225:
316:, especially when standing or walking. The fore feet have four digits while the hind feet have three digits. Its tail is short, depressed, and hairless. It has a gray dorsal pelage with a white patch on the rump separated from the dorsal fur by a black area. In addition, the mara has a white underside with a somewhat orange flank and chin. The average Patagonian mara has a head and body length of 69–75 cm (27–30 in) with a tail of 4–5 cm (1.6–2.0 in). It weighs 8–16 kg (18–35 lb). Unlike most other
50:
528:
time for around one hour and the other parents circle around the den. One or two pups are nursed at a time by a female. A female may sometimes nurse a young from another pair. While a female may prevent young other than her own from nursing her, some young are able to steal milk. Mothers do not actively cooperate in raising their young. For the first three weeks, young remain near the den. At this time, distance between individuals is low, with frequent body contact, huddling,
500:
539:. Maras tooth chatter and emit low grunts when threatened. They also produce a series of short grunts when grooming. Scent marking is used by maras for complex and intense social interactions. Maras stretch and sniff the soil and then sit upright with an arched back and the anogenital area flattened to the ground, a process known as anal digging. In addition, a male stands on his hind legs and urinates on a female's rump, to which the female responds by
103:
78:
479:
512:
556:
341:
491:
shared by up to 29 pairs. Maras breed, at least in southern
Argentina, from August to January. Gestation lasts 100 days in wild. Most births in Patagonia occur between September and October, which is before the summer dry season and after the winter rains. Females produce one litter each year in the
438:
and around 46% of their daily activities involve feeding. Their temporal activity rhythms are related to environmental factors. Light, precipitation, and temperature have positive effects on annual activities, while darkness and relative humidity have negative effects. The daily activity of maras is
527:
Dens are dug during the breeding season for the young to be raised. Litters from one to 22 pairs are grouped together in these dens. Communal living provides protection from predators with the survival rate for young being higher in larger groups than in smaller groups. One pair visits the den at a
486:
Their social organizations have a unique combination of monogamy and communal breeding. Being monogamous, pairs of maras stay together for life, with replacement of partners only occurring after one's death. The male has almost the sole responsibility in maintaining the pair by following the female
443:
in winter and biomodal in other seasons. Their preferred temperature is around 20 °C or 68 °F. Females spend more time feeding than the males, due to the demands of gestation and lactation. Males spend most of the day sitting and being vigilant for predators. Predators of maras,
487:
wherever she goes. A male marks his female with urine and marks the ground around her with secretions from his glands and with feces, making the grounds around the female a mobile territory. Pairs breed together alone or with other pairs in
887:
813:
1826:
535:
Maras make a number of vocalizations during grazing or slow locomotion. When seeking contact, a mara emits an inflected "wheet" while a low, repetitive grunt is made when following a
543:
into the face of the male. The male's urination is meant to repel other males from his partner, while the female's urination is a rejection of any approaching male when she is not
1592:
1801:
1670:
378:, and well-developed sensory organs making them capable of running and communicating in these open habitats. When running, maras have been compared to deer and
648:
IUCN (International Union for
Conservation of Nature) 2008. Dolichotis patagonum. In: IUCN 2015. The IUCN Red List of Threatened Species. Version 2015.2.
1760:
1566:
532:, and extended play among the pups. After this, the young are able to leave the den and graze with their parents. Young are weaned after 13 weeks.
1618:
1044:
214:
681:
1017:
1821:
304:. It has distinctive long ears and long limbs. Its hind limbs are longer and more muscular than its fore limbs and it has a longer
1644:
966:
Rood. J. P. (1972) Ecological and behavioral comparisons of three genera of
Argentine cavies, Animal Behavior Monographs 5:1-83.
224:
1662:
1623:
1146:
1203:
1816:
1811:
1235:
1657:
1605:
1219:
536:
663:
547:. Both anal digging and urination are more frequent during the breeding season and are more commonly done by males.
102:
1282:
1227:
1037:
540:
1675:
1508:
49:
1778:
356:. They prefer to live in habitats with shrub cover, but they also inhabit overgrazed and barren soils in the
830:
Kufner. M. B. (1995) "Temporal activity of the mara (Dolichotis patagonum) in the Monte Desert, Argentina".
