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Lolium rigidum

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254: 33: 376:. This nematode causes galls on the grass flower spikes and it is in these that the bacteria multiply. Infected galls are present in the winter, but become more toxic as the inflorescence dies in spring. The disease occurs when the pasture is grazed at this toxic stage. Topping the pasture before grazing may prevent the condition, but the seed heads are still toxic when preserved as hay. The symptoms of poisoning are neurological, often causing a mortality rate of forty to fifty percent in infected animals. 46: 814: 422:
has been grown as a forage crop in southern Australia since the nineteenth century and seeds readily, it grows ubiquitously in the region with large, genetically diverse populations. Now that arable crops are increasingly been grown, it has become an important weed of these crops. The main reason for
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is a native of the Mediterranean area. Its natural range includes Western, Southern and Central Europe, northern Africa, the Near East, western Asia and the Indian sub-continent. It has spread to many other parts of the world and is considered invasive in some regions. It is planted in Australia as a
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is an annual grass that grows in open tussocks. It has fibrous roots and can grow up to a metre tall. The plant form is usually erect but may be prostrate. The stems are often geniculate (with a knee-like bend) and are purplish at the base. The leaves are 5 to 25 cm (2 to 10 in) long, and 3
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is grown as a forage crop in suitable areas. About 80% of the seed germinates in the autumn, soon after the first significant rains, and about 5% may remain dormant for twelve months. The plant has vigorous growth, and flowering is initiated when the day length is at least eight hours. This means
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its success in competing with crops is its ability to produce prolific numbers of seeds per plant – up to 45,000 seeds per 1 m (11 sq ft) – and also its ability to compete with crops at a very early stage of their life cycle.
238:. It is a native of southern Europe, northern Africa, the Middle East and the Indian subcontinent and is grown as a forage crop, particularly in Australia, where it is also a serious and economically damaging crop weed. 301:
forage crop but not usually in Europe. It was introduced as a forage crop in Australia around 1880, but has since proven to be an economically damaging weed that infests other more important crops, such as
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The young leaves are rolled when in bud, the auricles are small and the ligule is white and translucent, wider than it is long. The unbranched flower spike is up to 30 cm (12 in) long, with the
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is a diploid grass with a chromosome number of n=7 (2n=14). It exhibits much genetic variability and grows readily in a variety of situations and habitats. It can hybridise with both
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to 5 mm (0.12 to 0.20 in) wide; the upper surface is glossy dark green, flat and hairless with longitudinal veins, and the underside is shiny and smooth.
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has become one of the most serious and economically damaging weeds in southern Australia. A 2021 study listed annual ryegrass as fourth on the list of
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or annual ryegrass staggers, which is known to occur in the west and south of Australia and in South Africa. The infection is caused by the bacterium
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which cost Australian farmers the most over a 60-year period, at over a billion Australian dollars.
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may become infected by a certain species of bacteria, which results in the production of
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which are toxic to livestock; ingestion of infected material causes a disease, known as
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is a similar shape with two nerve and a few fine hairs. The three anthers are yellow.
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that the crop has a relatively uniform flowering period in late winter/early spring.
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The spikelets bear up to twelve florets, mostly with a single
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In addition, it serves as a host for species of the bacteria
564:. Springer Science & Business Media. pp. 213–216. 342:. It propagates solely by seed; freshly shed seed displays 558:
Boller, Beat; Posselt, Ulrich K.; Veronesi, Fabio (2010).
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Department of Primary Industries and Regional Development
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Murrumbidgee Catchment Management Authority (2008).
