295:, a mountainous area that passes through Cambodia and Vietnam. Its habitat is mostly characterized by evergreen forest in the mountains, in the middle to upper canopy. They move around quadrupedally and by brachiation up in the trees. This species is unique with its coloration among the doucs as it has a bluish face with yellow rings around its eyes a blue scrotum and a pink penis. Like other doucs, this species has a tail as long as its body and head length. Black-shanked douc have been observed in groups ranging from 3 to 30 individuals, depending on their habitat. Group tend to have a fission-fusion dynamic that changes with food availability. Their diet varies from dry to wet season. Regardless of the season, their diet consists mostly of leaves, but they have also been found to consume considerable amounts of fruits and flowers during wet season. The species changed conservation status in 2015 from
398:, one-male units averaged 7.5 individuals, and were composed of a single male, several females and offspring. Their social hierarchy is therefore male dominant. Black-shanked doucs have a fission-fusion social system, with fusion of one-male units into bands occurring more commonly in the wet season due to higher food availability. Group size can vary from 3 up to 35 individuals. The reason for such variability in size has been associated with the quality of their habitat and the level of human disturbance. Groups that were observed in the wild were of multi-male, multi-female disposition, with a higher ratio of males, the bigger the group was.
382:
light grey. Their rest of their body is black, which includes legs and arms, excluding the pubic patch area which is white, their scrotum is blue, and their penis is pink. While the majority of individuals have those standard colors, there has been color variations observed in few individuals, who were confirmed by genetic sampling to be black-shanked doucs. Amongst those observations, color variations occurred for the forearms which were partially or completely white and the lower leg which was dark red. Though it is not fully understood, an explanation for those color variations would be due to hybrid between douc species.
357:, but the black-shanked douc is only found in Cambodia and Vietnam. Studies on douc species have shown differentiation in ecological niche. The findings were that each douc species are spread out from North to South of Vietnam, each having their own latitudinal range. For the black-shanked douc, they estimated its geographic range is at a narrow latitudinal range of 11° N to 13.5°N, in Eastern Cambodia and Southern Vietnam. They can be found living in different types of forest: broad-leafed evergreen, semievergreen, mixed, dipterocarp lowland. More specifically, those species of the
418:. During the dry season, the consumption of leaves increased, with a smaller amounts of other food consumed like flowers and fruits. During wet season, the consumption of leaves had higher diversity of plant species and the consumption of fruits and flowers was higher. This can be explained by Studies showed that variation of food intake also varied throughout the day. Leaves are consumed consistently throughout the day, but fruits are consumed more before midday, a habit common in folivorous non-human primate species for a more efficient energy intake needed in the morning.
365:, a mountainous landscape in Eastern Indochina. Like all douc species, the black-shanked douc is an arboreal species that moves around quadrupedally and through brachiation as locomotion. It is mostly found in the middle to upper forest canopy for most of the time. The range of elevation of the terrain where they have been seen goes from sea level to 1500 meters. However, it is important to note that most of the forest at lower elevation have been torn down over the years. Therefore, most sightings occur at higher elevations.
323:
410:. Studies in Cambodia and Vietnam have shown their diet to consist primarily of leaves, from a wide diversity of plant species. Studies showed they consume as many as 150 different plant species, which shows they do not have particular preferences in plants, they consume whichever is available. Fruits, flowers, and seeds were reported as the remaining portion of their diet at various proportions, according to food availability. Indeed, studies found that their diet changed from
111:
69:
449:, an increase in agricultural demands caused further deforestation. Increasing demands from the timber industry and mining industry also spurred deforestation. In addition, urbanization of areas in Vietnam have caused great habitat loss and fragmentation to convert forests into urban areas. Illegal conversion of protected areas into agricultural land like banana farms is still occurring in Vietnam.
42:
457:
during the 21st century. Its spatial distribution currently does not cover a wide area and the species harbours a habitat that is currently getting more depleted. Therefore, habitat modelling of climate change has shown it will be difficult for black-shanked douc to find habitat suitable for their survival.