949:
TABER, B. E., AND D. W. MACDONALD. (1984) "Scent dispersing papillae and associated behaviour in the mara,
1470:
1266:
466:
374:. They have adapted well to a cursorial lifestyle on the open plains and steppe, with long legs, reduced
312:. The feet are compressed, making them hoof-like, which contributes to Patagonian maras resembling small
1709:
1030:
580:
197:
1018:
https://web.archive.org/web/20120905133613/http://nationalzoo.si.edu/Animals/SmallMammals/fact-mara.cfm
910:
1553:
1517:
1274:
1130:
992:
983:), food availability and effects of an extended drought in Northern Patagonia (Mendoza, Argentina)".
851:
816:. Dolichotis patagonum. Ph. D. Thesis, Belliol College, University of Oxford, Oxford, United Kingdom.
492:
wild, but can produce as many as four litters a year in captivity. Young can walk almost immediately
285:
1211:
353:
349:
289:
67:
627:
499:
332:
of the mara are between the anus and the base of the tail rather than being anterior to the anus.
1170:
1154:
575:. Nevertheless, maras have been greatly affected by hunting and habitat alteration and have been
544:
504:
97:
1714:
560:
1597:
933:
Eisenberg, J. F. (1974). "The function and motivational basis of hystricomorph vocalizations".
1696:
1540:
687:
677:
1522:
1022:
671:
360:
biome. In northwestern
Argentina, they primarily inhabit lowland habitats such as forest and
1724:
1701:
1138:
1000:
979:
Puig, Silvia; Cona, Monica I.; Videla, Fernando; Mendez, Eduardo (2010). "Diet of the mara (
622:
576:
572:
371:
708:
1806:
1683:
1636:
1454:
1396:
1316:
568:
529:
488:
396:
82:
996:
1435:
1369:
1093:
571:
species. Historically, maras have ranged from north-central
Argentina south almost to
1795:
1631:
1299:
1184:
667:
613:
457:
387:
305:
253:
87:
604:
1751:
1545:
1379:
1352:
1340:
357:
1004:
1649:
1579:
1502:
516:
503:
Young mara and older one sniffing each other, from the zoological garden of the
440:
402:
391:
383:
269:
58:
39:
478:
17:
1249:
1162:
511:
493:
435:
408:
329:
321:
301:
174:
1493:
776:
Claudia M. Campos and
Ricardo A. Ojeda, (1997) "Dispersal and germination of
1427:
691:
281:
277:
114:
1610:
747:
Campos. C. M., Tognelli. M. F., Ojeda. R. A., (2001) Dolichotis patagonu,
386:. They feed primarily on green vegetation and fruit. In the Monte Desert,
1745:
1688:
1532:
1487:
1388:
1103:
1075:
1054:
461:
449:
427:
414:
379:
375:
361:
325:
317:
313:
164:
134:
911:"Juvenile development as part of the extraordinary social system of the
555:
1773:
1765:
1571:
1410:
520:
309:
1584:
1087:
1081:
1069:
445:
421:
366:
249:
154:
144:
124:
1464:
394:
make up 30%. Preferred grass species eaten are those of the genera
340:
1113:
554:
510:
477:
673:
Mammal
Species of the World: A Taxonomic and Geographic Reference
288:, possibly as a result of escaped pets or captive animals. It is
1558:
273:
1468:
1026:
676:(3rd ed.). Johns Hopkins University Press. p. 1555.
598:
596:
453:
649:
709:"On the existence of pursuit invitation signals in mammals"
292:, but often breeds in warrens shared by several pairs.
276:-like animal is found in open and semiopen habitats in
284:. A population has also been recorded in the northern
583:. Mara skins have been used for bedspreads and rugs.
1827:
Taxa named by
Eberhard August Wilhelm von Zimmermann
1735:
1477:
1408:
1377:
1368:
1338:
1314:
1298:
1247:
1182:
1111:
1102:
348:Patagonian maras are found only in Argentina, from
780:(Fabaceae) seeds by desert mammals in Argentina",
905:
903:
460:. Maras are also hosts for parasites such as the
929:
927:
370:. Maras prefer sandy and low shrub habitat in
1038:
882:
880:
878:
876:
874:
872:
870:
868:
772:
770:
743:
741:
628:10.2305/IUCN.UK.2016-2.RLTS.T6785A22190337.en
8:
945:
943:
846:
844:
842:
840:
832:Studies on Neotropical Fauna and Environment
826:
824:
822:
762:Mammals of the Neotropics: The Southern Cone
760:Redford, K. H., and J. F. Eisenberg, (1992)
739:
737:
735:
733:
731:
729:
727:
725:
723:
721:
703:
701:
808:
806:
804:
802:
800:
798:
796:
794:
792:
790:
1465:
1374:
1311:
1108:
1045:
1031:
1023:
850:Taber, A. B. and Macdonald, D. W. (1992),
223:
76:
48:
31:
626:
567:Patagonian maras are considered to be a
498:
339:
592:
1204:Southern highland yellow-toothed cavy
662:Woods, C.A.; Kilpatrick, C.W. (2005).
