Rodrigues solitaire

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their turns, and the young is not hatch'd till at seven Weeks' end: All the while they are sitting upon it, or are bringing up their young one, which is not able to provide itself in several Months, they will not suffer any other Bird of their Species to come within two hundred Yards round of the Place; But what is very singular, is, the Males will never drive away the Females, only when he perceives one he makes a noise with his Wings to call to the Female, and she drives the unwelcome Stranger away, not leaving it till 'tis without her Bounds. The Female do's the same as to the Males, whom she leaves to the Male, and he drives them away. We have observ'd this several Times, and I affirm it to be true. The Combats between them on this occasion last sometimes pretty long, because the Stranger only turns about, and do's not fly directly from the Nest. However, the others do not forsake it till they have quite driven it out of their Limits. After these Birds have rais'd their young One, and left it to itself, they are always together, which the other Birds are not, and tho' they happen to mingle with other Birds of the same Species, these two Companions never disunite. We have often remark'd, that some Days after the young leaves the Nest, a Company of thirty or forty brings another young one to it, and the now fledg'd Bird, with its Father and Mother joyning with the Band, march to some bye Place. We frequently follow'd them, and found that afterwards the old ones went each their way alone, or in Couples, and left the two young ones together, which we call'd a Marriage.

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little longer in proportion than a turkey's when it lifts up his head. Its eye is black and lively, and its head without comb on cop. They never fly, their wings are too little to support the weight of their bodies ; they serve only to beat themselves and flutter when they call one another. They will whirl about for twenty or thirty times together on the same side during the space of 4 or 5 minutes. The motions of their wings make then a noise very like that of a rattle, and one may hear it two hundred paces off. The bone of their wings grows greater towards the extremity, and forms a little round mass under the feathers as big as a musket ball. That and its beak are the chief defences of this bird. 'Tis very hard to catch in the woods, but easy in open places, because we run faster than they, and sometimes we approach them without much trouble. From March to September they are very fat, and taste admirably well, especially while they are young, some of the males weigh 45 pounds.

1361:. They presumably settled disputes by striking each other with their wings; to aid this purpose, they used the knobs on their wrists. Fractures in their wing bones also indicate that they were used in combat. It has also been suggested that these fractures may have been the result of a hereditary bone disease rather than battle-injuries. But in all extant birds where carpal spurs and knobs are present, these are used as weapons without exceptions. Though some dodo bones have been found with healed fractures, it had weak pectoral muscles and more reduced wings in comparison with the Rodrigues solitaire. Since Rodrigues receives less rainfall and has more seasonal variation than Mauritius, which would have affected the availability of resources on the island, the Rodrigues solitaire would have more reason to evolve aggressive territorial behaviour. Several accounts state that they also defended themselves with a powerful bite. 1679:. However, size dimorphism does occur in some monogamous birds; most other pigeons are monogamous as well. A 2015 article proposed that males invited females into their territories as secondary mates, which would result in the resident female acting aggressively towards the newcomer. Similar behaviour is seen in species that practice resource-defence polygyny. The territories probably provided all the food the birds needed in addition to acting as breeding-areas, and there was probably intense competition for favourable territories. The fact that Rodrigues island shrank by 90% at the end of the 1662:

for I have not been able to discover where they lay. But I have never seen but one little one alone with them, and, if any one tried to approach it, they would bite him very severely. These birds live on seeds and leaves of trees, which they pick up on the ground. They have a gizzard larger than the fist, and what is surprising is that there is found in it a stone of the size of a henn's egg, of oval shape, a little flattened, although this animal cannot swallow anything larger than a small cherry-stone. I have eaten them: they are tolerably well tasted.

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Their weight may have varied substantially due to fat cycles, meaning that individuals were fat during cool seasons, but slim during hot seasons, and may have been as low as 21 kg in males and 13 kg in females. Though male pigeons are usually larger than females, there is no direct evidence for the largest specimens actually being the males of the species, and this has only been assumed based on early works. Though the male was probably largest, this can only be confirmed by molecular sexing techniques, and not skeletal morphology alone.

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in pairs, they preen their plumage or fur with their beak, and keep themselves very clean. They have their toes furnished with hard scales, and run with quickness, mostly among the rocks, where a man, however agile, can hardly catch them. They have a very short beak, of about an inch in length, which is sharp. They, nevertheless, do not attempt to hurt anyone, except when they find someone before them, and, when hardly pressed, try to bite him. They have a small stump of a wing, which has a sort of bullet at its extremity, and serves as a defence.