381:
genus. Their most unique characteristic is their bluish face with large yellow eye-rings. It also has thin and short whiskers. The outline of its face has a thin chestnut coloration before the back of its head which is black. The region around its throat is white. Its back is dark grey. Its belly is
452:
Poaching, hunting and wildlife trade is the biggest threat to primates in
Vietnam, as they are hunted and sold they can serve as food, medicine and pets. Even though those practices are illegal, they are still an issue in Vietnam and Cambodia. Conservation efforts have been made by establishing gun
385:
It has a tail as long as its body and head length. Few individuals were measured but the results for the male black-shanked douc had an average head and body length of 50 cm (20 in), an average tail length of 69 cm (27 in) and an average weight of 5.3 kg (12 lb). While
456:
Climate change has also raised concern about the conservation of the black-shanked douc as endemic species tend to be more at risk of extinction due to climate change. Indeed, it has been modelled that black-shanked douc would be a vulnerable species with an increase of global average temperatures
441:
due to a more alarming rapid rate of population decline. The main drivers of the population decline for black-shanked doucs has been habitat loss and fragmentation. Other factors such as poaching and wildlife trade have been added as threatening factors for the conservation of this specie but they
252:
315:, a population that has remained stable over the last decade. The largest populations estimated to be in Vietnam is around 500-600 individuals. The biggest challenges that the black-shanked douc faces in terms of conservation are
307:. This reassessment is due to an increase of the rate of population decline. No global population estimate exists. The majority of the population can be found in Cambodia, with smaller populations in Vietnam. In fact, the
802:. Frankfurt Zoological Society - Vietnam Primate Conservation Program Endangered Primate Rescue Center, Cuc Phuong National Park and Conservation International, Indo-Burma Program: 91–97.
2337:
611:
Nuttall, Matthew N.; Griffin, Olly; Fewster, Rachel M.; McGowan, Philip J. K.; Abernethy, Katharine; O'Kelly, Hannah; Nut, Menghor; Sot, Vandoeun; Bunnefeld, Nils (2021).
319:
and illegal poaching. Conservation efforts are being made to control illegal poaching and trade in
Vietnam by putting laws against hunting and trading threatened species.
930:"Modelling the change in the distribution of the black-shanked douc, Pygathrix nigripes (Milne-Edwards) in the context of climate change: Implications for conservation"
377:, the black-shanked douc is characterized by its colorations. The colors on its head and neck are what differentiates the black-shanked douc from other species of the
846:
671:
2362:
2257:
780:. Vietnam: Frankfurt Zoological Society & Cuc Phuong National Park Conservation Program & Fauna & Flora International, Vietnam Program. p. 226.
991:
394:
Four major social units occur in the species: one-male units, bands comprising several one-male units, bachelor groups and lone males. In a study in
2179:
928:
Tran, Dung V.; Vu, Thinh T.; Tran, Bao Q.; Nguyen, Manh D.; Vu, Phuong T.; Tran, Trang H.; Nguyen, Hoa T.; Pham, Thong V.; Nguyen, Thanh C. (2020).
2205:
386:
the female black-shanked douc had an average head and body length of 56 cm (22 in) and an average weight of 5.4 kg (12 lb).
2357:
969:
1956:
492:
2231:
445:
Habitat loss and habitat fragmentation in
Vietnam has been intensive during the 20th century due to deforestation. Following the
2249:
2210:
1692:
1157:
984:
676:
308:
73:
2347:
395:
312:
594:
50:
2352:
2244:
2140:
1746:
1386:
1737:
482:
2262:
2051:
110:
1983:
1728:
1274:
1974:
1947:
1660:
1507:
1395:
1292:
977:
1247:
1184:
2132:
1543:
1534:
1238:
1175:
1063:
2105:
2067:
1965:
1701:
1588:
1579:
1229:
241:
2145:
1606:
2296:
1818:
1710:
438:
322:
300:
225:
1265:
847:"Color variation in black-shanked douc langurs (Pygathrix nigripes), and some behavioural observations"
2166:
2114:
1854:
1773:
1764:
1651:
1220:
1211:
1193:
1166:
543:
1633:
1480:
1413:
1368:
1256:
1202:
1148:
316:
58:
929:
1827:
1404:
1283:
907:
825:
The socio-ecology of the black-shanked douc (Pygathrix nigripes) in
Mondulkiri Province, Cambodia
745:
691:
652:
434:
427:
296:
105:
2301:
1809:
1791:
828:
2283:
2153:
2031:
2022:
1930:
1915:
1525:
1498:
1462:
1453:
1377:
1359:
952:
899:
737:
644:
613:"Long-term monitoring of wildlife populations for protected area management in Southeast Asia"
498:
488:
2288:
2007:
1998:
1897:
1800:
1001:
944:
891:
729:
681:
634:
624:
538:
192:
2223:
2119:
2270:
1836:
1489:
1116:
598:
251:
2342:
1845:
1755:
1719:
1642:
1471:
1437:
1422:
1107:
1089:
453:
control laws, and prohibition of threatened species hunting and trade laws in
Vietnam.
362:
292:
520:
2331:
2218:
1624:
1615:
1570:
1342:
1316:
1307:
1080:
795:
717:
695:
656:
529:
478:
431:
304:
78:
911:
749:
2158:
1782:
1098:
474:
41:
17:
2314:
2236:
2192:
2099:
1552:
1131:
446:
686:
1045:
895:
794:
Bang, Tran Van; Hoang Minh, Duc; Luu, Hong Truong; Covert, Herbert H. (2010).
