321:
60:
403:
395:
40:
421:
for an adult of this genus is 25 cm (10 in), but most species reach 7–15 cm (2.8–5.9 in) and some species do not surpass 5 cm (2 in). In appearance they may resemble miniature
320:
1374:
446:
matter. Breeding is by egg scattering and takes place close to the bottom, near or within areas of dense plant growth. They do not show parental care and adults may eat the young.
1777:
1659:
1685:
1372:
The fishes of the inland waters of southeast Asia: A catalogue and core bibliography of the fishes known to occur in freshwaters, mangroves and estuaries.
1371:
1633:
1672:
1184:
1262:
1724:
1767:
1677:
807:
794:
1772:
1607:
1594:
836:
1762:
1412:
888:
1690:
1222:
1169:
1095:
982:
690:
148:
59:
964:
952:
1432:"Early Cenozoic Cyprinoids (Ostariophysi: Cypriniformes: Cyprinidae and Danionidae) from Sumatra, Indonesia"
740:
369:
215:
956:
819:
1534:
1045:
947:
880:
832:
465:
969:
884:
1757:
1711:
1612:
1276:
1145:
1104:
1023:
1009:
811:
786:
772:
720:
1344:
1280:
1109:
1729:
1581:
1431:
1249:
1137:
1067:
1018:
924:
916:
902:
875:
840:
815:
790:
669:
633:
574:
538:
309:
1231:
1206:
1192:
1178:
1155:
1075:
1050:
977:
709:
612:
1469:
1151:
1004:
937:
749:
699:
655:
481:
54:
1716:
1123:
995:
853:
754:
725:
304:
515:
with regard to the
Cyprininae and better merged there at least for the largest part (including
1698:
1664:
1620:
1451:
1244:
1236:
1198:
867:
627:
589:
568:
469:
402:
1054:
1752:
1703:
1443:
1289:
896:
781:
734:
597:
554:
418:
383:
345:
204:
1378:
1303:
1271:
1257:
1118:
990:
861:
768:
762:
675:
374:
200:
165:
290:
Previously, it was assumed that the earliest record of this genus was the fossil species
1473:(Teleostei: Cyprinidae), a new species of barb from Manimala River, Kerala, South India.
1393:
1131:
1089:
1031:
911:
848:
802:
647:
641:
618:
394:
101:
1746:
1572:
1340:
1216:
1062:
827:
583:
501:
111:
1514:
1487:
526:
512:
439:
247:
157:
1625:
1447:
1646:
1586:
1566:
1309:
1294:
1081:
1036:
929:
661:
603:
560:
497:
406:
325:
1599:
684:
521:
435:
361:
211:
207:
121:
1557:
1455:
1163:
505:
486:
365:
277:
271:
253:
241:
71:
20:
1416:, a new genus of barbs (Teleostei: Cyprinidae) from Western Ghats of India.
39:
1551:
1468:
Krishna Kumar, K., Benno
Pereira, F.G. & Radhakrishnan, K.V. (2012):
1353:
508:
283:
259:
235:
131:
91:
1638:
492:
299:
226:
1651:
1504:
Egypt. Acad. J. Biolog. Sci. (B. Zoology) Vol. 12(2) pp: 93-102(2020)
460:
379:
295:
265:
219:
81:
1528:
1491:, a new fish species (Cypriniformes; Cyprinidae) from Kerala, India.
1518:(Cypriniformes, Cyprinidae), a new fish species from Kerala, India.
484:. The closest living relatives of the spotted barbs are the genus
443:
401:
393:
319:
196:
534:
423:
302:, but a 2020 study reclassified this species into the new genus
1532:
1411:
Raghavan, R., Philip, S., Ali, A. & Dahanukar, N. (2013):
1392:
Pethiyagoda, R., Meegaskumbura, M. & Maduwage, K. (2012):
549:
These are the currently recognized species in this genus:
1401:
Ichthyological
Exploration of Freshwaters, 23 (1): 69–95.