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careful to adjust themselves, and make them all even with their beaks. The feathers on their thighs are round like shells at the end, and being there very thick, have an agreeable effect. They have two risings on their craws , and the feathers are whiter there than the rest, which livelily represents the fine neck of a beautiful woman. They walk with so much stateliness and good grace that one cannot help admiring them and loving them, by which means their fine mien often saves their lives.

1589: 1633: 1538:. The risings on the crop of the female may have covered glands that produced the crop milk. If the theory is correct, the birds may have practiced a division of labour, where the female stayed and fed the young crop milk, while the male collected food in the crop and delivered it to the female. It has been suggested that the maximum size attained by the solitaire and the dodo was limited by the amount of crop milk they were able to produce for their young during early growth. 801: 122: 5222: 1333: 1402: 1186:. One study measured the largest knob to be 32.9 millimetres (1.30 in) in diameter. The knobs vary in size across individuals, and were entirely absent from 58% of specimens examined for the study. These are thought to be immature birds, or birds without territory. The carpometacarpi of males without the knobs were smaller on average than those with it, but there was little difference between the females. In life, the knobs would have been covered by tough 176: 622: 1387: 5174: 1550:

of the bigness of a Hen's Egg, 'tis somewhat rough, flat on one side and round on the other, heavy and hard. We believe this Stone was there when they were hatched, for let them be never so young, you meet with it always. They never have but one of 'em, and besides, the Passage from the Craw to the Gizard is so narrow, that a like Mass of half Bigness cou'd not pass. It serv'd to whet our Knives better than any other Stone Whatsoever.

1706: 1265: 1345: 5210: 5098: 150: 1645: 5122: 3517: 1697: 5186: 3430: 3347: 3074: 2958: 2742: 2512: 2241: 2045: 1914: 1244:. Its neck and legs were proportionally longer, and the dodo did not possess an equivalent to the carpal knob of the Rodrigues solitaire. Many skeletal features of the Rodrigues solitaire and dodo that are unique among pigeons have evolved to adapt to flightlessness. Their pelvic elements were thicker than those of flighted birds (to support their greater weight), and their 5134: 1787:. The young ones that were brought had the head made more or less like the latter animal, but their feet were similar to those of turkeys, instead of that of the ostrich which is forked and cloven in the shape of a hind's foot. These two birds, when skinned, had an inch of fat on the body. One was made into a pie, which turned out to be so tough that it was uneatable. 5110: 3505: 5162: 1197:, which would have made them appear even larger. Carpal spurs and knobs are also known from other extant as well as extinct birds. Within Columbidae, the crowned pigeons and the Viti Levu giant pigeon have outgrowths on the carpometacarpus which are similar to those of the female Rodrigues solitaire. Other well known examples are the 1087:. The Nicobar and spotted green pigeon were placed at the base of a lineage leading to the Raphinae, which indicates the flightless raphines had ancestors that were able to fly, were semi-terrestrial, and inhabited islands. This in turn supports the hypothesis that the ancestors of those birds reached the Mascarene islands by 1423:, writing about his time on the island around 1735, stated that a captive Rodrigues solitaire (which he described as having a melancholic appearance) would always walk in the same line until running out of space, and then return. The species may have lived primarily in the island's woodlands, rather than on the shores. 322: 1373:

such lesions would likely not occur only in a specific part of the skeleton, but would appear in any growing bone tissue. If such a disease was due to inbreeding, it would also be present in other isolated island bird populations, but it is not. The authors instead suggested that the wing bones contained

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Tho' these Birds will sometimes very familiarly come up near enough to one, when we do not run after them, yet they will never grow Tame. As soon as they are caught they shed Tears without Crying, and refuse all sustenance till they die. We find in the Gizzards of both Male and Female, a brown Stone,

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In 1869, the Newton brothers suggested that the carpal knobs may have been formed through continuous injuries, as they resemble diseased bone. It has also been claimed that the carpal knobs were instead formed due to a hereditary disease caused by inbreeding. This was dismissed in a 2013 study, since

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Our men told of having seen goats and a large quantity of birds of different kinds: they brought, amongst others, two of which were bigger by a third than the largest turkey; they appeared, nevertheless to be still quite young, still having down on the neck and head; their wingtips were but sparsely