733:
437:
due to the rate of its population decline. In 2015, its status was updated as
415:
411:
172:
2090:
956:
903:
741:
648:
1675:
1008:
879:
591:
502:
122:
948:
565:
2275:
2084:
1027:
776:
Nadler, Tilo; Momberg, Frank; Dang, Nguyen Xuan; Lormee, Nicolas (2003).
407:
346:
342:
284:
142:
487:(3rd ed.). Baltimore: Johns Hopkins University Press. p. 173.
2184:
1039:
354:
288:
276:
162:
639:
2197:
2127:
1033:
1021:
778:
Vietnam
Primate Conservation Status Review 2002 - Part 2 Leaf Monkeys
629:
612:
182:
152:
132:
2061:
716:
Bett, Nolan N.; Blair, Mary E.; Sterling, Eleanor J. (August 2012).
519:
Duc, H.; Quyet, L.K.; Rawson, B.M.; O'Brien, J.; Covert, H. (2021).
2309:
321:
95:
89:
672:"Status of Key Species in Keo Seima Wildlife Sanctuary 2010-2020"
484:
Mammal
Species of the World: A Taxonomic and Geographic Reference
2171:
1880:
350:
336:
280:
202:
2065:
973:
878:
Duc, Hoang Minh; Baxter, G. S.; Page, Manda J. (2009-02-01).
829:
https://openresearch-repository.anu.edu.au/handle/1885/49396
796:"Conservation status of primates in Ta Kou nature reserve"
718:"Ecological Niche Conservatism in Doucs (Genus Pygathrix)"
291:. The region they are mostly found in is called the
2074:
2020:
1996:
1928:
1878:
1871:
1673:
1435:
1340:
1333:
1305:
1129:
1061:
1054:
880:"Diet of Pygathrix nigripes in Southern Vietnam"
426:In 2000, the black-shanked douc was assessed as
311:reports almost 25,000 individuals in Cambodia's
544:10.2305/IUCN.UK.2021-1.RLTS.T39828A196138291.en
985:
592:Unexpected Large Monkey Population Discovered
8:
287:, with some smaller populations in Southern
2338:IUCN Red List critically endangered species
2062:
1875:
1337:
1058:
992:
978:
970:
250:
67:
40:
31:
685:
638:
628:
542:
442:are not as detrimental as habitat loss.
670:Griffin, O.; Nuttall, M. (2020-12-04).
601:Newswise, Retrieved on August 28, 2008.
466:
283:found mostly in the forests of Eastern
514:
512:
923:
921:
873:
871:
869:
867:
840:
838:
836:
819:
817:
815:
813:
811:
809:
800:Conservation of Primates in Indochina
7:
2363:Taxa named by Alphonse Milne-Edwards
884:International Journal of Primatology
789:
787:
771:
769:
767:
765:
763:
761:
759:
722:International Journal of Primatology
711:
709:
707:
705:
587:
585:
326:Black-shanked douc in Safery Phuquoc
1957:Black-and-white snub-nosed monkey (
530:IUCN Red List of Threatened Species
432:IUCN Red List of Threatened Species
305:IUCN Red List of Endangered Species
25:
854:Vietnamese Journal of Primatology
617:Conservation Science and Practice
109:
49:Black-shanked douc with baby at
1693:Black-crested Sumatran langur (
406:Black-shanked doucs are mostly
361:genus are mostly found in the
345:species, that can be found in
1:
1747:East Sumatran banded langur (
1387:Northern plains gray langur (
677:Wildlife Conservation Society
309:Wildlife Conservation Society
1158:Miss Waldron's red colobus (
396:Keo Seima Wildlife Sanctuary
331:Geographic range and habitat
313:Keo Seima Wildlife Sanctuary
1984:Myanmar snub-nosed monkey (
937:Raffles Bulletin of Zoology
27:Species of Old World monkey
2379:
1975:Tonkin snub-nosed monkey (
1948:Golden snub-nosed monkey (
1738:Robinson's banded langur (
1661:Indochinese black langur (
1508:Selangor silvered langur (
1396:Black-footed gray langur (
1293:Ulindi River red colobus (
687:10.19121/2020.Report.38511
2358:Mammals described in 1871
2048:
1535:Indochinese grey langur (
1239:Niger Delta red colobus (
1016:
1000:Extant species of family
896:10.1007/s10764-008-9325-y
734:10.1007/s10764-012-9622-3
259:Black-shanked douc range
258:
249:
231:
224:
106:Scientific classification
104:
87:
65:
56:
48:
39:
34:
1966:Gray snub-nosed monkey (
1729:Raffles' banded langur (
1702:Black-and-white langur (
1275:Oustalet's red colobus (
1230:Tana River red colobus (
1071:(Black and white colobi)
481:; Reeder, D. M. (eds.).