839: 398:) which can cause serious losses in cereal crops. 758:Wegulo, Stephen N; Carlson, Michael P. (2011). 583: 581: 434:(ARGT), and can be infected by the poisonous 8: 699:in Australia: a very resistance prone weed!" 612: 610: 449:have met with an unexpectedly high level of 367:, which is introduced into the grass by the 266:on alternating sides and edgeways-on to the 210:. Common names by which it is known include 270:(stem), pressed into recesses in the stem. 827: 31: 20: 767:University of Nebraska–Lincoln Extension 637: 635: 485: 523: 521: 519: 285:is oblong and has five nerves and the 689: 687: 7: 1397:9c039cf8-8ef4-45de-9570-94be7262c1ea 1260:e893ec60-02c5-44a9-936f-bf23b711a365 723: 721: 719: 790:Australian Broadcasting Corporation 14: 1348:urn:lsid:ipni.org:names:407503-1 812: 642:Simpson, Wayne (1 August 2013). 590:Annual ryegrass (Lolium rigidum) 561:Fodder Crops and Amenity Grasses 44: 382:is susceptible to infection by 461:are being explored, including 336:), as well as some species of 1: 442:in humans and other mammals. 445:Attempts to control it with 537:Invasive Species Compendium 1464: 670:"Annual ryegrass toxicity" 644:"Annual ryegrass staggers" 780:Khan, Jo (31 July 2021). 178: 171: 41:Scientific classification 39: 30: 23: 504:. Biosecurity Queensland 432:annual ryegrass toxicity 359:annual ryegrass toxicity 293:Distribution and habitat 646:. Merck. Archived from 464:Pyrenophora seminiperda 396:Gaeumannomyces graminis 214:, a name also given to 258: 735:. Western Australia. 364:Rathayibacter toxicus 256: 823:at Wikimedia Commons 733:Agriculture and Food 451:herbicide resistance 349:The flower spike of 650:on 11 December 2015 502:Weeds of Australia 494:"Annual ryegrass: 388:Claviceps purpurea 259: 257:Annotated spikelet 220:Lolium multiflorum 1430: 1429: 1294:Open Tree of Life 833:Taxon identifiers 817:Media related to 739:. 7 December 2020 729:"Annual ryegrass" 618:"Annual ryegrass" 571:978-1-4419-0760-8 533:(rigid ryegrass)" 195: 194: 1455: 1448:Flora of Lebanon 1423: 1422: 1410: 1409: 1400: 1399: 1387: 1386: 1377: 1376: 1364: 1363: 1351: 1350: 1338: 1337: 1325: 1324: 1312: 1311: 1302: 1301: 1289: 1288: 1276: 1275: 1263: 1262: 1253: 1252: 1240: 1239: 1227: 1226: 1214: 1213: 1211:NBNSYS0000004868 1201: 1200: 1188: 1187: 1175: 1174: 1162: 1161: 1149: 1148: 1136: 1135: 1123: 1122: 1110: 1109: 1097: 1096: 1084: 1083: 1074: 1073: 1061: 1060: 1048: 1047: 1035: 1034: 1022: 1021: 1009: 1008: 996: 995: 983: 982: 970: 969: 957: 956: 944: 943: 934: 933: 921: 920: 911: 910: 898: 897: 885: 884: 875: 874: 873: 860: 859: 858: 828: 816: 801: 800: 798: 796: 777: 771: 770: 764: 755: 749: 748: 746: 744: 725: 714: 713: 711: 709: 703:Plants in Action 691: 682: 681: 679: 677: 666: 660: 659: 657: 655: 639: 630: 629: 627: 625: 614: 605: 604: 602: 600: 594: 585: 576: 575: 555: 549: 548: 546: 544: 525: 514: 513: 511: 509: 490: 475:invasive species 414:Damaging effects 330:Italian ryegrass 236:Wimmera ryegrass 216:Italian ryegrass 184: 49: 48: 35: 21: 16:Species of grass 1463: 1462: 1458: 1457: 1456: 1454: 1453: 1452: 1433: 1432: 1431: 1426: 1418: 1413: 1405: 1403: 1395: 1390: 1382: 1380: 1372: 1367: 1359: 1354: 1346: 1341: 1333: 1328: 1320: 1315: 1307: 1305: 1297: 1292: 1284: 1281:Observation.org 1279: 1271: 1266: 1258: 1256: 1248: 1243: 1235: 1230: 1222: 1217: 1209: 1204: 1196: 1191: 1183: 1178: 1170: 1165: 1157: 1152: 1144: 1139: 1131: 1126: 1118: 1113: 1105: 1100: 1092: 1087: 1079: 1077: 1069: 1064: 1056: 1051: 