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The solitaire is a large bird, which weighs about forty or fifty pounds. They have a very big head, with a sort of frontlet, as if of black velvet. Their feathers are neither feathers nor fur; they are of light grey colour, with a little black on their backs. Strutting proudly about, either alone or

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attempted to obtain a live specimen, as he had been assured the Rodrigues solitaire still survived in remote areas of the island. Though trying for 18 months, and offering large rewards, none could be found. He noted that cats were blamed for decimating the species, but suspected that it was due to

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tissue able to form the knob. This development was either in response to continuous impacts during combat, or to hormones released when individuals paired up and acquired territories. It appears a male which had long held a territory would possess especially large carpal knobs, and that their mates

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Several of Leguat's observations were later confirmed through study of subfossil Rodrigues solitaire remains. The curved contour lines of the pelvis also support the roundness of its hind parts, which he compared to that of a horse. Also, a ridged surface appears at the base of the beak, indicating

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They do not fly at all, having no feathers to their wings, but they flap them, and make a great noise with their wings when angry, and the noise is something like thunder in the distance. They only ly, as I am led to suppose, but once in the year, and only one egg. Not that I have seen their eggs,

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The females are wonderfully beautiful, some fair, some brown. I call them fair, because they are the colour of fair hair ; they have a sort of peak like a widow's, upon their breasts , which is of a dun colour. No one feather is straggling from the other all over their bodies, they being very

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refugees, who were the first to colonise the island from 1691 to 1693, after they were marooned there by their captain. His description of the Rodrigues solitaire and its behaviour is the most detailed account of the bird in life, and he also described other species that are now extinct. Leguat's

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When these Birds build their Nests, they choose a clean Place, gather together some Palm-Leaves for that purpose, and heap them up a foot and a half high from the Ground, on which they sit. They never lay but one Egg, which is much bigger than that of a Goose. The Male and Female both cover it in

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In addition to their use as weapons, both sexes of the Rodrigues solitaire also used their wings for communication. The wings could create low-frequency sounds for communicating with mates, or to warn rivals, but it is unknown exactly how this sound was created. The sound could be heard 200 yards

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The feathers of the males are of a brown-grey colour, the feet and beak are like a turkey's, but a little more crooked. They have scarce any tail, but their hind part covered with feathers is roundish, like the crupper (rump) of a hare. They are taller than turkeys. Their neck is straight, and a

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bird. Males were considerably larger than females, measuring 75.7–90 centimetres (30–35 inches) in height and weighing up to 28 kilograms (62 pounds), whereas females were 63.8–70 centimetres (25–28 in) and weighed 17 kilograms (37 lb). This is only 60% of the weight of a mature male.

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By the time the discovery of subfossil Rodrigues solitaire bones from 1786 onwards confirmed Leguat's descriptions, no living residents of Rodrigues remembered having seen live specimens. In 1831, a man who had lived on Rodrigues for 40 years said that he had never seen birds large enough to be

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Apart from Leguat's rather simple depiction, the life appearance of the Rodrigues solitaire is only known from a handful of descriptions; no soft-tissue remains survive. Leguat devoted three pages of his memoirs to the Rodrigues solitaire, and was clearly impressed by the bird. He described its

689:. In spite of the evidence, some later scholars doubted Leguat's story, and the existence of the Rodrigues solitaire. In 1921, the American linguist Geoffroy Atkinson claimed Leguat's memoir was merely a novel, and that the man had never even existed, and in 1955, the British ecologist 1138:

The beak of the Rodrigues solitaire was slightly hooked, and its neck and legs were long. One observer described it as the size of a swan. The skull was 170 mm (6.7 in) long, flattened at the top with the fore and hind parts elevated into two bony ridges structured with

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in 1848. They dissected the only known dodo specimen with soft tissue, comparing it with the few Rodrigues solitaire remains then available. Strickland stated that, although not identical, these birds shared many distinguishing features in the leg bones otherwise only known in

1580:, but hardly flat on one side as described by Leguat. This could be due to its association with a young individual. Although Leguat asserted that the bird hatched with the gizzard stone already inside, in reality adults most likely fed the stones to their hatchlings. 1666:

The size difference between sexes has led to the suggestion that the Rodrigues solitaire was not monogamous as stated by Leguat, and that this deeply religious man attributed the trait to the bird for moral reasons. It has been proposed that it was instead