341:genus are all Indochina
1819:Sabah grizzled langur (
1711:Black Sumatran langur (
1248:Bouvier's red colobus (
1185:Thollon's red colobus (
1855:Natuna Island surili (
1774:White-fronted surili (
1765:White-thighed surili (
1544:Phayre's leaf monkey (
1221:Zanzibar red colobus (
1212:Udzungwa red colobus (
1176:Preuss's red colobus (
949:10.26107/RBZ-2020-0088
823:Rawson, B. M. (2009).
537:: e.T39828A196138291.
327:
51:Cát Tiên National Park
2297:Paleobiology Database
1634:White-headed langur (
1589:Gee's golden langur (
1580:Shortridge's langur (
1414:Purple-faced langur (
1369:Kashmir gray langur (
1257:Semliki red colobus (
1203:Ugandan red colobus (
1149:Western red colobus (
845:Nadler, Tilo (2008).
439:Critically Endangered
325:
301:critically endangered
74:Critically Endangered
1938:(Snub-nosed monkeys)
1907:Black-shanked douc (
1828:Maroon leaf monkey (
1405:Tufted gray langur (
1284:Lomami red colobus (
1266:Lang's red colobus (
566:"Appendices | CITES"
373:Like all species of
2348:Mammals of Cambodia
2032:Pig-tailed langur (
1916:Gray-shanked douc (
1652:Delacour's langur (
1526:Dusky leaf monkey (
1499:Tenasserim lutung (
1463:West Javan langur (
1454:East Javan langur (
1378:Tarai gray langur (
1360:Nepal gray langur (
1194:Foa's red colobus (
1167:Pennant's colobus (
1004:(Old World monkeys)
275:) is an endangered
59:Conservation status
35:Black-shanked douc
2353:Mammals of Vietnam
2133:pygathrix-nigripes
2120:Pygathrix_nigripes
2106:Pygathrix nigripes
2076:Pygathrix nigripes
2008:Proboscis monkey (
1898:Red-shanked douc (
1607:François' langur (
1481:Germain's langur (
597:2011-10-04 at the
523:Pygathrix nigripes
328:
272:Pygathrix nigripes
267:black-shanked douc
235:Pygathrix nigripes
18:Pygathrix nigripes
2325:
2324:
2284:Open Tree of Life
2068:Taxon identifiers
2059:
2058:
2044:
2043:
1941:
1891:
1867:
1866:
1837:Mentawai langur (
1810:Miller's langur (
1792:Thomas's langur (
1686:
1490:Annamese langur (
1353:
1329:
1328:
1142:
1117:Mantled guereza (
1074:
1005:
827:. Retrieved from
335:Species from the
263:
262:
99:
82:
16:(Redirected from
2370:
2318:
2317:
2305:
2304:
2292:
2291:
2279:
2278:
2266:
2265:
2253:
2252:
2240:
2239:
2227:
2226:
2214:
2213:
2201:
2200:
2188:
2187:
2175:
2174:
2162:
2161:
2149:
2148:
2136:
2135:
2123:
2122:
2110:
2109:
2108:
2095:
2094:
2093:
2063:
1940:
1939:
1935:
1890:
1889:
1885:
1876:
1846:Siberut langur (
1756:Sarawak surili (
1720:Mitered langur (
1685:
1684:
1680:
1643:Laotian langur (
1636:T. leucocephalus
1627:T. poliocephalus
1510:T. selangorensis
1472:Silvery lutung (
1423:Nilgiri langur (
1352:
1351:
1347:
1338:
1141:
1140:
1136:
1108:Ursine colobus (
1090:Angola colobus (
1073:
1072:
1068:
1059:
1003:
1002:Cercopithecidae
994:
987:
980:
971:
961:
960:
934:
925:
916:
915:
875:
862:
861:
851:
842:
831:
821:
804:
803:
791:
782:
781:
773:
754:
753:
713:
700:
699:
689:
667:
661:
660:
642:
632:
630:10.1111/csp2.614
608:
602:
589:
580:
579:
577:
576:
562:
556:
555:
553:
551:
546:
516:
507:
506:
471:
254:
242:A. Milne-Edwards
237:
217:P. nigripes
114:
113:
93:
76:
71:
70:
44:
32:
21:
2378:
2377:
2373:
2372:
2371:
2369:
2368:
2367:
2328:
2327:
2326:
2321:
2313:
2308:
2300:
2295:
2287:
2282:
2274:
2271:Observation.org
2269:
2261:
2256:
2248:
2243:
2235:
2230:
2222:
2217:
2209:
2204:
2196:
2191:
2183:
2178:
2170:
2165:
2157:
2152:
2144:
2139:
2131:
2126:
2118:
2113:
2104:
2103:
2098:
2089:
2088:
2083:
2070:
2060:
2055:
2040:
2016:
1992:
1937:
1936:
1934:
1924:
1887:
1886:
1884:
1872:Odd-nosed group
1863:
1801:Hose's langur (
1682:
1681:
1679:
1669:
1625:Cat Ba langur (
1616:Hatinh langur (
1571:Capped langur (
1431:
1349:
1348:
1346:
1325:
1317:Olive colobus (
1301:
1259:P. semlikiensis
1232:P. rufomitratus
1205:P. tephrosceles
1138:
1137:
1135:
1125:
1081:Black colobus (
1070:
1069:
1067:
1050:
1012:
998:
967:
965:
964:
943:(452): 769778.