468:
remains the same in these – except in cases of
529:
and its relatives – for some time included in
1541:
1382:Raffles Bulletin of Zoology, Suppl. No. 27: 1–663.
458:have been classified in several genera, including
426:and are sometimes brightly coloured or patterned.
1521:Journal of Research in Biology, 3 (7): 1093–1104.
1494:Journal of Research in Biology, 4 (8): 1581–1588.
1394:A synopsis of the South Asian fishes referred to
1419:Journal of Threatened Taxa, 5 (15): 4932–4938.
1366:
1364:
8:
1513:Plamoottil, M. & Abraham, N.P. (2014):
1529:
38:
27:
843:, S. Phurailatpam & M. S. Bisht, 2004
233:have been moved to other genera such as
1778:Taxa named by Francis Buchanan-Hamilton
1323:
972:, K. P. Verma & R. B. Sharma, 1977
714:G. M. Yazdani & M. Babu Rao, 1976
704:G. M. Yazdani & D. F. Singh, 1994
464:. Despite the reclassifications, the
7:
1335:
1333:
1331:
1329:
1327:
1436:Journal of Vertebrate Paleontology
14:
942:Arunkumar & Tombi Singh, 1998
504:were formerly often separated as
1430:Murray, Alison M. (2020-01-02).
533:– appears to be a kind of
496:. These and the other "typical"
58:
413:), male below and female above.
364:(west to Pakistan and south to
454:Historically, many species of
1:
1476:Biosystematica, 5 (2): 31-37.
1448:10.1080/02724634.2020.1762627
398:Puntius of West Bengal, India
324:Puti maach (Puntius fish) in
438:; their diet includes small
308:, possibly in the subfamily
1014:(Malabar black-backed barb)
519:). In particular the genus
511:, but this group is highly
1794:
348:term for small cyprinids.
18:
372:, with a single species,
181:
176:
163:
156:
55:Scientific classification
53:
46:
37:
30:
1343:; Pauly, Daniel (eds.).
1211:Nguyen & Doan, 1969)
1000:(Travancore yellow barb)
965:Puntius muzaffarpurensis
19:Not to be confused with
1486:Plamoottil, M. (2014):
370:Mainland Southeast Asia
216:Mainland Southeast Asia
1768:Fish of Southeast Asia
1046:Puntius paucimaculatus
948:Puntius mudumalaiensis
490:and perhaps the genus
414:
399:
329:
1712:Paleobiology Database
1398:(Pisces: Cyprinidae).
1370:Kottelat, M. (2013):
1105:Puntius sahyadriensis
730:(Longfin Kerala barb)
721:Puntius dolichopterus
579:(Scarlet-banded barb)
405:
397:
323:
1471:Puntius madhusoodani
1357:. June 2016 version.
1285:(Spotfin green barb)
925:Puntius melanostigma
876:Puntius madhusoodani
670:Puntius cauveriensis
539:evolved convergently
356:Fishes of the genus
1232:Puntius tetraspilus
1207:Puntius takhoaensis
1193:Puntius spilopterus
1179:Puntius sophoroides
1076:Puntius punjabensis
1026:& Vineeth, 2020
978:Puntius nangalensis
745:(Long-snouted barb)
710:Puntius deccanensis
613:Puntius bimaculatus
229:formerly placed in
1773:Fish of South Asia
1489:Puntius nigronotus
1377:2015-01-06 at the
1345:"Species in genus
1299:(Greenstripe barb)
1146:Puntius somphongsi
1005:Puntius nigronotus
938:Puntius morehensis
750:Puntius euspilurus
700:Puntius crescentus
656:Puntius burmanicus