412:, while dicots that are eaten include
1802:IUCN Red List near threatened species
814:"The Behavioural Ecology of the Mara"
7:
1779:E177503B-5EB6-4375-B47B-0DA85858C88B
886:Genest, H. & Dubost, G. (1974)
614:IUCN Red List of Threatened Species
25:
909:U. GANSLOSSER, S. WEHNELT (1997)
541:spraying a jet of urine backwards
390:make up 70% of their diet, while
852:"Communal breeding in the mara,
474:Social behavior and reproduction
300:The Patagonian mara resembles a
101:
764:, University of Chicago Press.
1:
652:. Downloaded on 09 July 2015.
1220:Lowland yellow-toothed cavy
1147:Santa Catarina´s guinea pig
1005:10.1016/j.mambio.2009.12.003
782:Journal of Arid Environments
559:A Patagonian mara family in
444:particularly the young, are
1391:(Hydrochoerus hydrochaeris)
1236:Spix's yellow-toothed cavy
1228:Common yellow-toothed cavy
953:(Rodentia: Caviomorpha) ".
664:"Infraorder Hystricognathi"
280:, including large parts of
1843:
888:"Pair living in the mara (
650:http://www.iucnredlist.org
260:. It is also known as the
1822:Mammals described in 1780
1451:
1064:
1053:Extant species of family
913:Mara Dolichotis patagonum
579:in some areas, including
231:
222:
203:
196:
98:Scientific classification
96:
74:
65:
56:
47:
34:
1283:Shipton's mountain cavy
248:) is a relatively large
1399:(Hydrochoerus isthmius)
1355:(Pediolagus salinicola)
1267:Southern mountain cavy
1329:(Dolichotis patagonum)
1269:(Microcavia australis)
915:(Rodentia: Caviidae) "
890:Dolichotis paragonum Z
670:; Reeder, D.M (eds.).
564:
524:
508:
483:
345:
232:Patagonian mara range
1710:Paleobiology Database
1285:(Microcavia shiptoni)
1275:Andean mountain cavy
1222:(Galea leucoblephara)
1131:Brazilian guinea pig
937:. London 34, 211-248.
581:Buenos Aires Province
558:
514:
502:
481:
343:
1817:Mammals of Argentina
1812:Mammals of Patagonia
1523:Dolichotis_patagonum
1509:Dolichotis patagonum
1479:Dolichotis patagonum
1212:Yellow-toothed cavy
981:Dolichotis patagonum
951:Dolichotis patagonum
854:Dolichotis patagonum
621:: e.T6785A22190337.
607:Dolichotis patagonum
482:Mara pair with young
467:Wellcomia dolichotis
434:Maras are primarily
382:. Maras are largely
336:Ecology and activity
286:United Arab Emirates
245:Dolichotis patagonum
207:Dolichotis patagonum
1430:(Kerodon rupestris)
1230:(Galea musteloides)
1171:Montane guinea pig
1155:Greater guinea pig
997:2010MamBi..75..389P
713:American Naturalist
707:Smythe. N., (1970)
563:- note pup nursing.
68:Conservation status
1438:(Kerodon acrobata)
1277:(Microcavia niata)
1149:(Cavia intermedia)
955:Journal of Zoology
860:Journal of Zoology
603:Roach, N. (2016).
565:
525:
509:
505:Jardin des plantes
484:
346:
1789:
1788:
1697:Open Tree of Life
1471:Taxon identifiers
1462:
1461:
1447:
1446:
1421:
1364:
1363:
1308:
1294:
1293:
1260:
1257:(Mountain cavies)
1214:(Galea flavidens)
1197:
1165:(Cavia porcellus)
1139:Shiny guinea pig
1124:
1058:
985:Mammalian Biology
778:Prosopis flexuosa
749:Mammalian Species
715:104(938): 491-94.