932:
927:
926:
919:
877:
876:
865:
849:
844:
843:
834:
822:
807:
793:
792:
785:
775:
774:
757:
715:
714:
703:
669:
668:
664:
610:
609:
605:
599:Wayback Machine
590:
583:
574:
572:
564:
563:
559:
549:
547:
518:
517:
510:
495:
473:
472:
468:
463:
424:
404:
392:
371:
333:
245:
239:
233:
220:
193:Cercopithecidae
108:
100:
83:
72:
68:
61:
28:
23:
22:
15:
12:
11:
5:
2376:
2374:
2366:
2365:
2360:
2355:
2350:
2345:
2340:
2330:
2329:
2323:
2322:
2320:
2319:
2306:
2293:
2280:
2267:
2254:
2241:
2228:
2215:
2202:
2189:
2176:
2163:
2150:
2137:
2124:
2111:
2096:
2080:
2078:
2072:
2071:
2066:
2057:
2056:
2049:
2046:
2045:
2042:
2041:
2039:
2038:
2028:
2026:
2018:
2017:
2015:
2014:
2004:
2002:
1994:
1993:
1991:
1990:
1981:
1972:
1963:
1954:
1944:
1942:
1926:
1925:
1923:
1922:
1913:
1904:
1894:
1892:
1873:
1869:
1868:
1865:
1864:
1862:
1861:
1852:
1843:
1834:
1825:
1816:
1807:
1798:
1789:
1783:Javan surili (
1780:
1771:
1762:
1758:P. chrysomelas
1753:
1744:
1735:
1726:
1717:
1708:
1699:
1689:
1687:
1671:
1670:
1668:
1667:
1658:
1649:
1640:
1631:
1622:
1618:T. hatinhensis
1613:
1596:
1595:
1586:
1582:T. shortridgei
1577:
1560:
1559:
1550:
1541:
1537:T. crepusculus
1532:
1515:
1514:
1505:
1496:
1487:
1478:
1469:
1460:
1443:
1441:
1438:Trachypithecus
1433:
1432:
1430:
1429:
1420:
1411:
1402:
1393:
1384:
1375:
1366:
1362:S. schistaceus
1356:
1354:
1350:(Gray langurs)
1335:
1331:
1330:
1327:
1326:
1324:
1323:
1313:
1311:
1303:
1302:
1300:
1299:
1290:
1286:P. parmentieri
1281:
1272:
1263:
1254:
1245:
1236:
1227:
1218:
1209:
1200:
1191:
1182:
1173:
1164:
1155:
1145:
1143:
1127:
1126:
1124:
1123:
1114:
1105:
1099:King colobus (
1096:
1087:
1077:
1075:
1056:
1052:
1051:
1049:
1048:
1042:
1036:
1030:
1024:
1017:
1014:
1013:
999:
997:
996:
989:
982:
974:
963:
962:
917:
863:
832:
805:
783:
755:
728:(4): 972–988.