482:grammatical gender
415:
400:
330:
1763:Cyprinidae genera
1740:
1739:
1699:Open Tree of Life
1535:Taxon identifiers
1313:
1298:
1284:
1266:
1253:
1245:Puntius thermalis
1240:
1226:
1212:
1202:
1188:
1173:
1159:
1141:
1127:
1113:
1099:
1085:
1071:
1058:
1040:
1027:
1013:
999:
986:
973:
960:
943:
933:
920:
906:
892:
871:
857:
844:
823:
798:
776:
758:
744:
729:
715:
705:
694:
679:
665:
651:
637:
628:Puntius bramoides
622:
607:
593:
578:
569:Puntius amphibius
564:
525:, containing the
434:These fishes are
188:
187:
171:F. Hamilton, 1822
152:
1785:
1733:
1732:
1720:
1719:
1707:
1706:
1694:
1693:
1681:
1680:
1668:
1667:
1655:
1654:
1642:
1641:
1629:
1628:
1616:
1615:
1603:
1602:
1590:
1589:
1577:
1576:
1575:
1562:
1561:
1560:
1530:
1523:
1511:
1505:
1502:
1496:
1484:
1478:
1466:
1460:
1459:
1427:
1421:
1409:
1403:
1390:
1384:
1368:
1359:
1358:
1337:
1307:
1293:
1290:Puntius vittatus
1275:
1261:
1248:
1235:
1220:
1210:
1196:
1182:
1167:
1149:
1135:
1122:
1108:
1093:
1079:
1066:
1049:
1035:
1022:
1008:
994:
981:
968:
951:
941:
928:
915:
900:
897:Puntius mahecola
879:
865:
852:
839:, S. P. Uniyal,
831:
806:
785:
782:Puntius kamalika
766:
753:
738:
735:Puntius dorsalis
724:
713:
703:
688:
673:
659:
645:
631:
616:
601:
598:Puntius arenatus
587:
572:
558:
555:Puntius ambassis
466:specific epithet
384:species richness
166:Cyprinus sophore
147:
63:
62:
42:
28:
1793:
1792:
1788:
1787:
1786:
1784:
1783:
1782:
1743:
1742:
1741:
1736:
1728:
1723:
1715:
1710:
1702:
1697:
1689:
1684:
1676:
1671:
1663:
1658:
1650:
1645:
1637:
1632:
1624:
1619:
1611:
1606:
1598:
1593:
1585:
1580:
1571:
1570:
1565:
1556:
1555:
1550:
1537:
1527:
1526:
1516:Puntius viridis
1512:
1508:
1503:
1499:
1485:
1481:
1467:
1463:
1442:(1): e1762627.
1429:
1428:
1424:
1410:
1406:
1391:
1387:
1379:Wayback Machine
1369:
1362:
1339:
1338:
1325:
1320:
1304:Puntius waageni
1272:Puntius viridis
1258:Puntius titteya
1164:Puntius sophore
1119:Puntius sanctus
1068:S. O. Kullander
1019:Puntius ocellus
991:Puntius nelsoni
907:(Mahecola barb)
862:Puntius layardi
763:Puntius fraseri
608:(Arenatus barb)
547:
452:
432:
411:Puntius titteya
392:
382:. The greatest
354:
318:
310:Smiliogastrinae
201:freshwater fish
172:
169:
146:
57:
24:
17:
16:Genus of fishes
12:
11:
5:
1791:
1789:
1781:
1780:
1775:
1770:
1765:
1760:
1755:
1745:
1744:
1738:
1737:
1735:
1734:
1721:
1708:
1695:
1682:
1669:
1656:
1643:
1630:
1617:
1604:
1591:
1578:
1563:
1547:
1545:
1539:
1538:
1533:
1525:
1524:
1506:
1497:
1479:
1461:
1422:
1404:
1385:
1360:
1341:Froese, Rainer
1322:
1321:
1319:
1316:
1315:
1314:
1300:
1286:
1268:
1254:
1241:
1228:
1227:(Onespot barb)
1213:
1203:
1189:
1175:
1160:
1142:
1132:Puntius sealei
1128:
1115:
1101:
1090:Puntius puntio
1086:
1072:
1059:
1042:
1032:Puntius parrah
1028:
1015:
1001:
987:
974:
961:
944:
934:
921:
912:Puntius masyai
908:
893:
872:
858:
849:Puntius kyphus
845:
824:
812:N. K. A. Silva
803:Puntius kelumi