683:978-0-8018-8221-0
236:
235:
189:D. patagonum
91:
27:Species of rodent
16:(Redirected from
1834:
1782:
1781:
1769:
1768:
1756:
1755:
1754:
1728:
1727:
1718:
1717:
1705:
1704:
1692:
1691:
1679:
1678:
1666:
1665:
1653:
1652:
1640:
1639:
1627:
1626:
1614:
1613:
1601:
1600:
1588:
1587:
1575:
1574:
1562:
1561:
1549:
1548:
1536:
1535:
1526:
1525:
1513:
1512:
1511:
1498:
1497:
1496:
1466:
1420:
1419:
1415:
1397:Lesser capybara
1375:
1327:Patagonian mara
1312:
1307:
1306:
1302:
1259:
1258:
1254:
1196:
1195:
1189:
1173:(Cavia tschudii)
1123:
1122:
1118:
1109:
1056:
1047:
1040:
1033:
1024:
1008:
967:
964:
958:
947:
938:
931:
922:
907:
898:
884:
863:
848:
835:
828:
817:
812:TABER AB (1987)
810:
785:
774:
765:
758:
752:
745:
716:
705:
696:
695:
659:
653:
646:
640:
639:
637:
635:
630:
600:
573:Tierra del Fuego
372:Valdes Peninsula
227:
209:
106:
105:
85:
80:
79:
52:
38:Temporal range:
32:
21:
1842:
1841:
1837:
1836:
1835:
1833:
1832:
1831:
1792:
1791:
1790:
1785:
1777:
1772:
1764:
1759:
1750:
1749:
1744:
1737:Cavia patagonum
1731:
1723:
1721:
1713:
1708:
1700:
1695:
1687:
1684:Observation.org
1682:
1674:
1669:
1661:
1656:
1648:
1643:
1635:
1630:
1622:
1617:
1609:
1604:
1596:
1591:
1583:
1578:
1570:
1565:
1557:
1552:
1544:
1539:
1531:
1529:
1521:
1516:
1507:
1506:
1501:
1492:
1491:
1486:
1473:
1463:
1458:
1443:
1436:Acrobatic cavy
1417:
1416:
1414:
1404:
1360:
1346:
1334:
1320:
1304:
1303:
1290:
1256:
1255:
1253:
1243:
1193:
1192:(Yellow-toothed
1191:
1190:
1188:
1178:
1141:(Cavia fulgida)
1120:
1119:
1117:
1098:
1060:
1051:
1014:
978:
976:
974:Further reading
971:
970:
965:
961:
948:
941:
935:Symp. Zool. Soc
932:
925:
908:
901:
885:
866:
862:, 227: 439–452.
849:
838:
829:
820:
811:
788:
784:, 35(4):707-14.
775:
768:
759:
755:
746:
719:
706:
699:
684:
661:
660:
656:
647:
643:
633:
631:
602:
601:
594:
589:
569:near threatened
561:Southwick's Zoo
553:
476:
338:
298:
266:Patagonian hare
262:Patagonian cavy
240:Patagonian mara
218:
211:
205:
192:
100:
92:
83:Near Threatened
81:
77:
70:
43:
36:
35:Patagonian mara
28:
23:
22:
18:Patagonian Cavy
15:
12:
11:
5:
1840:
1838:
1830:
1829:
1824:
1819:
1814:
1809:
1804:
1794:
1793:
1787:
1786:
1784:
1783:
1770:
1757:
1741:
1739:
1733:
1732:
1730:
1729:
1719:
1706:
1693:
1680:
1667:
1654:
1641:
1628:
1615:
1602:
1589:
1576:
1563:
1550:
1537:
1527:
1514:
1499:
1483:
1481:
1475:
1474:
1469:
1460:
1459:
1452:
1449:
1448:
1445:
1444:
1442:
1441:
1433:
1424:
1422:
1406:
1405:
1403:
1402:
1394:
1385:
1383:
1372:
1370:Hydrochoerinae
1366:
1365:
1362:
1361:
1359:
1358:
1349:
1347:
1336:
1335:
1333:
1332:
1323:
1321:
1309:
1296:
1295:
1292:
1291:
1289:
1288:
1280:
1272:
1263:
1261:
1245:
1244:
1242:
1241:
1238:(Galea spixii)
1233:
1225:
1217:
1209:
1200:
1198:
1180:
1179:
1177:
1176:
1168:
1160:
1152:
1144:
1136:
1133:(Cavia aperea)
1127:
1125:
1106:
1100:
1099:
1097:
1096:
1094:Hystricomorpha
1090:
1084:
1078:
1072:
1065:
1062:
1061:
1052:
1050:
1049:
1042:
1035:
1027:
1021:
1020:
1013:
1012:External links
1010:
991:(5): 389–398.