1043: 1038: 1030: 1025: 1017: 1012: 1004: 999: 991: 986: 978: 973: 965: 960: 952: 947: 939: 937: 929: 924: 916: 914: 906: 901: 893: 888: 880: 878: 869: 868: 863: 854: 853: 848: 835: 809: 804: 794: 792: 779: 778: 774: 762: 757: 756: 752: 742: 740: 727: 726: 717: 707: 705: 693: 692: 685: 675: 673: 668: 667: 663: 653: 651: 641: 640: 633: 623: 621: 616: 615: 608: 598: 596: 592: 587: 586: 579: 572: 557: 556: 552: 542: 540: 527: 526: 517: 507: 505: 492: 491: 487: 483: 438:, which causes 416: 404: 390:), and also by 373:Anguina funesta 315: 295: 244: 212:annual ryegrass 191: 186: 180: 167: 164:L. rigidum 43: 17: 12: 11: 5: 1461: 1459: 1451: 1450: 1445: 1435: 1434: 1428: 1427: 1425: 1424: 1420:wfo-0000878467 1411: 1401: 1388: 1378: 1365: 1352: 1339: 1326: 1313: 1303: 1290: 1277: 1264: 1254: 1241: 1237:Lolium~rigidum 1228: 1215: 1202: 1189: 1176: 1163: 1150: 1137: 1124: 1111: 1098: 1085: 1075: 1062: 1058:Lolium rigidum 1049: 1036: 1023: 1010: 997: 984: 980:Lolium_rigidum 971: 958: 945: 935: 922: 912: 908:lolium-rigidum 899: 886: 876: 871:Lolium rigidum 861: 845: 843: 841:Lolium rigidum 837: 836: 831: 825: 824: 820:Lolium rigidum 808: 807:External links 805: 803: 802: 772: 750: 715: 697:Lolium rigidum 683: 672:. SheepConnect 661: 631: 606: 577: 570: 550: 531:Lolium rigidum 515: 496:Lolium rigidum 484: 482: 479: 430:, which cause 415: 412: 407:Lolium rigidum 403: 400: 334:L. multiflorum 318:Lolium rigidum 314: 311: 298:Lolium rigidum 294: 291: 247:Lolium rigidum 243: 240: 232:Swiss ryegrass 224:rigid ryegrass 199:Lolium rigidum 193: 192: 187: 182:Lolium rigidum 176: 175: 169: 168: 161: 159: 155: 154: 147: 143: 142: 137: 133: 132: 127: 123: 122: 117: 113: 112: 107: 100: 99: 94: 87: 86: 81: 74: 73: 68: 61: 60: 55: 51: 50: 37: 36: 28: 27: 25:Lolium rigidum 15: 13: 10: 9: 6: 4: 3: 2: 1460: 1449: 1446: 1444: 1441: 1440: 1438: 1421: 1416: 1412: 1408: 1402: 1398: 1393: 1389: 1385: 1379: 1375: 1370: 1366: 1362: 1357: 1353: 1349: 1344: 1340: 1336: 1331: 1327: 1323: 1318: 1314: 1310: 1304: 1300: 1295: 1291: 1287: 1282: 1278: 1274: 1269: 1265: 1261: 1255: 1251: 1246: 1242: 1238: 1233: 1229: 1225: 1220: 1216: 1212: 1207: 1203: 1199: 1194: 1190: 1186: 1181: 1177: 1173: 1168: 1164: 1160: 1155: 1151: 1147: 1142: 1138: 1134: 1129: 1125: 1121: 1116: 1112: 1108: 1103: 1099: 1095: 1090: 1086: 1082: 1076: 1072: 1067: 1063: 1059: 1054: 1050: 1046: 1041: 1037: 1033: 1028: 1024: 1020: 1015: 1011: 1007: 1002: 998: 994: 989: 985: 981: 976: 972: 968: 963: 959: 955: 950: 946: 942: 936: 932: 927: 923: 919: 913: 909: 904: 900: 896: 891: 887: 883: 877: 872: 866: 862: 857: 851: 847: 846: 844: 842: 838: 834: 829: 822: 821: 815: 811: 810: 806: 791: 787: 783: 776: 773: 768: 761: 754: 751: 738: 734: 730: 724: 722: 720: 716: 704: 700: 698: 690: 688: 684: 671: 665: 662: 649: 645: 638: 636: 632: 619: 613: 611: 607: 591: 584: 582: 578: 573: 567: 563: 562: 554: 551: 538: 534: 532: 524: 522: 520: 516: 503: 499: 497: 489: 486: 480: 478: 476: 472: 468: 466: 465: 460: 459:bioherbicides 456: 452: 448: 443: 441: 437: 433: 429: 424: 421: 413: 411: 408: 401: 399: 397: 393: 389: 385: 381: 377: 375: 374: 370: 366: 365: 360: 356: 352: 347: 345: 341: 340: 335: 331: 327: 323: 319: 312: 310: 308: 304: 299: 292: 290: 288: 284: 280: 276: 271: 269: 265: 255: 251: 248: 241: 239: 237: 233: 229: 225: 221: 217: 213: 209: 205: 201: 200: 190: 185: 183: 177: 174: 173:Binomial name 170: 166: 165: 160: 157: 156: 153: 152: 148: 145: 144: 141: 138: 135: 134: 131: 128: 125: 124: 121: 118: 115: 114: 111: 108: 105: 102: 101: 98: 95: 92: 89: 88: 85: 82: 79: 76: 75: 72: 71:Tracheophytes 69: 66: 63: 62: 59: 56: 53: 52: 47: 42: 38: 34: 29: 26: 22: 19: 840: 819: 793:. Retrieved 785: 775: 766: 753: 741:. Retrieved 732: 706:. Retrieved 702: 696: 674:. Retrieved 664: 652:. Retrieved 648:the original 622:. Retrieved 620:. HerbiGuide 597:. Retrieved 560: 553: 541:. Retrieved 536: 530: 506:. Retrieved 501: 495: 488: 470: 469: 467:, a fungus. 