799:
787:N. K. A. Silva
778:
759:
746:
731:
717:
706:
696:
681:
680:(Cauvery barb)
666:
652:
642:Puntius brevis
638:
624:
623:(Redside barb)
609:
594:
580:
565:
546:
543:
480:have the same
451:
448:
431:
428:
391:
388:
353:
350:
317:
314:
292:Puntius bussyi
186:
185:
179:
178:
174:
173:
170:
161:
160:
154:
153:
139:
135:
134:
129:
125:
124:
119:
115:
114:
109:
105:
104:
102:Actinopterygii
99:
95:
94:
89:
85:
84:
79:
75:
74:
69:
65:
64:
51:
50:
44:
43:
35:
34:
15:
13:
10:
9:
6:
4:
3:
2:
1790:
1779:
1776:
1774:
1771:
1769:
1766:
1764:
1761:
1759:
1756:
1754:
1751:
1750:
1748:
1731:
1726:
1722:
1718:
1713:
1709:
1705:
1700:
1696:
1692:
1687:
1683:
1679:
1674:
1670:
1666:
1661:
1657:
1653:
1648:
1644:
1640:
1635:
1631:
1627:
1622:
1618:
1614:
1609:
1605:
1601:
1596:
1592:
1588:
1583:
1579:
1574:
1568:
1564:
1559:
1553:
1549:
1548:
1546:
1544:
1540:
1536:
1531:
1522:
1519:
1517:
1510:
1507:
1501:
1498:
1495:
1492:
1490:
1483:
1480:
1477:
1474:
1472:
1465:
1462:
1457:
1453:
1449:
1445:
1441:
1437:
1433:
1426:
1423:
1420:
1417:
1415:
1408:
1405:
1402:
1399:
1397:
1389:
1386:
1383:
1380:
1376:
1373:
1367:
1365:
1361:
1356:
1355:
1350:
1348:
1342:
1336:
1334:
1332:
1330:
1328:
1324:
1317:
1311:
1306:
1305:
1301:
1296:
1292:
1291:
1287:
1282:
1278:
1274:
1273:
1269:
1267:(Cherry barb)
1264:
1260:
1259:
1255:
1251:
1247:
1246:
1242:
1238:
1234:
1233:
1229:
1224:
1219:
1218:
1217:Puntius terio
1214:
1209:
1208:
1204:
1200:
1195:
1194:
1190:
1186:
1181:
1180:
1176:
1171:
1166:
1165:
1161:
1157:
1153:
1148:
1147:
1143:
1139:
1134:
1133:
1129:
1125:
1121:
1120:
1116:
1114:(Khavli barb)
1111:
1107:
1106:
1102:
1100:(Puntio barb)
1097:
1092:
1091:
1087:
1083:
1078:
1077:
1073:
1069:
1065:
1064:
1063:Puntius pugio
1060:
1056:
1052:
1048:
1047:
1043:
1041:(Parrah barb)
1038:
1034:
1033:
1029:
1025:
1021:
1020:
1016:
1011:
1007:
1006:
1002:
997:
993:
992:
988:
984:
980:
979:
975:
971:
967:
966:
962:
958:
954:
950:
949:
945:
940:
939:
935:
931:
927:
926:
922:
918:
914:
913:
909:
904:
899:
898:
894:
890:
889:Radhakrishnan
886:
885:Benno Pereira
882:
881:Krishna Kumar
878:
877:
873:
869:
864:
863:
859:
855:
851:
850:
846:
842:
838:
834:
830:
829:
828:Puntius khohi
825:
821:
820:Meegaskumbura
817:
813:
809:
805:
804:
800:
796:
792:
788:
784:
783:
779:
777:(Dharna barb)
774:
770:
765:
764:
760:
756:
752:
751:
747:
742:
737:
736:
732:
727:
723:
722:
718:
716:(Deccan barb)
712:
711:
707:
702:
701:
697:
692:
687:
686:
685:Puntius chola
682:
677:
672:
671:
667:
663:
658:
657:
653:
649:
644:
643:
639:
635:
630:
629:
625:
620:
615:
614:
610:
605:
600:
599:
595:
591:
586:
585:
584:Puntius aphya
581:
576:
571:
570:
566:
562:
557:
556:
552:
551:
550:
544:
542:
540:
536:
532:
528:
524:
523:
518:
514:
510:
507:
503:
499:
495:
494:
489:
488:
483:
479:
475:
471:
467:
463:
462:
457:
449:
447:
445:
441:
440:invertebrates
437:
429:
427:
425:
420:
412:
408:
404:
396:
389:
387:
386:is in India.