701:
662:
603:
581:
557:
508:
493:
465:
464:
462:
459:
423:
420:
403:
400:
391:
388:
370:
367:
363:Annamite Range
332:
329:
293:Annamite Range
261:
260:
256:
255:
247:
246:
240:
229:
228:
222:
221:
214:
212:
208:
207:
200:
196:
195:
190:
186:
185:
180:
176:
175:
170:
166:
165:
160:
156:
155:
150:
146:
145:
140:
136:
135:
130:
126:
125:
120:
116:
115:
102:
101:
88:
85:
84:
66:
63:
62:
57:
54:
53:
46:
45:
37:
36:
26:
24:
14:
13:
10:
9:
6:
4:
3:
2:
2375:
2364:
2361:
2359:
2356:
2354:
2351:
2349:
2346:
2344:
2341:
2339:
2336:
2335:
2333:
2316:
2311:
2307:
2303:
2298:
2294:
2290:
2285:
2281:
2277:
2272:
2268:
2264:
2259:
2255:
2251:
2246:
2242:
2238:
2233:
2229:
2225:
2220:
2216:
2212:
2207:
2203:
2199:
2194:
2190:
2186:
2181:
2177:
2173:
2168:
2164:
2160:
2155:
2151:
2147:
2142:
2138:
2134:
2129:
2125:
2121:
2116:
2112:
2107:
2101:
2097:
2092:
2086:
2082:
2081:
2079:
2077:
2073:
2069:
2064:
2054:
2053:
2047:
2037:
2035:
2030:
2029:
2027:
2025:
2024:
2019:
2013:
2011:
2006:
2005:
2003:
2001:
2000:
1995:
1989:
1987:
1982:
1980:
1978:
1973:
1971:
1969:
1964:
1962:
1960:
1955:
1953:
1951:
1946:
1945:
1943:
1933:
1932:
1931:Rhinopithecus
1927:
1921:
1919:
1914:
1912:
1910:
1905:
1903:
1901:
1896:
1895:
1893:
1883:
1882:
1877:
1874:
1870:
1860:
1858:
1853:
1851:
1849:
1844:
1842:
1840:
1839:P. potenziani
1835:
1833:
1831:
1826:
1824:
1822:
1817:
1815:
1813:
1808:
1806:
1804:
1799:
1797:
1795:
1790:
1788:
1786:
1781:
1779:
1777:
1772:
1770:
1768:
1763:
1761:
1759:
1754:
1752:
1750:
1745:
1743:
1741:
1736:
1734:
1732:
1727:
1725:
1723:
1718:
1716:
1714:
1709:
1707:
1705:
1700:
1698:
1696:
1695:P. melalophos
1691:
1690:
1688:
1678:
1677:
1672:
1666:
1664:
1659:
1657:
1655:
1650:
1648:
1646:
1641:
1639:
1637:
1632:
1630:
1628:
1623:
1621:
1619:
1614:
1612:
1610:
1604:
1602:
1598:
1597:
1594:
1592:
1587:
1585:
1583:
1578:
1576:
1574:
1568:
1566:
1562:
1561:
1558:
1556:
1553:Popa langur (
1551:
1549:
1547:
1542:
1540:
1538:
1533:
1531:
1529:
1523:
1521:
1517:
1516:
1513:
1511:
1506:
1504:
1502:
1497:
1495:
1493:
1488:
1486:
1484:
1479:
1477:
1475:
1470:
1468:
1466:
1461:
1459:
1457:
1451:
1449:
1445:
1444:
1442:
1440:
1439:
1434:
1428:
1426:
1421:
1419:
1417:
1412:
1410:
1408:
1403:
1401:
1399:
1398:S. hypoleucos
1394:
1392:
1390:
1385:
1383:
1381:
1376:
1374:
1372:
1367:
1365:
1363:
1358:
1357:
1355:
1345:
1344:
1343:Semnopithecus
1339:
1336:
1332:
1322:
1320:
1315:
1314:
1312:
1310:
1309:
1304:
1298:
1296:
1295:P. lulindicus
1291:
1289:
1287:
1282:
1280:
1278:
1273:
1271:
1269:
1264:
1262:
1260:
1255:
1253:
1251:
1246:
1244:
1242:
1237:
1235:
1233:
1228:
1226:
1224:
1219:
1217:
1215:
1214:P. gordonorum
1210:
1208:
1206:
1201:
1199:
1197:
1192:
1190:
1188:
1183:
1181:
1179:
1174:
1172:
1170:
1165:
1163:
1161:
1156:
1154:
1152:
1147:
1146:
1144:
1134:
1133:
1128:
1122:
1120:
1115:
1113:
1111:
1110:C. vellerosus
1106:
1104:
1102:
1097:
1095:
1093:
1092:C. angolensis
1088:
1086:
1084:
1079:
1078:
1076:
1066:
1065:
1060:
1057:
1055:African group
1053:
1047:
1043:
1041:
1037:
1035:
1031:
1029:
1025:
1023:
1019:
1018:
1015:
1010:
1006:
995:
990:
988:
983:
981:
976:
975:
972:
968:
958:
954:
950:
946:
942:
938:
931:
924:
922:
918:
913:
909:
905:
901:
897:
893:
889:
885:
881:
874:
872:
870:
868:
864:
859:
855:
848:
841:
839:
837:
833:
830:
826:
820:
818:
816:
814:
812:
810:
806:
801:
797:
790:
788:
784:
779:
772:
770:
768:
766:
764:
762:
760:
756:
751:
747:
743:
739:
735:
731:
727:
723:
719:
712:
710:
708:
706:
702:
697:
693:
688:
683:
679:
678:
673:
666:
663:
658:
654:
650:
646:
641:
636:
631:
626:
622:
618:
614:
607:
604:
600:
596:
593:
588:
586:
582:
571:
567:
561:
558:
545:
540:
536:
532:
531:
526:
524:
515:
513:
509:
504:
500:
496:
494:0-801-88221-4
490:
486:
485:
480:
479:Wilson, D. E.