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864:
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786:
766:
753:
717:
697:
682:
654:
641:
591:
590:
588:
585:
552:
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472:
337:
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1510:
1504:
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1476:
1472:
1467:
1457:
1456:
1450:
1440:
1439:
1434:
1432:
1431:
1426:
1425:
1423:
1418:(Rock cavies)
1413:
1412:
1407:
1401:
1400:
1395:
1393:
1392:
1387:
1386:
1384:
1382:
1381:
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1371:
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1353:Chacoan mara
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1348:
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1344:
1343:
1337:
1331:
1330:
1325:
1324:
1322:
1319:
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1234:
1232:
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1226:
1224:
1223:
1218:
1216:
1215:
1210:
1208:
1207:
1206:(Galea comes)
1202:
1201:
1199:
1187:
1186:
1181:
1175:
1174:
1169:
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1157:(Cavia magna)
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1380:Hydrochoerus
1378:
1354:
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1328:
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1315:
1300:Dolichotinae
1284:
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1205:
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957:203:298-301.
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672:
668:Wilson, D.E.
657:
644:
632:. Retrieved
618:
612:
606:
566:
534:
530:allogrooming
526:
485:
465:
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358:Monte Desert
347:
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206:
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188:
187:
175:
61:, Argentina
59:Temaikèn Zoo
29:
1580:iNaturalist
1503:Wikispecies
1163:Guinea pig
634:12 November
537:conspecific
517:Yerevan Zoo
425:spp., and
403:Pappophorum
384:herbivorous
330:anal glands
322:guinea pigs
296:Description
272:, somewhat
270:herbivorous
40:Pleistocene
1796:Categories
1752:Q122548972
1428:Rock cavy
1342:Pediolagus
1317:Dolichotis
1250:Microcavia
1092:Suborder:
587:References
577:extirpated
494:postpartum
409:Trichloris
320:, such as
302:jackrabbit
290:monogamous
258:Dolichotis
215:Zimmermann
176:Dolichotis
1389:Capybara
1068:Kingdom:
1055:Caviidae
834:30:37–43.
751:, 625:1-5
545:receptive
326:capybaras
314:ungulates
282:Patagonia
278:Argentina
183:Species:
121:Kingdom:
115:Eukaryota
1746:Wikidata
1663:13400212
1598:10764118
1530:BioLib:
1488:Wikidata
1455:Category
1104:Caviinae
1088:Rodentia
1082:Mammalia
1076:Chordata
1074:Phylum:
1070:Animalia
1057:(Cavies)
921:61:3-15.
919:Mammalia
895:Mammalia
692:62265494
515:Mara at
507:in Paris
462:nematode
441:unimodal
428:Prosopis
415:Atriplex
388:monocots
380:antelope
376:clavicle
364:bush or
362:creosote
165:Caviidae
161:Family:
155:Rodentia
145:Mammalia
135:Chordata
131:Phylum:
125:Animalia
111:Domain:
88:IUCN 3.1
42:- Recent
1774:ZooBank
1766:8838269
1650:1001207
1572:2437619
1559:1038700
1494:Q194313
1411:Kerodon
1305:(Maras)
1194:cavies)
1086:Order:
1080:Class:
993:Bibcode
521:Armenia
489:warrens
450:grisons
436:diurnal
397:Chloris
318:caviids
310:humerus
268:. This
252:in the
217:, 1780)
171:Genus:
151:Order:
141:Class:
86: (
1807:Cavies
1725:106451
1722:uBio:
1715:231492
1702:754336
1624:584722
690:
680:
551:Status
456:, and
446:felids
422:Lycium
406:, and
392:dicots
367:larrea
328:, the
306:radius
274:rabbit
256:genus
250:rodent
1689:83194
1676:29091
1611:85192
1593:IRMNG
1585:43802
1546:377CX
1533:37027
1185:Galea
1114:Cavia
666:. In
464:worm
454:foxes
431:spp.
417:lampa
308:than
1761:GBIF
1671:NCBI
1637:6785
1632:IUCN
1619:ITIS
1567:GBIF
688:OCLC
678:ISBN
636:2021
619:2016
354:50°S
324:and
254:mara
238:The
1658:MSW
1645:MDD
1606:ISC
1554:EoL
1541:CoL
1518:ADW
1001:doi
892:) "
623:doi
496:.
352:to
264:or
57:At
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1556::
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999:.
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987:.
942:^
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902:^
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720:^
711:.
700:^
686:.
617:.
611:.
595:^
519:,
470:.
452:,
448:,
419:,
400:,
350:28
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694:.
638:.
625::
609:"
605:"
523:.
242:(
213:(
90:)
20:)
Text is available under the Creative Commons Attribution-ShareAlike License. Additional terms may apply.