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was home not only to a white dodo, but also to a white bird similar to the Rodrigues solitaire, both of which are now believed to be misinterpretations of old reports of the Réunion ibis. An atypical 17th-century description of a dodo and bones found on

455:. Males were much larger than females and measured up to 75.7–90 centimetres (30–35 inches) in height and 28 kilograms (62 pounds) in weight, contrasting with 63.8–70 centimetres (25–28 in) and 17 kilograms (37 lb) for females. Its 1418:

birds have slower growth rates, indicates that the Rodrigues solitaire may have had a protracted development period. Based on mass estimates, it has been suggested the male could reach the age of 28, and the female 17. The French economist

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of the island is heavily damaged. Before humans arrived, forests covered the island entirely, but very little remains today due to deforestation. The Rodrigues solitaire lived alongside other recently extinct birds such as the

1290:". Before fossils of the carpal knob were found, Strickland noted that the keel of the sternum of the Rodrigues solitaire was so well-developed as to almost indicate it had possessed the power of flight; however, since the 1724:

mentioned "dodos" on Rodrigues again in 1634, and they were also mentioned in 1700. The next account, which was the first referring to the bird as the "solitaire", was published in François Leguat's 1708 memoir,

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In 1877 three stones were found in a cavern on Rodrigues, each near a Rodrigues solitaire skeleton, and were inferred to be the gizzard stones mentioned by Leguat. One of the stones was examined and found to be

1756:, which has been attributed to Julien Tafforet, a mariner marooned on Rodrigues in 1726. His observations are considered credible, though it is known he had a copy of Leguat's memoirs with him during his stay. 1795:

sent a "solitaire" to France from the nearby island of Réunion around 1740. Since the Réunion solitaire is believed to have become extinct by this date, the bird may actually have been a Rodrigues solitaire.

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of the bird in detail. Thousands of bones were excavated, and mounted skeletons were composed from the remains of several specimens. Study of skeletal features by the Newtons indicated that the solitaire was

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The Dutch viceadmiral Hans Hendricksz Bouwer was the first to list "dodos", most likely referring to the Rodrigues solitaire, as part of the fauna of Rodrigues in 1601. The English traveller

1624:, which may have followed foraging adults as part of the learning process. A study of subfossil remains found that the carpal knob only developed after the bird reached skeletal maturity. 1182:-like in appearance, and consisted of up to two or three lobes. The knobs were about half the length of the metacarpus, were larger in males than females, and described as the size of a 1545:. Dodos also did this, which may imply a similar diet. Leguat described the stones in the following passage, mentioning that Rodrigues solitaires refused to feed in captivity: 1083:

origin and are less than 10 million years old. Therefore, the ancestors of both birds probably remained capable of flight for a considerable time after the separation of their

486:, and was extinct by the late 18th century. Apart from Leguat's account and drawing, and a few other contemporary descriptions, nothing was known about the bird until a few 459:

was grey and brown; the female was paler than the male. It had a black band at the base of its slightly hooked beak, and its neck and legs were long. Both sexes were highly

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observations are considered some of the first cohesive accounts of animal behaviour in the wild. He later left for Mauritius, but was too late to observe dodos there.

4010: 3362:"Digital reconstruction of Rodrigues Solitaire (Pezophaps solitaria) (Aves: Columbidae) physical appearance based on early descriptive observation and other evidence" 1812:

trade between 1730 and 1750; traders burnt off vegetation, hunted solitaires and imported cats and pigs that preyed on eggs and chicks. In 1755, the French engineer

1046:) at the base of the clade. Based on behavioural and morphological evidence, Jolyon C. Parish proposed that the dodo and Rodrigues solitaire should be placed in the 6308: 5254: 2175:

The Dodo and Its Kindred; or the History, Affinities, and Osteology of the Dodo, Solitaire, and Other Extinct Birds of the Islands Mauritius, Rodriguez, and Bourbon

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has since led to the dissolution of the family Raphidae, and the dodo and solitaire are now placed in their own subfamily, Raphinae, within the family Columbidae.

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Tafforet's account confirms Leguat's description of reproductive behaviour, adding that Rodrigues solitaires would even attack humans approaching their chicks:

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Leguat continued with an elaborate description of the female Rodrigues solitaire, which also appears to be the sex depicted in his illustration of the bird:

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doubted aspects of the bird's biology mentioned by Leguat. Today, it is widely accepted that Leguat's memoirs are credible observations of the bird in life.