476:
475:Groves, C. P.
470:
467:
460:
458:
454:
450:
448:
443:
440:
436:
433:
429:
421:
419:
417:
413:
409:
401:
399:
397:
389:
387:
383:
380:
376:
368:
366:
364:
360:
356:
352:
348:
344:
340:
339:
330:
324:
320:
318:
314:
310:
306:
302:
298:
294:
290:
286:
282:
278:
274:
273:
268:
257:
253:
248:
243:
238:
236:
230:
227:
226:Binomial name
223:
219:
218:
213:
210:
209:
206:
205:
201:
198:
197:
194:
191:
188:
187:
184:
181:
178:
177:
174:
171:
168:
167:
164:
161:
158:
157:
154:
151:
148:
147:
144:
141:
138:
137:
134:
131:
128:
127:
124:
121:
118:
117:
112:
107:
103:
97:
91:
86:
80:
75:
64:
60:
55:
52:
47:
43:
38:
33:
30:
19:
2075:
2050:
2033:
2021:
2009:
1997:
1985:
1977:R. avunculus
1976:
1967:
1958:
1950:R. roxellana
1949:
1929:
1917:
1908:
1906:
1899:
1879:
1856:
1847:
1838:
1830:P. rubicunda
1829:
1820:
1811:
1802:
1793:
1784:
1775:
1767:P. siamensis
1766:
1757:
1748:
1740:P. robinsoni
1739:
1731:P. femoralis
1730:
1721:
1713:P. sumatrana
1712:
1703:
1694:
1674:
1662:
1654:T. delacouri
1653:
1644:
1635:
1626:
1617:
1609:T. francoisi
1608:
1601:T. francoisi
1600:
1599:
1590:
1581:
1572:
1564:
1563:
1554:
1545:
1536:
1527:
1519:
1518:
1509:
1500:
1492:T. margarita
1491:
1482:
1474:T. cristatus
1473:
1465:T. mauritius
1464:
1455:
1448:T. cristatus
1447:
1446:
1436:
1424:
1415:
1406:
1397:
1388:
1379:
1370:
1361:
1341:
1334:Langur group
1318:
1306:
1294:
1285:
1277:P. oustaleti
1276:
1267:
1258:
1249:
1240:
1231:
1222:
1213:
1204:
1195:
1186:
1177:
1169:P. pennantii
1168:
1160:P. waldronae
1159:
1150:
1139:(Red colobi)
1132:Piliocolobus
1130:
1118:
1109:
1101:C. polykomos
1100:
1091:
1082:
1062:
966:
940:
936:
890:(1): 15–28.
887:
883:
857:
853:
824:
799:
777:
725:
721:
675:
665:
620:
616:
606:
573:. Retrieved
569:
560:
548:. Retrieved
534:
528:
522:
483:
469:
455:
451:
444:
425:
422:Conservation
405:
393:
384:
378:
374:
372:
358:
337:
334:
317:habitat loss
271:
270:
266:
264:
234:
232:
216:
215:
203:
179:Infraorder:
29:
2193:iNaturalist
2100:Wikispecies
2034:S. concolor
2010:N. larvatus
1986:R. strykeri
1968:R. brelichi
1909:P. nigripes
1812:P. canicrus
1776:P. frontata
1573:T. pileatus
1565:T. pileatus
1528:T. obscurus
1520:T. obscurus
1483:T. germaini
1389:S. entellus
1250:P. bouvieri
1187:P. tholloni
1007:(subfamily
623:(2): e614.
550:19 November
447:Vietnam War
369:Description
183:Simiiformes
2332:Categories
1918:P. cinerea
1900:P. nemaeus
1857:P. natunae
1794:P. thomasi
1749:P. percura
1722:P. mitrata
1704:P. bicolor
1546:T. phayrei
1456:T. auratus
1416:S. vetulus
1308:Procolobus
1178:P. preussi
1119:C. guereza
1083:C. satanas
1046:Haplorhini
1044:Suborder:
640:1893/33780
575:2022-01-14
461:References
435:Endangered
428:Endangered
416:wet season
412:dry season
408:folivorous
297:endangered
173:Haplorhini
169:Suborder:
92:Appendix I
1881:Pygathrix
1848:P. siberu
1821:P. sabana
1785:P. comata
1683:(Surilis)
1676:Presbytis
1663:T. ebenus
1645:T. laotum
1501:T. barbei
1425:S. johnii
1380:S. hector
1241:P. epieni
1223:P. kirkii
1151:P. badius
1020:Kingdom:
1009:Colobinae
957:0217-2445
904:1573-8604
742:0164-0291
696:229677607
657:245405123
649:2578-4854
570:cites.org
390:Behaviour
379:Pygathrix
375:Pygathrix
359:Pygathrix
338:Pygathrix
211:Species:
204:Pygathrix
129:Kingdom:
123:Eukaryota
2310:Species+
2250:12100684
2085:Wikidata
2052:Category
1959:R. bieti
1803:P. hosei
1407:S. priam
1319:P. verus
1268:P. langi
1040:Primates
1034:Mammalia
1028:Chordata
1026:Phylum:
1022:Animalia
912:44304812
860:: 71–76.
750:17954281
595:Archived
503:62265494
477:(2005).
347:Cambodia
285:Cambodia
189:Family:
163:Primates
153:Mammalia
143:Chordata
139:Phylum:
133:Animalia
119:Domain:
79:IUCN 3.1
2237:1000681
2185:4267166
2172:4453746
2091:Q959360
1999:Nasalis
1888:(Doucs)
1591:T. geei
1555:T. popa
1371:S. ajax
1196:P. foai
1064:Colobus
1038:Order:
1032:Class:
430:by the
355:Vietnam
343:endemic
303:in the
289:Vietnam
277:species
199:Genus:
159:Order:
149:Class:
94: (
77: (
2302:232434
2289:473243
2276:201166
2263:310352
2211:944257
2146:493872
2128:ARKive
2023:Simias
955:
910:
902:
748:
740:
694:
655:
647:
501:
491:
353:, and
244:, 1871
2343:Doucs
2224:39828
2198:74992
2159:4QNHG
1603:group
1567:group
1522:group
1450:group
933:(PDF)
908:S2CID
850:(PDF)
746:S2CID
692:S2CID
653:S2CID
96:CITES
90:CITES
2315:5093
2258:NCBI
2219:IUCN
2206:ITIS
2180:GBIF
2141:BOLD
953:ISSN
900:ISSN
738:ISSN
645:ISSN
552:2021
535:2021
499:OCLC
489:ISBN
402:Diet
351:Laos
281:douc
265:The
2245:MSW
2232:MDD
2167:EoL
2154:CoL
2115:ADW
945:doi
892:doi
730:doi
682:doi
635:hdl
625:doi
539:doi
414:to
299:to
279:of
2334::
2312::
2299::
2286::
2273::
2260::
2247::
2234::
2221::
2208::
2195::
2182::
2169::
2156::
2143::
2130::
2117::
2102::
2087::
1605::
1569::
1524::
1452::
951:.
941:68
939:.
935:.
920:^
906:.
898:.
888:30
886:.
882:.
866:^
856:.
852:.
835:^
808:^
798:.
786:^
758:^
744:.
736:.
726:33
724:.
720:.
704:^
690:.
680:.
674:.
651:.
643:.
633:.
619:.
615:.
584:^
568:.
533:.
527:.
511:^
497:.
349:,
2036:)
2012:)
1988:)
1979:)
1970:)
1961:)
1952:)
1920:)
1911:)
1902:)
1859:)
1850:)
1841:)
1832:)
1823:)
1814:)
1805:)
1796:)
1787:)
1778:)
1769:)
1760:)
1751:)
1742:)
1733:)
1724:)
1715:)
1706:)
1697:)
1665:)
1656:)
1647:)
1638:)
1629:)
1620:)
1611:)
1593:)
1584:)
1575:)
1557:)
1548:)
1539:)
1530:)
1512:)
1503:)
1494:)
1485:)
1476:)
1467:)
1458:)
1427:)
1418:)
1409:)
1400:)
1391:)
1382:)
1373:)
1364:)
1321:)
1297:)
1288:)
1279:)
1270:)
1261:)
1252:)
1243:)
1234:)
1225:)
1216:)
1207:)
1198:)
1189:)
1180:)
1171:)
1162:)
1153:)
1121:)
1112:)
1103:)
1094:)
1085:)
1011:)
993:e
986:t
979:v
959:.
947::
914:.
894::
858:2
752:.
732::
698:.
684::
659:.
637::
627::
621:4
578:.
554:.
541::
525:"
521:"
505:.
269:(
98:)
81:)
20:)
Text is available under the Creative Commons Attribution-ShareAlike License. Additional terms may apply.
