46:
33:
371:
594:
251:(hairs). The inside of the claw of the crab is smooth, although many ridges and fine granules are present on the movable portion; these ridges and granules are smaller in female and
460:
473:
275:
P. L. Mfilinge & M. Tsuchiya (2008). "Effect of temperature on leaf litter consumption by grapsid crabs in a subtropical mangrove (Okinawa, Japan)".
659:
307:
K. K. Pillay & Y. Ono (1978). "The breeding cycles of two species of grapsid crabs (Crustacea: Decapoda) from the North Coast of Kyushu, Japan".
566:
434:
664:
669:
486:
635:
525:
375:
337:
45:
478:
309:
277:
628:
389:
160:
530:
674:
654:
252:
517:
621:
137:
40:
201:
465:
499:
421:
504:
318:
286:
256:
213:
605:
177:
32:
648:
240:
557:
426:
87:
290:
512:
447:
107:
412:
197:
127:
77:
57:
370:
551:
406:
259:) of the walking legs of this crab are broad relative to the lower limb.
244:
236:
209:
97:
571:
439:
322:
221:
217:
593:
452:
67:
383:
224:. It eats leaf litter and other vegetation. The breeding season of
220:, and living in small crevices and abandoned holes made by other
601:
491:
248:
205:
193:
117:
387:
609:
541:
396:
247:that wear down as the crab ages, and some short
629:
8:
302:
300:
255:crabs than in adult males. The top portion (
636:
622:
384:
31:
20:
267:
348:. Marine Species Identification Portal
7:
590:
588:
14:
660:Crustaceans of the Pacific Ocean
592:
369:
44:
208:. This crab typically inhabits
228:is between May and September.
1:
665:Crustaceans described in 1833
212:swamps, preferring the upper
670:Taxa named by Wilhem de Haan
608:. You can help Knowledge by
291:10.1016/j.seares.2007.07.004
204:), which is endemic to East
231:Like other sesarmid crabs,
691:
587:
166:
159:
41:Scientific classification
39:
30:
23:
278:Journal of Sea Research
243:are covered with scaly
380:at Wikimedia Commons
196:, belonging to the
398:Parasesarma pictum
377:Parasesarma pictum
340:Parasesarma pictum
323:10.1007/BF00390606
200:family (subfamily
189:Parasesarma pictum
170:Parasesarma pictum
25:Parasesarma pictum
617:
616:
582:
581:
500:Open Tree of Life
390:Taxon identifiers
374:Media related to
214:intertidal region
185:
184:
682:
638:
631:
624:
596:
589:
575:
574:
562:
561:
560:
534:
533:
521:
520:
508:
507:
495:
494:
482:
481:
469:
468:
456:
455:
443:
442:
430:
429:
417:
416:
415:
385:
373:
358:
357:
355:
353:
333:
327:
326:
304:
295:
294:
272:
172:
49:
48:
35:
21:
690:
689:
685:
684:
683:
681:
680:
679:
645:
644:
643:
642:
585:
583:
578:
570:
565:
556:
555:
550:
537:
529:
524:
516:
511:
503:
498:
490:
485:
477:
472:
464:
459:
451:
446:
438:
433:
425:
420:
411:
410:
405:
392:
366:
361:
351:
349:
335:
334:
330:
306:
305:
298:
285:(1–2): 94–102.
274:
273:
269:
265:
181:
174:
168:
155:
43:
17:
16:Species of crab
12:
11:
5:
688:
686:
678:
677:
672:
667:
662:
657:
647:
646:
641:
640:
633:
626:
618:
615:
614:
597:
580:
579:
577:
576:
563:
547:
545:
543:Grapsus pictum
539:
538:
536:
535:
522:
509:
496:
483:
470:
457:
444:
431:
418:
402:
400:
394:
393:
388:
382:
381:
365:
364:External links
362:
360:
359:
346:Crabs of Japan
328:
317:(3): 237–248.
310:Marine Biology
296:
266:
264:
261:
233:P. pictum
226:P. pictum
183:
182:
175:
164:
163:
157:
156:
152:P. pictum
149:
147:
143:
142:
135:
131:
130:
125:
121:
120:
115:
111:
110:
105:
101:
100:
95:
91:
90:
85:
81:
80:
75:
71:
70:
65:
61:
60:
55:
51:
50:
37:
36:
28:
27:
15:
13:
10:
9:
6:
4:
3:
2:
687:
676:
673:
671:
668:
666:
663:
661:
658:
656:
653:
652:
650:
639:
634:
632:
627:
625:
620:
619:
613:
611:
607:
604:article is a
603:
598:
595:
591:
586:
573:
568:
564:
559:
553:
549:
548:
546:
544:
540:
532:
527:
523:
519:
514:
510:
506:
501:
497:
493:
488:
484:
480:
475:
471:
467:
462:
458:
454:
449:
445:
441:
436:
432:
428:
423:
419:
414:
408:
404:
403:
401:
399:
395:
391:
386:
379:
378:
372:
368:
367:
363:
347:
343:
341:
332:
329:
324:
320:
316:
312:
311:
303:
301:
297:
292:
288:
284:
280:
279:
271:
268:
262:
260:
258:
254:
250:
246:
242:
238:
235:has a square
234:
229:
227:
223:
219:
215:
211:
207:
203:
199:
195:
192:is a mudflat
191:
190:
179:
173:
171:
165:
162:
161:Binomial name
158:
154:
153:
148:
145:
144:
141:
140:
136:
133:
132:
129:
126:
123:
122:
119:
116:
113:
112:
109:
106:
103:
102:
99:
96:
93:
92:
89:
86:
83:
82:
79:
76:
73:
72:
69:
66:
63:
62:
59:
56:
53:
52:
47:
42:
38:
34:
29:
26:
22:
19:
610:expanding it
599:
584:
542:
397:
376:
350:. Retrieved
345:
339:
331:
314:
308:
282:
276:
270:
232:
230:
225:
188:
187:
186:
169:
167:
151:
150:
138:
114:Infraorder:
88:Malacostraca
24:
18:
513:SeaLifeBase
448:iNaturalist
139:Parasesarma
108:Pleocyemata
675:Crab stubs
655:Sesarmidae
649:Categories
558:Q109655217
352:19 January
336:K. Sakai.
263:References
202:Sesarminae
198:Sesarmidae
128:Sesarmidae
104:Suborder:
78:Arthropoda
245:tubercles
218:estuaries
146:Species:
118:Brachyura
64:Kingdom:
58:Eukaryota
552:Wikidata
466:11708370
413:Q6618961
407:Wikidata
253:juvenile
237:carapace
210:mangrove
124:Family:
98:Decapoda
74:Phylum:
68:Animalia
54:Domain:
572:5974921
505:2981211
479:1904478
440:5974920
222:species
180:, 1833)
178:De Haan
134:Genus:
94:Order:
84:Class:
531:444637
518:172269
492:444637
453:496429
241:chelae
239:. Its
600:This
526:WoRMS
461:IRMNG
427:4DBZH
257:merus
249:setae
606:stub
602:crab
567:GBIF
487:OBIS
474:NCBI
435:GBIF
354:2012
206:Asia
194:crab
422:CoL
319:doi
287:doi
216:of
651::
569::
554::
528::
515::
502::
489::
476::
463::
450::
437::
424::
409::
344:.
315:45
313:.
299:^
283:59
281:.
637:e
630:t
623:v
612:.
356:.
342:"
338:"
325:.
321::
293:.
289::
176:(
Text is available under the Creative Commons Attribution-ShareAlike License. Additional terms may apply.