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Rodrigues solitaires. Rodrigues covers only 104 square kilometres (40 sq mi), making it implausible that the bird would have survived undetected.

2597:(L.), Aves, Columbiformes] from Mauritius Cave Shelters Highlights Human Predation, with a Comment on the Status of the Family Raphidae Wetmore, 1930" 510:

was the first to refer to the bird as the "solitaire" (referring to its solitary habits), but it has been suggested that he borrowed the name from a 1689

1813: 463:, with large bony knobs on their wings that were used in combat. The Rodrigues solitaire laid a single egg that was incubated in turn by both sexes. 6282: 6321: 4005: 2984: 1791:

Unlike the dodo, no Rodrigues solitaires are known to have been sent to Europe alive. However, it has been claimed that the French naval officer

637:, since an observation station was located on the island. Many of these excavations were requested by the English ornithologists (and brothers) 6429: 1464: 700:

of their own, the Raphidae (formerly Dididae), because their exact relationships with other pigeons were unresolved. Each was also placed in a

3304: 3279: 3181: 3101: 3027: 2901: 2818: 2459: 2420: 1118:. It was only slightly smaller than the Rodrigues solitaire and the dodo, and it too is thought to have been related to the crowned pigeons. 6454: 1599:

The most detailed account of the reproductive habits of the Rodrigues solitaire is Leguat's. He described mating and nesting as follows:

5247: 2709:"Mitochondrial and nuclear DNA sequences support a Cretaceous origin of Columbiformes and a dispersal-driven radiation in the Paleogene" 1774:

noted that some Rodrigues solitaire remains bore traces of having been broken by a human or perhaps another large predator, to extract

2150: 1778:. Writing in 1735, the French lieutenant Gennes de la Chancelière described the capture and consumption of two specimens as follows: 6439: 1143:. A black band (a contemporary description described it as a "frontlet") appeared on its head just behind the base of the beak. The 548:, which caused confusion, until they were compared with other bones from Rodrigues that were found to belong to the same species. 6459: 5460: 1740:

The Huguenots praised the Rodrigues solitaires for their flavour, especially that of the young, and used their gizzard stones as

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offspring, which required extensive parental care until maturity. The gathering of unrelated juveniles suggests that they formed

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feathered, without any proper tail. Three sailors told me of having seen two others, of the same species, as big as the biggest

1420: 1102:. The dodo lost the ability to fly owing to the lack of mammalian predators on Mauritius. Another large, flightless pigeon, the 3549: 2634:

Shapiro, B.; Sibthorpe, D.; Rambaut, A.; Austin, J.; Wragg, G. M.; Bininda-Emonds, O. R. P.; Lee, P. L. M.; Cooper, A. (2002).

1064:) was analysed, and it was found to be a close relative of the Nicobar pigeon, and thus also the dodo and Rodrigues solitaire. 572:

and cared for its nestlings also supported this relationship. Strickland recognised its generic distinction and named the new

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bones were found in a cave in 1786. Thousands of bones have subsequently been excavated. It is the only extinct bird with a

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pigeons and others, in agreement with the genetic evidence In 2014, DNA of the only known specimen of the recently extinct

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shows the closest relationships of the dodo and the Rodrigues solitaire within Columbidae, based on Shapiro et al., 2002:

1974: 1236:. It differed in other aspects; it was taller and more slender than the dodo and had a smaller skull and beak, a flatter 1714:

Leguat's 1708 maps of Rodrigues and his settlement. Rodrigues solitaires are distributed across the maps, many in pairs

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was described as consisting of a single egg; given the bird's large size, this led to proposals that the solitaire was

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It has been proposed that Leguat's comparison between the crop of the female Rodrigues solitaire and the "beautiful

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was very short he inferred that this was instead related to Leguat's claim that they used their wings for defence.

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The Rodrigues solitaire shared traits with the dodo, its closest relative, such as size and features in the skull,

1039: 175: 5232: 1752:. The second most detailed description of the bird was found in an anonymous document rediscovered in 1874 called 1365:(182 m) away, and this may therefore be the size of the territory of an individual. Other species of birds (e.g., 1310:

of a woman" (changed to "fine neck" in some editions of his memoirs) was out of longing for female companionship.