385:
381:
377:
376:
371:
367:
363:
360:are found in
359:
351:
349:
347:
343:
339:
335:
327:
322:
315:
313:
311:
307:
306:
301:
297:
293:
288:
286:
285:
280:
279:
274:
273:
268:
267:
262:
261:
256:
255:
250:
249:
244:
243:
238:
237:
232:
228:
223:
221:
218:, as well as
217:
213:
209:
206:
202:
198:
194:
193:
184:
180:
175:
168:
167:
162:
159:
155:
150:
145:
144:
140:
137:
136:
133:
130:
127:
126:
123:
120:
117:
116:
113:
112:Cypriniformes
110:
107:
106:
103:
100:
97:
96:
93:
90:
87:
86:
83:
80:
77:
76:
73:
70:
67:
66:
61:
56:
52:
49:
48:Puntius chola
45:
41:
36:
33:
29:
26:
22:
1758:Barbs (fish)
1542:
1520:
1515:
1509:
1500:
1493:
1488:
1482:
1475:
1470:
1464:
1439:
1435:
1425:
1418:
1413:
1407:
1400:
1395:
1388:
1381:
1352:
1346:
1302:
1288:
1270:
1263:Deraniyagala
1256:
1250:Valenciennes
1243:
1230:
1215:
1205:
1191:
1177:
1162:
1144:
1130:
1117:
1103:
1088:
1074:
1061:
1044:
1030:
1017:
1003:
989:
976:
963:
946:
936:
923:
910:
903:Valenciennes
895:
874:
860:
847:
826:
801:
780:
761:
748:
733:
719:
708:
698:
695:(Swamp barb)
683:
668:
654:
640:
634:Valenciennes
626:
611:
596:
582:
575:Valenciennes
567:
553:
548:
541:with barbs.
530:
527:tinfoil barb
520:
516:
513:paraphyletic
491:
485:
477:
473:
459:
455:
453:
433:
419:maximum size
416:
410:
373:
357:
355:
341:
340:(Pronounced
337:
333:
331:
303:
291:
289:
282:
276:
270:
264:
258:
252:
248:Desmopuntius
246:
240:
234:
230:
224:
191:
190:
189:
182:
164:
158:Type species
142:
141:
47:
31:
25:
1647:iNaturalist
1567:Wikispecies
1223:F. Hamilton
1174:(Pool barb)
1170:F. Hamilton
1096:F. Hamilton
917:H. M. Smith
841:H. K. Joshi
808:Pethiyagoda
795:Pethiyagoda
691:F. Hamilton
472:– as
407:Cherry barb
390:Description
336:comes from
326:West Bengal
149:F. Hamilton
128:Subfamily:
1747:Categories
1318:References
1277:Plamoottil
1156:Klausewitz
1124:Plamoottil
1051:Y. H. Wang
1024:Plamoottil
1010:Plamoottil
996:Plamoottil
970:Srivastava
854:Plamootitl
755:Plamoottil
726:Plamoottil
522:Barbonymus
470:homonymies
436:omnivorous
375:P. snyderi
362:South Asia
305:Pauciuncus
212:South Asia
210:native to
208:Cyprinidae
122:Cyprinidae
1456:0272-4634
1414:Sahyadria
957:Rema Devi
537:that has
506:subfamily
487:Cyprinion
366:Sri Lanka
332:The name
316:Etymology
294:from the
278:Sahyadria
272:Puntigrus
254:Haludaria
242:Dawkinsia
199:of small
183:See text
78:Kingdom:
72:Eukaryota
21:Puntigrus
1552:Wikidata
1375:Archived
1354:FishBase
837:R. Singh
833:Dobriyal
816:Maduwage
791:Maduwage
509:Barbinae
450:Taxonomy
430:Behavior
328:, India.
284:Systomus
260:Oliotius
236:Barbodes
177:Species
132:Barbinae
118:Family:
92:Chordata
88:Phylum:
82:Animalia
68:Domain:
1753:Puntius
1665:1432917
1639:2363821
1600:Puntius
1587:Puntius
1573:Puntius
1558:Q131627
1543:Puntius
1396:Puntius
1347:Puntius
1312:, 1872)
1281:Abraham
1252:, 1844)
1239:, 1868)
1237:Günther
1225:, 1822)
1201:, 1934)
1187:, 1868)
1185:Günther
1172:, 1822)
1158:, 1962)
1140:, 1933)
1098:, 1822)
1084:, 1871)
983:Jayaram
905:, 1844)
870:, 1868)
868:Günther
775:, 1938)
743:, 1849)
728:, 2015
693:, 1822)
678:, 1937)
664:, 1878)
650:, 1849)
648:Bleeker
636:, 1842)
621:, 1863)
619:Bleeker
606:, 1878)
592:, 1868)
590:Günther
577:, 1842)
563:, 1869)
545:Species
531:Puntius
517:Puntius
502:barbels
493:Capoeta
478:Puntius
456:Puntius
358:Puntius
334:Puntius
300:Sumatra
231:Puntius
227:species
203:in the
192:Puntius
143:Puntius
138:Genus:
108:Order:
98:Class:
32:Puntius
1730:270668
1717:364559
1704:230492
1678:163706
1454:
1310:F. Day
1297:, 1865
1295:F. Day
1283:, 2014
1279:&
1265:, 1929
1199:Fowler
1154:&
1112:, 1953
1082:F. Day
1070:, 2008
1057:, 1982
1053:&
1039:, 1865
1037:F. Day
1012:, 2014
998:, 2014
985:, 1990
959:, 1992
955:&
932:, 1878
930:F. Day
919:, 1945
891:, 2012
887:&
856:, 2019
822:, 2008
818:&
797:, 2008
793:&
771:&
741:Jerdon
662:F. Day
604:F. Day
561:F. Day
474:Barbus
461:Barbus
380:Taiwan
368:) and
346:Bangla
338:Pungti
296:Eocene
266:Pethia
220:Taiwan
205:family
151:, 1822
1725:WoRMS
1691:27707
1660:IRMNG
1652:91587
1613:86734
1138:Herre
1110:Silas
1055:Y. Ni
953:Menon
773:Misra
498:barbs
444:plant
378:, in
352:Range
344:), a
225:Many
197:genus
195:is a
1686:NCBI
1673:ITIS
1634:GBIF
1626:75HG
1608:BOLD
1452:ISSN
1152:Benl
1126:2020
769:Hora
757:2016
676:Hora
535:carp
500:and
476:and
442:and
424:carp
417:The
342:Puti
281:and
214:and
1621:CoL
1595:AFD
1582:ADW
1444:doi
298:of
1749::
1727::
1714::
1701::
1688::
1675::
1662::
1649::
1636::
1623::
1610::
1597::
1584::
1569::
1554::
1450:.
1440:40
1438:.
1434:.
1363:^
1351:.
1326:^
883:,
835:,
814:,
810:,
789:,
312:.
287:.
275:,
269:,
263:,
257:,
251:,
245:,
239:,
222:.
1458:.
1446::
1349:"
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1221:(
1197:(
1183:(
1168:(
1150:(
1136:(
1094:(
1080:(
901:(
866:(
767:(
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689:(
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660:(
646:(
632:(
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602:(
588:(
573:(
559:(
409:(
23:.
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