 294:planktonic 187:spoon worm 125:Echiuridae 101:Subclass: 95:Sedentaria 743:Echiurans 402:diversity 359:flounders 348:predators 286:nephridia 280:into the 263:ecosystem 245:substrate 202:proboscis 143:Species: 55:Kingdom: 49:Eukaryota 642:10003784 615:Q2339954 609:Wikidata 395:polluted 337:and the 258:bacteria 121:Family: 69:Annelida 65:Phylum: 59:Animalia 45:Domain: 694:1029498 629:5234373 399:benthic 311:Ecology 278:gametes 236:Biology 131:Genus: 111:Order: 105:Echiura 720:266792 681:266792 655:155070 536:  468:  355:preyed 328:, the 300:larvae 282:coelom 274:gonads 272:. The 715:WoRMS 707:60643 668:61850 637:IRMNG 507:(PDF) 431:WoRMS 391:fauna 249:cilia 206:setae 89:Clade 76:Clade 676:OBIS 663:NCBI 650:ITIS 624:GBIF 534:ISBN 466:ISBN 321:crab 290:eggs 214:anus 572:doi 462:263 373:). 739:: 717:: 704:: 691:: 678:: 665:: 652:: 639:: 626:: 611:: 568:22 566:. 548:^ 515:34 513:. 509:. 480:^ 464:. 433:. 429:. 265:. 91:: 78:: 578:. 574:: 542:. 474:. 443:. 423:"

Index

Scientific classification
Edit this classification
Eukaryota
Animalia
Annelida
Pleistoannelida
Sedentaria
Echiura
Echiuroidea
Echiuridae
Listriolobus
Binomial name
spoon worm
proboscis
setae
nephridiopores
anus
respiration
substrate
cilia
peristaltic
bacteria
ecosystem
gonads
gametes
coelom
nephridia
eggs
planktonic
trochophore

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