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Voyages et Avantures de François Leguat & de ses Compagnons, en Deux Isles Desertes des Indes Orientales, etc

1826: 1131: 556: 2060:"In the footsteps of the bone collectors: Nineteenth-century cave exploration on Rodrigues Island, Indian Ocean" 4018: 4000: 1833:

after the bird to commemorate the journey. Although the Rodrigues solitaire is the only extinct bird to have a

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of the bird were so large that Strickland thought they belonged to two species, naming the smaller female bird

6172: 3521: 4023: 2836:"On the Comparative Ecological and Evolutionary Significance of Total and Mass-Specific Rates of Metabolism" 1818: 1524:, whereas Tafforet mentioned seeds and leaves. No other accounts mention diet. It has been suggested it ate 527: 311: 6207: 6134: 3990: 3802: 3752: 3485: 1103: 728:

and colleagues analysed the DNA of the dodo and the Rodrigues solitaire for the first time. Comparison of

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may also have contributed to such competition over territories, and thereby furthered sexual dimorphism.

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The 2002 study indicated that the ancestors of the Rodrigues solitaire and the dodo diverged around the

634: 552: 491: 291: 267: 2105:"XVI. On some Bones of Birds allied to the Dodo, in the Collection of the Zoological Society of London" 1484: 633:

Additional subfossils were recovered during the 1860s, but more complete remains were found during the

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sometime between the 1730s and 1760s; the exact date is unknown. Its disappearance coincided with the

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At one point it was suggested that the skeleton of this species is the best described after that of

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Many other of the endemic species of Rodrigues became extinct after the arrival of humans, so the

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of the Rodrigues solitaire was described as grey and brown. Females were paler than males and had

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Leguat's statements were confirmed by another description by Julien Tafforet, who wrote in 1726:

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Many old accounts mention that Rodrigues solitaires were hunted by humans. The Danish zoologist

1252:(underdeveloped, retaining juvenile features). However, the skull, trunk, and pelvic limbs were 507: 471: 130: 6391: 6313: 6010: 5962: 5648: 5593: 5114: 3322:"A possible connection between crop milk and the maximum size attainable by flightless pigeons" 2667: 1759: 1401: 6449: 6360: 6339: 6243: 5553: 5407: 5214: 4894: 4841: 4791: 3934: 3924: 3867: 3667: 3662: 3300: 3275: 3177: 3097: 3023: 2897: 2863: 2814: 2791: 2730: 2658: 2455: 2416: 2386: 2179: 2146: 1830: 1763: 1214: 1076: 952: 800: 750: 611: 495: 452: 5788: 6365: 6186: 5747: 5740: 4760: 3902: 3574: 3463: 3416: 3373: 3333: 3237: 3145: 3060: 2938: 2847: 2781: 2771: 2720: 2650: 2616: 2565: 2484: 2378: 2345: 2310: 2279: 2227: 2116: 2074: 2031: 1989: 1946: 1900: 1456: 1436: 1269: 1099: 697: 470:

First mentioned during the 17th century, the Rodrigues solitaire was described in detail by

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to Leguat's 1708 memoir, showing his settlement on Rodrigues, and a solitaire in the middle

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by his sponsor Marquis Henri Duquesne, which used the name "solitaire" in reference to the

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intermediate between the dodo and ordinary pigeons, but differed from them in its unique

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Worthy, T. H. (2001). "A giant flightless pigeon gen. Et sp. Nov. And a new species of

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competing for resources on these islands allowed the solitaire and the dodo to attain

6423: 6404: 6334: 5897: 5695: 5630: 5478: 5202: 5040: 5024: 4947: 4939: 4773: 4571: 4421: 4308: 4266: 4066: 4058: 3980: 3944: 3862: 3786: 3361: 2504: 2451: 2173: 2086: 1937: 1609: 1592: 1542: 1495: 1218: 1187: 739: 670:, now known to have belonged to the Rodrigues solitaire, led the British taxidermist 642: 638: 580: 464: 160: 72: 3385: 2950: 2875: 2835: 1344: 1332: 621: 6248: 6046: 5797: 5761: 5709: 5533: 5513: 5166: 5126: 4870: 4715: 4531: 4369: 4317: 4189: 4150: 3919: 3907: 3897: 3887: 3584: 3015: 2889: 2443: 1632: 1530: 1521: 1249: 1202: 1198: 1126: 742: 732: 729: 725: 713: 709: 121: 17: 6235: 3297: