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Pygmy short-horned lizard

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318:(SVL) and is a flat-bodied, squat lizard with short (but not sharp) spikes crowning the head. It has a "snub-nosed" side profile compared to the sleeker snouts of many lizards, and relatively short legs. The trunk is fringed by one row of pointed scales, while the belly scales are smooth. Their colour is greyish-beige, yellowish, or reddish-brown, depending on seasonality and region. Additionally, there are two rows of larger, dark spots on the back. When ready to mate, or threatened or reacting defensively, this species' colours become more intense (commonly known as "fired-up") and, like all horned lizards, the pygmy short-horned lizard has the ability to squirt blood from its eyes in an effort confuse or scare predators. 75: 31: 50: 735: 306:
from Canada). When placed together, the two are easily distinguished at maturity, as the pygmy short-horned lizard is considerably smaller. The greater short-horned lizard is a highly variable species, as well, with different geographic populations exhibiting differences in colouration and
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evidence was revealed, the greater short-horned lizard was considered to be the same species as the pygmy short-horned lizard, but the two are now considered distinct, with the pygmy species occupying the northwest portion of the U.S. and, formerly, the extreme south of
349:(89% of their diet), while it has been documented that mature lizards consume fewer ants (around 72% or the diet). Yearlings, perhaps in an "experimental" phase, consume the lowest proportion of ants (60%), supplementing their diets with various 435:
Within its family, given that its habitat often freezes during winter, the pygmy short-horned lizard is known as a unique species that tolerates lower temperatures better than others, thus it can inhabit
385:. They may also browse on plant matter. Overall, throughout their lives, pygmy short-horned lizards always consume ants, though total quantities of a particular prey species will vary seasonally. 969: 293:, as both species have much of the same physiology, such as a squat body and small, "horn"-like, pointed scales located along the head, back and side body. Until recent 925: 676: 1002: 1022: 860: 886: 1017: 579: 551: 1027: 519: 1012: 891: 811: 485: 286: 74: 930: 393: 789: 654: 274: 456: 697: 260: 196: 751: 608: 604: 179: 507: 798: 1007: 563: 39: 917: 658: 535: 472: 239:. Like other horned lizards, it is often called a "horned toad" or "horny toad," but it is not a 69: 514:
California Natural History Guides. Oakland, California: University of California Press. 191 pp.
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The pygmy short-horned lizard measures roughly 1.25–2.5 in (3.2–6.4 cm) from snout-to-
634: 630: 329:, Canada, according to the Committee on the Status of Endangered Wildlife in Canada (COSEWIC). 951: 834: 575: 547: 515: 294: 803: 979: 964: 956: 716: 650: 468: 397: 303: 299: 225: 904: 739: 637:(April 2008). "Ecology and ontogenetic variation of diet in the pigmy short-horned lizard ( 341:
varies from region to region, by seasonal availability, and by an individual's age or sex;
30: 938: 401: 268: 228: 146: 531: 996: 899: 232: 221: 156: 59: 54: 662: 839: 413: 136: 677:"Cold-Blooded and Cold-Tolerant: Finding Reptiles In the North Cascades Ecosystem" 574:. The Peterson Field Guide Series. Boston and New York: Houghton Mifflin Company. 912: 873: 783: 382: 378: 734: 816: 322: 308: 307:
patterning, in addition to overall size; some authorities describe up to five
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The pygmy short-horned lizard is often mistaken for its close relative, the
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A Field Guide to Western Reptiles and Amphibians: Third Edition
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10.1674/0003-0031(2008)159[327:EAOVOD]2.0.CO;2
704:, brought from the Columbia River by Mr. Douglass ". 325:
of pygmy short-horned lizard is a species at risk in
758: 512:Introduction to Horned Lizards of North America. 706:Transactions of the Linnean Society of London 546:. Baltimore: Johns Hopkins University Press. 440:not generally accessible to related species. 8: 605:"Phrynosoma douglassi (Short-horned lizard)" 503: 501: 499: 473:10.2305/IUCN.UK.2007.RLTS.T64075A12741891.en 231:. The species is native to the northwestern 746: 345:and hatchlings feed almost exclusively on 48: 29: 20: 700:(1828). "Description of a new Species of 570:", pp. 303–304 + Plate 34 + Map 101. in 449: 7: 598: 596: 594: 592: 590: 588: 1003:IUCN Red List least concern species 1023:Fauna of the Western United States 14: 544:The Eponym Dictionary of Reptiles 733: 73: 467:Red List of Threatened Species 1: 1018:Reptiles of the United States 538:; Grayson, Michael. (2011). " 643:American Midland Naturalist 287:greater short-horned lizard 1044: 1028:Reptiles described in 1828 420:, and potentially larger, 235:and adjacent southwestern 24:Pygmy short-horned lizard 740:Pygmy short-horned lizard 603:Cossel Jr., John (1997). 493:www.reptile-database.org. 394:long-nosed leopard lizard 208:pygmy short-horned lizard 185: 178: 70:Scientific classification 68: 46: 37: 28: 23: 1013:Lizards of North America 681:North Cascades Institute 711:: 105–107 + Plate X. (" 609:Idaho State University 715:", new species). (in 639:Phrynosoma douglasii) 804:Phrynosoma_douglasii 790:Phrynosoma douglasii 760:Phrynosoma douglasii 568:Phrynosoma douglasii 491:The Reptile Database 486:Phrynosoma douglasii 461:Phrynosoma douglasii 333:Behavior and ecology 213:Phrynosoma douglasii 189:Phrynosoma douglasii 540:Phrynosoma douglasi 508:Sherbrooke, Wade C. 40:Conservation status 564:Stebbins, Robert C 990: 989: 952:Open Tree of Life 752:Taxon identifiers 580:978-0-395-98272-3 552:978-1-4214-0135-5 295:mitochondrial DNA 267:, is in honor of 204: 203: 171:P. douglasii 63: 16:Species of lizard 1035: 983: 982: 973: 972: 960: 959: 947: 946: 934: 933: 921: 920: 908: 907: 895: 894: 882: 881: 869: 868: 856: 855: 843: 842: 830: 829: 820: 819: 807: 806: 794: 793: 792: 779: 778: 777: 747: 738:Data related to 737: 713:Agama Douglassii 685: 684: 673: 667: 666: 627: 621: 620: 618: 616: 600: 583: 561: 555: 536:Watkins, Michael 529: 523: 505: 494: 481: 475: 454: 402:northern shrikes 300:British Columbia 243:at all. It is a 191: 78: 77: 57: 52: 51: 33: 21: 1043: 1042: 1038: 1037: 1036: 1034: 1033: 1032: 993: 992: 991: 986: 978: 976: 968: 963: 955: 950: 942: 939:Observation.org 937: 929: 924: 916: 911: 903: 898: 890: 885: 877: 872: 864: 859: 851: 846: 838: 833: 825: 823: 815: 810: 802: 797: 788: 787: 782: 773: 772: 767: 754: 730: 694: 692:Further reading 689: 688: 675: 674: 670: 635:Beck, Daniel D. 631:Lahti, Megan E. 629: 628: 624: 614: 612: 607:. www.ISU.edu ( 602: 601: 586: 562: 558: 530: 526: 506: 497: 482: 478: 457:Hammerson, G.A. 455: 451: 446: 335: 283: 257: 229:Phrynosomatidae 200: 193: 187: 174: 147:Phrynosomatidae 72: 64: 53: 49: 42: 17: 12: 11: 5: 1041: 1039: 1031: 1030: 1025: 1020: 1015: 1010: 1005: 995: 994: 988: 987: 985: 984: 974: 961: 948: 935: 922: 909: 896: 883: 870: 857: 844: 831: 821: 808: 795: 780: 764: 762: 756: 755: 750: 744: 743: 742:at Wikispecies 729: 728:External links 726: 725: 724: 693: 690: 687: 686: 668: 649:(2): 327–339. 