462: 454: 444: 436:ergot fungus 425: 419: 417: 406: 405: 395: 387: 379: 378: 371: 362: 355:corynetoxins 350: 348: 337: 333: 325: 317: 316: 297: 296: 272: 260: 246: 245: 235: 231: 228:stiff darnel 227: 223: 219: 211: 198: 197: 196: 181: 179: 163: 162: 150: 103: 90: 77: 64: 24: 18: 1193:NatureServe 1128:iNaturalist 865:Wikispecies 676:11 December 428:Clavibacter 242:Description 206:of annual 136:Subfamily: 110:Commelinids 84:Angiosperms 1437:Categories 1322:kew-422874 1317:Plant List 938:Calflora: 708:8 December 654:9 December 624:6 December 543:6 December 508:6 December 481:References 471:L. rigidum 455:L. rigidum 420:L. rigidum 380:L. rigidum 351:L. rigidum 326:L. perenne 1102:GrassBase 1071:242416784 1045:242416784 1027:FloraBase 447:herbicide 322:perennial 264:spikelets 158:Species: 54:Kingdom: 1443:Pooideae 1392:VicFlora 1381:VASCAN: 1374:25511246 1369:Tropicos 1232:NSWFlora 1198:2.133208 1159:10203772 1146:407503-1 1107:imp06040 975:eFloraSA 915:BioLib: 856:Q3156354 850:Wikidata 786:ABC News 440:ergotism 418:Because 394:fungus ( 392:take-all 369:nematode 344:dormancy 140:Pooideae 126:Family: 97:Monocots 1245:NTFlora 1094:2706218 993:1114468 962:Ecocrop 795:30 July 743:31 July 599:31 July 339:Festuca 313:Ecology 204:species 146:Genus: 130:Poaceae 116:Order: 58:Plantae 1361:1272-5 1330:PLANTS 1299:459976 1286:141435 1257:NZOR: 1133:164711 1081:LOLRIG 1078:FoIO: 1019:192473 931:434563 903:ARKive 895:111218 879:APDB: 568:  539:. CABI 328:) and 303:canola 281:, the 268:rachis 151:Lolium 120:Poales 1404:WoI: 1356:SANBI 1306:PFI: 1268:NZPCN 1224:89674 1185:40897 1172:31167 1154:IRMNG 1120:22498 1053:FoAO2 1014:EUNIS 1006:LOLRI 954:3VVXT 918:42404 882:52120 763:(PDF) 593:(PDF) 384:ergot 307:wheat 287:palea 283:lemma 275:glume 208:grass 202:is a 189:Gaud. 104:Clade 91:Clade 78:Clade 65:Clade 1407:1244 1384:7760 1343:POWO 1335:LORI 1309:7385 1273:3442 1250:4615 1219:NCBI 1180:ITIS 1141:IPNI 1115:GRIN 1089:GBIF 1001:EPPO 967:1363 941:4947 926:BOLD 890:APNI 797:2021 745:2021 710:2015 678:2015 656:2015 626:2015 601:2021 566:ISBN 545:2015 510:2015 402:Uses 305:and 234:and 1415:WFO 1206:NBN 1167:ISC 1066:FoC 1040:FNA 1032:478 988:EoL 949:CoL 279:awn 222:), 1439:: 1417:: 1394:: 1371:: 1358:: 1345:: 1332:: 1319:: 1296:: 1283:: 1270:: 1247:: 1234:: 1221:: 1208:: 1195:: 1182:: 1169:: 1156:: 1143:: 1130:: 1117:: 1104:: 1091:: 1068:: 1055:: 1042:: 1029:: 1016:: 1003:: 990:: 977:: 964:: 951:: 928:: 905:: 892:: 867:: 852:: 788:. 784:. 765:. 731:. 718:^ 701:. 686:^ 634:^ 609:^ 580:^ 535:. 518:^ 500:. 309:. 230:, 226:, 106:: 93:: 80:: 67:: 799:. 769:. 747:. 712:. 695:" 680:. 658:. 628:. 603:. 574:. 547:. 529:" 512:. 498:" 386:( 332:( 324:( 218:(

Index


Scientific classification
Edit this classification
Plantae
Tracheophytes
Angiosperms
Monocots
Commelinids
Poales
Poaceae
Pooideae
Lolium
Binomial name
Gaud.
species
grass
Italian ryegrass

spikelets
rachis
glume
awn
lemma
palea
canola
wheat
perennial
Italian ryegrass
Festuca
dormancy

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