518:
53:
31:
211:, most members of which lack permanently air-filled lungs as well. To emphasize the complete change in taxonomic concepts following from the molecular phylogenetic analyses but keep some continuity in the naming, Jörger et al. (2010) renamed the expanded "Pulmonata" as Panpulmonata
191:" grouped various, mostly marine euthyneuran lineages, often with reduced shell. Phylogenetic analyses have shown that none of these two groups was monophyletic. The former "Pulmonata" should also include several groups that lack an air-filled lung (
1213:
Jörger K. M., Stöger I., Kano Y., Fukuda H., Knebelsberger T. & Schrödl M. (2010). "On the origin of
Acochlidia and other enigmatic euthyneuran gastropods, with implications for the systematics of Heterobranchia".
1267:
Bouchet, Philippe; Rocroi, Jean-Pierre; Hausdorf, Bernhard; Kaim, Andrzej; Kano, Yasunori; Nützel, Alexander; Parkhaev, Pavel; Schrödl, Michael; Strong, Ellen E. (2017).
1166:
1378:
529:. The reason given is: Newer phylogenomic analyses resolve some of the branches differently, most importantly Siphonarioidea.
52:
1365:
1383:
1216:
919:
424:
208:
203:). The name "Pulmonata" also inaccurate when applied to some of the more traditional pulmonate taxa such as
1318:
1419:
1242:
573:
336:
261:
1298:
988:
497:
467:
47:
1269:"Revised Classification, Nomenclator and Typification of Gastropod and Monoplacophoran Families"
1356:
1391:
1290:
626:
1396:
1282:
1224:
39:
552:
shows phylogenic relations within the
Heterobranchia, as proposed by Jörger et al. (2010):
966:
879:
493:
379:
255:
200:
188:
531:
Please help update this article to reflect recent events or newly available information.
1171:
1047:
842:
789:
559:
411:
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384:
290:
279:
204:
162:
104:
1413:
1176:
941:
864:
738:
639:
416:
403:
371:
348:
324:
319:
311:
300:
267:
244:
232:
192:
1370:
1302:
365:
342:
305:
1268:
1350:
1273:
1010:
953:
577:
489:
479:
455:
390:
124:
804:
668:
607:
594:
449:
432:
354:
330:
273:
218:
196:
184:
166:
114:
94:
1341:
1294:
1228:
1032:
705:
690:
549:
501:
461:
226:
180:
64:
1335:
1286:
473:
438:
238:
84:
187:
gastropods which were considered to be "air-breathers". In contrast, "
154:
74:
1312:
30:
173:
158:
150:
1316:
1167:
Changes in the taxonomy of gastropods since 2005#Heterobranchia
511:
1187:
This article incorporates CC-BY-2.0 text from the reference.
584:
considered a clade in the study by Jörger et al. (2010).
339:
Neusser, Jörger, Lodde-Bensch, Strong & Schrödl, 2016
1325:
287:Superfamily Platyhedyloidea Salvini-Plawen, 1973
1209:
1207:
1205:
1203:
1201:
1199:
1197:
1195:
8:
1313:
29:
20:
446:Superfamily Lymnaeoidea Rafinesque, 1815
252:Superfamily Plakobranchoidea Gray, 1840
1191:
223:Superfamily Oxynooidea Stoliczka, 1868
214:The clade consists of following taxa:
172:Panpulmonata was established as a new
7:
1025:
1003:
981:
959:
934:
912:
857:
835:
828:
782:
775:
683:
661:
654:
632:
622:
600:
565:
555:
429:Superfamily Chilinoidea Dall, 1870
1243:"Time to say "Bye-bye Pulmonata"?"
176:by Jörger et al. in October 2010.
14:
314:Golding, Ponder & Byrne, 2007
516:
51:
1:
464:P. Fischer & Crosse, 1880
276:H. Adams & A. Adams, 1854
1172:Heterobranchia#2010 taxonomy
580:) - Lower Heterobranchia is
1177:Acochlidiacea#2010 taxonomy
362:Parhedyloidea Thiele, 1931
1436:
333:Swennen & Buatip, 2009
1045:
1030:
1023:
1008:
1001:
986:
979:
964:
957:
939:
932:
917:
910:
877:
862:
855:
840:
833:
826:
802:
787:
780:
773:
736:
703:
688:
681:
666:
659:
652:
637:
630:
620:
605:
598:
570:
563:
525:This section needs to be
183:" referred to a group of
48:Scientific classification
46:
37:
28:
23:
1241:Schödl, Michael (2014).
1229:10.1186/1471-2148-10-323
1217:BMC Evolutionary Biology
264:Ortea & Moro, 2015
1287:10.4002/040.061.0201
574:Lower Heterobranchia
293:Salvini-Plawen, 1973
139:Jörger et al., 2010
16:Clade of gastropods
989:Systellommatophora
498:Systellommatophora
1407:
1406:
1392:Open Tree of Life
1319:Taxon identifiers
1157:
1156:
1148:
1147:
1139:
1138:
1130:
1129:
1121:
1120:
1112:
1111:
1103:
1102:
1094:
1093:
1085:
1084:
1076:
1075:
1067:
1066:
1058:
1057:
899:
898:
890:
889:
815:
814:
759:
758:
750:
749:
725:
724:
716:
715:
627:Euopisthobranchia
546:
545:
282:K. B. Clark, 1984
241:E. A. Smith, 1885
165:within the clade
144:
143:
140:
1427:
1400:
1399:
1387:
1386:
1374:
1373:
1361:
1360:
1359:
1346:
1345:
1344:
1314:
1307:
1306:
1264:
1258:
1257:
1247:
1238:
1232:
1211:
1026:
1004:
982:
960:
935:
913:
858:
836:
829:
783:
776:
684:
662:
655:
633:
623:
601:
566:
556:
541:
538:
532:
520:
519:
512:
482:Rafinesque, 1815
452:Rafinesque, 1815
233:Cylindrobullidae
179:The older name "
138:
56:
55:
40:Cepaea nemoralis
33:
21:
1435:
1434:
1430:
1429:
1428:
1426:
1425:
1424:
1410:
1409:
1408:
1403:
1395:
1390:
1382:
1377:
1369:
1364:
1355:
1354:
1349:
1340:
1339:
1334:
1321:
1311:
1310:
1266:
1265:
1261:
1245:
1240:
1239:
1235:
1212:
1193:
1185:
1163:
1158:
1149:
1140:
1131:
1122:
1113:
1104:
1095:
1086:
1077:
1068:
1059:
967:Stylommatophora
900:
891:
880:Pyramidelloidea
816:
760:
751:
726:
717:
542:
536:
533:
530:
521:
517:
510:
494:Stylommatophora
476:Fitzinger, 1833
380:Pyramidelloidea
256:Plakobranchidae
229:Stoliczka, 1868
201:Pyramidelloidea
149:is a taxonomic
137:
50:
17:
12:
11:
5:
1433:
1431:
1423:
1422:
1412:
1411:
1405:
1404:
1402:
1401:
1388:
1375:
1362:
1347:
1331:
1329:
1323:
1322:
1317:
1309:
1308:
1281:(1–2): 1–526.
1259:
1233:
1190:
1189:
1184:
1181:
1180:
1179:
1174:
1169:
1162:
1159:
1155:
1154:
1151:
1150:
1146:
1145:
1142:
1141:
1137:
1136:
1133:
1132:
1128:
1127:
1124:
1123:
1119:
1118:
1115:
1114:
1110:
1109:
1106:
1105:
1101:
1100:
1097:
1096:
1092:
1091:
1088:
1087:
1083:
1082:
1079:
1078:
1074:
1073:
1070:
1069:
1065:
1064:
1061:
1060:
1056:
1055:
1052:
1051:
1048:Trimusculoidea
1044:
1041:
1040:
1037:
1036:
1029:
1024:
1022:
1019:
1018:
1015:
1014:
1007:
1002:
1000:
997:
996:
993:
992:
985:
980:
978:
975:
974:
971:
970:
963:
958:
956:
950:
949:
946:
945:
938:
933:
931:
928:
927:
924:
923:
916:
911:
909:
906:
905:
902:
901:
897:
896:
893:
892:
888:
887:
884:
883:
876:
873:
872:
869:
868:
861:
856:
854:
851:
850:
847:
846:
843:Glacidorboidea
839:
834:
832:
827:
825:
822:
821:
818:
817:
813:
812:
809:
808:
801:
798:
797:
794:
793:
790:Siphonarioidea
786:
781:
779:
774:
772:
766:
765:
762:
761:
757:
756:
753:
752:
748:
747:
744:
743:
735:
732:
731:
728:
727:
723:
722:
719:
718:
714:
713:
710:
709:
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695:
694:
687:
682:
680:
677:
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673:
672:
665:
660:
658:
653:
651:
648:
647:
644:
643:
636:
631:
629:
621:
619:
616:
615:
612:
611:
604:
599:
597:
591:
590:
587:
586:
569:
564:
562:
560:Heterobranchia
554:
544:
543:
524:
522:
515:
509:
506:
505:
504:
487:
486:
485:
484:
483:
477:
471:
470:Albrecht, 2017
465:
459:
453:
444:
443:
442:
436:
422:
421:
420:
412:Siphonarioidea
409:
408:
407:
399:Glacidorboidea
396:
395:
394:
388:
385:Pyramidellidae
377:
376:
375:
369:
360:
359:
358:
352:
346:
340:
337:Bathyhedylidae
334:
328:
317:
316:
315:
309:
298:
297:
296:
295:
294:
291:Platyhedylidae
285:
284:
283:
280:Costasiellidae
277:
271:
265:
259:
250:
249:
248:
242:
236:
230:
205:Siphonarioidea
189:Opistobranchia
163:Heterobranchia
142:
141:
132:
128:
127:
122:
118:
117:
112:
108:
107:
105:Heterobranchia
102:
98:
97:
92:
88:
87:
82:
78:
77:
72:
68:
67:
62:
58:
57:
44:
43:
35:
34:
26:
25:
15:
13:
10:
9:
6:
4:
3:
2:
1432:
1421:
1418:
1417:
1415:
1398:
1393:
1389:
1385:
1380:
1376:
1372:
1367:
1363:
1358:
1352:
1348:
1343:
1337:
1333:
1332:
1330:
1328:
1324:
1320:
1315:
1304:
1300:
1296:
1292:
1288:
1284:
1280:
1276:
1275:
1270:
1263:
1260:
1255:
1251:
1244:
1237:
1234:
1230:
1226:
1222:
1219:
1218:
1210:
1208:
1206:
1204:
1202:
1200:
1198:
1196:
1192:
1188:
1182:
1178:
1175:
1173:
1170:
1168:
1165:
1164:
1160:
1153:
1152:
1144:
1143:
1135:
1134:
1126:
1125:
1117:
1116:
1108:
1107:
1099:
1098:
1090:
1089:
1081:
1080:
1072:
1071:
1063:
1062:
1054:
1053:
1050:
1049:
1043:
1042:
1039:
1038:
1035:
1034:
1028:
1027:
1021:
1020:
1017:
1016:
1013:
1012:
1006:
1005:
999:
998:
995:
994:
991:
990:
984:
983:
977:
976:
973:
972:
969:
968:
962:
961:
955:
952:
951:
948:
947:
944:
943:
942:Acochlidiacea
937:
936:
930:
929:
926:
925:
922:
921:
915:
914:
908:
907:
904:
903:
895:
894:
886:
885:
882:
881:
875:
874:
871:
870:
867:
866:
865:Amphiboloidea
860:
859:
853:
852:
849:
848:
845:
844:
838:
837:
831:
830:
824:
823:
820:
819:
811:
810:
807:
806:
800:
799:
796:
795:
792:
791:
785:
784:
778:
777:
771:
768:
767:
764:
763:
755:
754:
746:
745:
742:
740:
739:Cephalaspidea
734:
733:
730:
729:
721:
720:
712:
711:
708:
707:
701:
700:
697:
696:
693:
692:
686:
685:
679:
678:
675:
674:
671:
670:
664:
663:
657:
656:
650:
649:
646:
645:
642:
641:
640:Umbraculoidea
635:
634:
628:
625:
624:
618:
617:
614:
613:
610:
609:
603:
602:
596:
593:
592:
589:
588:
585:
583:
579:
576:" (including
575:
568:
567:
561:
558:
557:
553:
551:
540:
528:
523:
514:
513:
507:
503:
499:
495:
491:
488:
481:
478:
475:
472:
469:
466:
463:
460:
457:
454:
451:
448:
447:
445:
440:
437:
434:
431:
430:
428:
427:
426:
423:
418:
417:Siphonariidae
415:
414:
413:
410:
405:
404:Glacidorbidae
402:
401:
400:
397:
392:
389:
386:
383:
382:
381:
378:
373:
372:Asperspinidae
370:
367:
364:
363:
361:
356:
353:
350:
349:Pseudunelidae
347:
344:
341:
338:
335:
332:
329:
326:
325:Acochlidiidae
323:
322:
321:
320:Acochlidiacea
318:
313:
312:Maningrididae
310:
307:
304:
303:
302:
301:Amphiboloidea
299:
292:
289:
288:
286:
281:
278:
275:
272:
269:
268:Limapontiidae
266:
263:
262:Jenseneriidae
260:
257:
254:
253:
251:
247:Pilsbry, 1895
246:
245:Volvatellidae
243:
240:
237:
234:
231:
228:
225:
224:
222:
221:
220:
217:
216:
215:
212:
210:
206:
202:
198:
194:
193:Acochlidiacea
190:
186:
182:
177:
175:
170:
168:
164:
161:in the clade
160:
156:
152:
148:
136:
133:
130:
129:
126:
123:
120:
119:
116:
113:
110:
109:
106:
103:
100:
99:
96:
93:
90:
89:
86:
83:
80:
79:
76:
73:
70:
69:
66:
63:
60:
59:
54:
49:
45:
42:
41:
36:
32:
27:
24:Panpulmonata
22:
19:
1420:Panpulmonata
1371:Panpulmonata
1357:Panpulmonata
1327:Panpulmonata
1326:
1278:
1272:
1262:
1253:
1249:
1236:
1220:
1215:
1186:
1046:
1031:
1009:
987:
965:
940:
918:
878:
863:
841:
803:
788:
770:Panpulmonata
769:
737:
704:
689:
667:
638:
606:
581:
571:
547:
534:
526:
458:Thiele, 1931
441:Hutton, 1882
406:Ponder, 1986
393:Ponder, 1987
374:Rankin, 1979
368:Thiele, 1931
366:Parhedylidae
357:Rankin, 1979
351:Rankin, 1979
345:Odhner, 1952
343:Hedylopsidae
306:Amphibolidae
235:Thiele, 1931
213:
178:
171:
147:Panpulmonata
146:
145:
135:Panpulmonata
134:
111:Infraclass:
38:
18:
1351:Wikispecies
1274:Malacologia
1011:Ellobioidea
954:Eupulmonata
578:Acteonoidea
537:August 2023
490:Eupulmonata
480:Planorbidae
468:Burnupiidae
456:Acroloxidae
391:Amathinidae
327:Küthe, 1935
185:euthyneuran
131:Subcohort:
125:Tectipleura
1256:: 161–164.
1183:References
920:Hygrophila
805:Sacoglossa
669:Runcinacea
608:Nudipleura
595:Euthyneura
450:Lymnaeidae
435:Dall, 1870
433:Chilinidae
425:Hygrophila
419:Gray, 1827
387:Gray, 1840
355:Tantulidae
331:Aitengidae
308:Gray, 1840
274:Hermaeidae
270:Gray, 1847
258:Gray, 1840
219:Sacoglossa
209:Hygrophila
197:Sacoglossa
167:Euthyneura
115:Euthyneura
101:Subclass:
95:Gastropoda
1295:0076-2997
1033:Otinoidea
706:Pteropoda
691:Anaspidea
550:cladogram
508:Cladogram
502:Ellobiida
462:Bulinidae
227:Oxynoidae
181:Pulmonata
71:Kingdom:
65:Eukaryota
1414:Category
1336:Wikidata
1303:91051256
1250:Spixiana
1161:See also
474:Physidae
439:Latiidae
239:Juliidae
121:Cohort:
85:Mollusca
81:Phylum:
75:Animalia
61:Domain:
1342:Q870378
1223:: 323.
527:updated
91:Class:
1397:933373
1384:977775
1301:
1293:
155:snails
1299:S2CID
1246:(PDF)
741:s.s.
548:This
174:taxon
159:slugs
151:clade
1379:NCBI
1291:ISSN
199:and
157:and
1366:AFD
1283:doi
1225:doi
582:not
207:or
153:of
1416::
1394::
1381::
1368::
1353::
1338::
1297:.
1289:.
1279:61
1277:.
1271:.
1254:37
1252:.
1248:.
1221:10
1194:^
500:,
496:,
492::
195:,
169:.
1305:.
1285::
1231:.
1227::
572:"
539:)
535:(
Text is available under the Creative Commons Attribution-ShareAlike License. Additional terms may apply.