622: 584: 556: 524: 520:978-0520228276 495: 476: 448: 447: 445: 442: 361:, terrestrial 334: 331: 291:P. hernandesi) 282: 281:Identification 279: 256: 253: 202: 201: 194: 183: 182: 176: 175: 168: 166: 162: 161: 154: 150: 149: 144: 140: 139: 134: 130: 129: 124: 120: 119: 114: 110: 109: 104: 100: 99: 94: 90: 89: 84: 80: 79: 66: 65: 47: 44: 43: 38: 35: 34: 26: 25: 15: 13: 10: 9: 6: 4: 3: 2: 1040: 1029: 1026: 1024: 1021: 1019: 1016: 1014: 1011: 1009: 1006: 1004: 1001: 1000: 998: 981: 975: 971: 966: 962: 958: 953: 949: 945: 940: 936: 932: 927: 923: 919: 914: 910: 906: 901: 897: 893: 888: 884: 880: 875: 871: 867: 862: 858: 854: 849: 845: 841: 836: 832: 828: 822: 818: 813: 809: 805: 800: 796: 791: 785: 781: 776: 770: 766: 765: 763: 761: 757: 753: 748: 741: 736: 732: 731: 727: 722: 718: 714: 710: 707: 703: 699: 696: 695: 691: 682: 678: 672: 669: 664: 660: 656: 652: 648: 644: 640: 636: 632: 626: 623: 610: 606: 599: 597: 595: 593: 591: 589: 585: 581: 577: 573: 569: 565: 560: 557: 553: 549: 545: 541: 537: 533: 528: 525: 521: 517: 513: 509: 504: 502: 500: 496: 492: 488: 487: 480: 477: 474: 470: 466: 462: 458: 453: 450: 443: 441: 439: 433: 431: 427: 423: 419: 415: 414:birds of prey 411: 407: 403: 399: 398:Stellar's jay 395: 391: 388:Predators of 386: 384: 380: 379:Lepidopterans 377:, as well as 376: 372: 368: 364: 360: 356: 352: 348: 344: 340: 332: 330: 328: 324: 319: 317: 312: 310: 305: 301: 296: 292: 288: 280: 278: 276: 275:David Douglas 273: 270: 266: 262: 261:specific name 254: 252: 250: 246: 242: 238: 234: 233:United States 230: 227: 223: 222:horned lizard 219: 215: 214: 209: 198: 192: 190: 184: 181: 180:Binomial name 177: 173: 172: 167: 164: 163: 160: 159: 155: 152: 151: 148: 145: 142: 141: 138: 135: 132: 131: 128: 125: 122: 121: 118: 115: 112: 111: 108: 105: 102: 101: 98: 95: 92: 91: 88: 85: 82: 81: 76: 71: 67: 61: 56: 55:Least Concern 45: 41: 36: 32: 27: 22: 19: 759: 712: 708: 705: 701: 698:Bell, Thomas 680: 671: 646: 642: 638: 625: 613:. Retrieved 571: 567: 559: 543: 539: 527: 511: 484: 479: 460: 452: 434: 392:include the 390:P. douglassi 389: 387: 383:caterpillars 339:P. douglasii 338: 337:The diet of 336: 321:The eastern 320: 313: 290: 284: 264: 258: 212: 211: 207: 205: 188: 186: 170: 169: 157: 18: 913:NatureServe 874:iNaturalist 784:Wikispecies 566:. (2003). " 532:Beolens, Bo 424:-consuming 357:and grubs, 1008:Phrynosoma 997:Categories 444:References 430:kingsnakes 428:, such as 381:and their 375:earthworms 323:subspecies 309:subspecies 304:extirpated 158:Phrynosoma 133:Suborder: 970:douglasii 542:", p. 75 459:(2007). " 351:arachnids 265:douglasii 255:Etymology 249:amphibian 247:, not an 220:of small 165:Species: 93:Kingdom: 87:Eukaryota 918:2.102494 824:BioLib: 769:Wikidata 663:54206550 510:(2003). 483:Species 422:squamate 363:mollusks 343:neonates 272:botanist 269:Scottish 143:Family: 127:Squamata 117:Reptilia 107:Chordata 103:Phylum: 97:Animalia 83:Domain: 60:IUCN 3.1 866:2451089 775:Q898028 717:English 683:. 2019. 463:". The 418:bobcats 406:coyotes 359:isopods 355:beetles 327:Alberta 245:reptile 224:in the 218:species 216:) is a 199:, 1828) 153:Genus: 137:Iguania 123:Order: 113:Class: 58: ( 980:212141 977:uBio: 957:968165 944:939223 892:564567 853:PRYNDO 827:185020 817:762927 661:  615:14 May 578:  550:  518:  438:biomes 426:snakes 373:) and 367:snails 237:Canada 226:family 931:43611 905:64075 879:36316 840:4GWSQ 721:Latin 702:Agama 659:S2CID 410:foxes 371:slugs 302:(now 926:NCBI 900:IUCN 887:ITIS 861:GBIF 848:EPPO 719:and 617:2024 576:ISBN 548:ISBN 516:ISBN 465:IUCN 347:ants 316:vent 311:. 259:The 241:toad 206:The 197:Bell 835:CoL 812:BHL 799:ADW 651:doi 647:159 641:". 489:at 469:doi 432:. 999:: 967:: 965:RD 954:: 941:: 928:: 915:: 902:: 889:: 876:: 863:: 850:: 837:: 814:: 801:: 786:: 771:: 723:). 709:16 679:. 657:. 645:. 633:; 587:^ 534:; 498:^ 416:, 412:, 408:, 404:, 400:, 396:, 369:, 353:, 277:. 263:, 251:. 665:. 653:: 619:. 611:) 582:. 554:. 522:. 471:: 365:( 289:( 210:( 195:( 62:)

Index


Conservation status
Least Concern
IUCN 3.1
Scientific classification
Edit this classification
Eukaryota
Animalia
Chordata
Reptilia
Squamata
Iguania
Phrynosomatidae
Phrynosoma
Binomial name
Bell
species
horned lizard
family
Phrynosomatidae
United States
Canada
toad
reptile
amphibian
specific name
Scottish
botanist
David Douglas
greater short-horned lizard

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